Parrot

For other uses, see Parrot (disambiguation).

Parrots, also known as psittacines (/ˈsɪtəsaɪnz/),^[1][2] are birds of the roughly 398 species^[3] in 92 genera comprising the order Psittaciformes (/ˈsɪtəsɪfɔːrmiːz/), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoidea ("true" parrots), the Cacatuoidea (cockatoos), and the Strigopoidea (New Zealand parrots). One-third of all parrot species are threatened by extinction, with higher aggregate extinction risk (IUCN Red List Index) than any other comparable bird group.^[4] Parrots have a generally pantropical distribution with several species inhabiting temperate regions in the Southern Hemisphere, as well. The greatest diversity of parrots is in South America^[5] and Australasia.^[6]

Parrots Temporal range: Eocene–Holocene 50–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Clade:	Psittacopasserae
Order:	Psittaciformes Wagler, 1830
Superfamilies
Vastanavis Quercypsitta Paleopsittacus Cacatuoidea (cockatoos) Psittacoidea (true parrots) Strigopoidea (New Zealand parrots)
Range of parrots, all species (red)

Characteristic features of parrots include a strong, curved bill, an upright stance, strong legs, and clawed zygodactyl feet. Many parrots are vividly coloured, and some are multi-coloured. Most parrots exhibit little or no sexual dimorphism in the visual spectrum. They form the most variably sized bird order in terms of length.

The most important components of most parrots' diets are seeds, nuts, fruit, buds, and other plant material. A few species sometimes eat animals and carrion, while the lories and lorikeets are specialised for feeding on floral nectar and soft fruits. Almost all parrots nest in tree hollows (or nest boxes in captivity), and lay white eggs from which hatch altricial (helpless) young.

Parrots, along with ravens, crows, jays, and magpies, are among the most intelligent birds, and the ability of some species to imitate human speech enhances their popularity as pets. Trapping wild parrots for the pet trade, as well as hunting, habitat loss, and competition from invasive species, has diminished wild populations, with parrots being subjected to more exploitation than any other group of birds. As of 2021, about 50 million parrots (half of all parrots) live in captivity, with the vast majority of these living as pets in people's homes.^[7] Measures taken to conserve the habitats of some high-profile charismatic species have also protected many of the less charismatic species living in the same ecosystems.

Parrots are the only creatures that display true tripedalism, using their necks and beaks as limbs with propulsive forces equal to or greater than those forces generated by the forelimbs of primates when climbing vertical surfaces. They can travel with cyclical tripedal gaits when climbing.^[8]

Taxonomy

Origins and evolution

 

Fossil dentary specimen UCMP 143274 restored as a parrot (left) or an oviraptorosaur

Psittaciform diversity in South America and Australasia suggests that the order may have evolved in Gondwana, centred in Australasia.^[9] The scarcity of parrots in the fossil record, however, presents difficulties in confirming the hypothesis. There is currently a higher amount of fossil remains from the northern hemisphere in the early Cenozoic.^[10] Molecular studies suggest that parrots evolved approximately 59 million years ago (Mya) (range 66–51 Mya) in Gondwana. The three major clades of Neotropical parrots originated about 50 Mya (range 57–41 Mya).^[11]

A single 15 mm (0.6 in) fragment from a large lower bill (UCMP 143274), found in deposits from the Lance Creek Formation in Niobrara County, Wyoming, had been thought to be the oldest parrot fossil and is presumed to have originated from the Late Cretaceous period, which makes it about 70 million years old.^[12] However, other studies suggest that this fossil is not from a bird, but from a caenagnathid oviraptorosaur (a non-avian dinosaur with a birdlike beak), as several details of the fossil used to support its identity as a parrot are not actually exclusive to parrots, and it is dissimilar to the earliest-known unequivocal parrot fossils.^[13][14]

It is generally assumed that the Psittaciformes were present during the Cretaceous–Paleogene extinction event (K-Pg extinction), 66 mya. They were probably generalised arboreal birds, and did not have the specialised crushing bills of modern species.^[10][15] Genomic analysis provides strong evidence that parrots are the sister group of passerines, forming the clade Psittacopasserae, which is the sister group of the falcons.^[16]

The first uncontroversial parrot fossils date to tropical Eocene Europe around 50 mya. Initially, a neoavian named Mopsitta tanta, uncovered in Denmark's Early Eocene Fur Formation and dated to 54 mya, was assigned to the Psittaciformes. However, the rather nondescript bone is not unequivocally psittaciform, and it may rather belong to the ibis genus Rhynchaeites, whose fossil legs were found in the same deposits.^[17]

 

Fossil skull of a presumed parrot relative from the Eocene Green River Formation in Wyoming

Several fairly complete skeletons of parrot-like birds have been found in England and Germany.^[18] These are probably not transitional fossils between ancestral and modern parrots, but rather lineages that evolved parallel to true parrots and cockatoos:^[19]

• Psittacopes
• Serudaptus
• Halcyornithidae
  • Cyrilavis
  • Halcyornis
  • Pulchrapollia
  • Pseudasturides
• Vastanavidae
  • Vastanavis
• Quercypsittidae
  • Quercypsitta
• Messelasturidae^[20]
  • Messelastur
  • Tynskya

The earliest records of modern parrots date to around 23–20 mya.^[21] The fossil record—mainly from Europe—consists of bones clearly recognisable as belonging to anatomically modern parrots.^[22] The Southern Hemisphere contains no known parrot-like remains earlier than the Early Miocene around 20 mya.^[21]

Etymology

The name 'Psittaciformes' comes from the ancient Greek for parrot, ψιττακός ('Psittacus'), whose origin is unclear. Ctesias (5th century BCE) recorded the name Psittacus after the Indian name for a bird, most likely a parakeet (now placed in the genus Psittacula). Pliny the Elder (23/24–79 CE) in his Natural History (book 10, chapter 58) noted that the Indians called the bird as "siptaces"; however, no matching Indian name has been traced.^[23][24]

Phylogeny

Parrots     Psittacoidea     Cacatuoidea     Strigopoidea     Other birds

Phylogenetic relationship between the three parrot superfamilies^[9]^[25]^[26]

The Psittaciformes comprise three main lineages: Strigopoidea, Psittacoidea and Cacatuoidea.^[27] The Strigopoidea were considered part of the Psittacoidea, but the former is now placed at the base of the parrot tree next to the remaining members of the Psittacoidea, as well as all members of the Cacatuoidea.^[9][25][26] The Cacatuoidea are quite distinct, having a movable head crest, a different arrangement of the carotid arteries, a gall bladder, differences in the skull bones, and lack the Dyck texture feathers that—in the Psittacidae—scatter light to produce the vibrant colours of so many parrots. Colourful feathers with high levels of psittacofulvin resist the feather-degrading bacterium Bacillus licheniformis better than white ones.^[28] Lorikeets were previously regarded as a third family, Loriidae,^[29]: 45 but are now considered a tribe (Loriini) within the subfamily Loriinae, family Psittaculidae. The two other tribes in the subfamily are the closely related fig parrots (two genera in the tribe Cyclopsittini) and budgerigar (tribe Melopsittacini).^[9][25][26]

Strigopoidea  Nestoridae  Strigopidae     Cacatuoidea  Cacatuidae  Psittacoidea  Psittacidae   Neotropical parrots   Psittacinae Psittrichasiidae   Psittrichadinae   Coracopseinae Psittaculidae Psittaculinae Broad-tailed parrots Fig parrots Budgerigar Lories and Lorikeets Bolbopsittacus Hanging parrots Lovebirds Psittacella

Phylogenetic relations between parrots^[9]

Systematics

Main article: List of parrots

The order Psittaciformes consists of roughly 393 species belonging to 92 genera.^{[30][9][25][27][31][32][33][34]}

 

Skeleton of a parrot

Superfamily Strigopoidea: New Zealand parrots

• Family Nestoridae: two genera with two living (kea and New Zealand kaka) and several extinct species of the New Zealand region
• Family Strigopidae: the flightless, critically endangered kakapo of New Zealand

Superfamily Cacatuoidea: cockatoos

• Family Cacatuidae
  • Subfamily Nymphicinae: one genus with one species, the cockatiel.
  • Subfamily Calyptorhynchinae: the black cockatoos
  • Subfamily Cacatuinae
    • Tribe Microglossini: one genus with one species, the black palm cockatoo
    • Tribe Cacatuini: four genera of white, pink, and grey species

Superfamily Psittacoidea: true parrots

• Family Psittacidae
  • Subfamily Psittacinae: two African genera, Psittacus and Poicephalus
  • Subfamily Arinae
    • Tribe Arini: 18 genera
    • Tribe Androglossini: seven genera.
• Family Psittrichasiidae
  • Subfamily Psittrichasinae: one species, Pesquet's parrot
  • Subfamily Coracopsinae: one genus with four species.
• Family Psittaculidae
  • Subfamily Platycercinae
    • Tribe Pezoporini: ground parrots and allies
    • Tribe Platycercini: broad-tailed parrots
  • Subfamily Psittacellinae: one genus (Psittacella) with several species
  • Subfamily Loriinae
    • Tribe Loriini: lories and lorikeets
    • Tribe Melopsittacini: one genus with one species, the budgerigar
    • Tribe Cyclopsittini: fig parrots
  •  

    Blue and gold macaw (Ara ararauna) skeleton on display at the Museum of Osteology.
    Subfamily Agapornithinae: three genera
  • Subfamily Psittaculinae
    • Tribe Polytelini: three genera
    • Tribe Psittaculini: Asian psittacines
    • Tribe Micropsittini: pygmy parrots

Morphology

 

Glossy black cockatoo showing the parrot's strong bill, clawed feet, and sideways-positioned eyes

Living species range in size from the buff-faced pygmy parrot, at under 10 g (0.4 oz) in weight and 8 cm (3.1 in) in length,^[29]: 149 to the hyacinth macaw, at 1 m (3.3 ft) in length,^[35] and the kakapo, at 4.0 kg (8.8 lb) in weight.^[36] Among the superfamilies, the three extant Strigopoidea species are all large parrots, and the cockatoos tend to be large birds, as well. The Psittacoidea parrots are far more variable, ranging the full spectrum of sizes shown by the family.^[36]

The most obvious physical characteristic is the strong, curved, broad bill. The upper mandible is prominent, curves downward, and comes to a point. It is not fused to the skull, which allows it to move independently, and contributes to the tremendous biting pressure the birds are able to exert. A large macaw, for example, has a bite force of 35 kg/cm² (500 lb/sq in), close to that of a large dog.^[37] The lower mandible is shorter, with a sharp, upward-facing cutting edge, which moves against the flat portion of the upper mandible in an anvil-like fashion. Touch receptors occur along the inner edges of the keratinised bill, which are collectively known as the "bill tip organ", allowing for highly dexterous manipulations. Seed-eating parrots have a strong tongue (containing similar touch receptors to those in the bill tip organ), which helps to manipulate seeds or position nuts in the bill so that the mandibles can apply an appropriate cracking force. The head is large, with eyes positioned high and laterally in the skull, so the visual field of parrots is unlike any other birds. Without turning its head, a parrot can see from just below its bill tip, all above its head, and quite far behind its head. Parrots also have quite a wide frontal binocular field for a bird, although this is nowhere near as large as primate binocular visual fields.^[38] Unlike humans, the vision of parrots is also sensitive to ultraviolet light.^[39]

Parrots have strong zygodactyl feet (two toes facing forward and two back) with sharp, elongated claws, which are used for climbing and swinging. Most species are capable of using their feet to manipulate food and other objects with a high degree of dexterity, in a similar manner to a human using their hands. A study conducted with Australian parrots has demonstrated that they exhibit "handedness", a distinct preference with regards to the foot used to pick up food, with adult parrots being almost exclusively "left-footed" or "right-footed", and with the prevalence of each preference within the population varying by species.^[40]

 

Eclectus parrots, male left and female right

Cockatoo species have a mobile crest of feathers on the top of their heads, which they can raise for display, and retract.^[41] No other parrots can do so, but the Pacific lorikeets in the genera Vini and Phigys can ruffle the feathers of the crown and nape, and the red-fan parrot (or hawk-headed parrot) has a prominent feather neck frill that it can raise and lower at will. The predominant colour of plumage in parrots is green, though most species have some red or another colour in small quantities. Cockatoos, however, are predominately black or white with some red, pink, or yellow.^[42] Strong sexual dimorphism in plumage is not typical among parrots, with some notable exceptions, the most striking being the eclectus parrot.^{[29]: 202–207} However it has been shown that some parrot species exhibit sexually dimorphic plumage in the ultraviolet spectrum, normally invisible to humans.^[43][44]

Distribution and habitat

 

Most parrot species are tropical, but a few species, like this austral parakeet, range deeply into temperate zones.

See also: List of Psittaciformes by population

Parrots are found on all tropical and subtropical continents and regions including Australia and Oceania,^[6] South Asia, Southeast Asia, Central America, South America,^[5] and Africa.^[45] Some Caribbean and Pacific islands are home to endemic species.^[46] By far the greatest number of parrot species come from Australasia and South America.^[47] The lories and lorikeets range from Sulawesi and the Philippines in the north to Australia and across the Pacific as far as French Polynesia, with the greatest diversity being found in and around New Guinea.^[46] The subfamily Arinae encompasses all the neotropical parrots, including the amazons, macaws, and conures, and ranges from northern Mexico and the Bahamas to Tierra del Fuego in the southern tip of South America.^[48] The pygmy parrots, tribe Micropsittini, form a small genus restricted to New Guinea and the Solomon Islands.^[49] The superfamily Strigopoidea contains three living species of aberrant parrots from New Zealand.^[50] The broad-tailed parrots, subfamily Platycercinae, are restricted to Australia, New Zealand, and the Pacific islands as far eastwards as Fiji.^[51] The true parrot superfamily, Psittacoidea, includes a range of species from Australia and New Guinea to South Asia and Africa.^[46] The centre of cockatoo biodiversity is Australia and New Guinea, although some species reach the Solomon Islands (and one formerly occurred in New Caledonia),^[52] Wallacea and the Philippines.^[53]

 

The kea is the only alpine parrot.

Several parrots inhabit the cool, temperate regions of South America and New Zealand. Three species—the thick-billed parrot, the green parakeet, and the now-extinct Carolina parakeet—have lived as far north as the southern United States. Many parrots, especially monk parakeets, have been introduced to areas with temperate climates, and have established stable populations in parts of the United States (including New York City),^[54] the United Kingdom,^[55] Belgium,^[56] Spain^[57][58] and Greece.^[59] These birds can be quite successful in introduced areas, such as the non-native population of red-crowned amazons in the U.S. which may rival that of their native Mexico.^[60] The only parrot to inhabit alpine climates is the kea, which is endemic to the Southern Alps mountain range on New Zealand's South Island.^[61]

Few parrots are wholly sedentary or fully migratory. Most fall somewhere between the two extremes, making poorly understood regional movements, with some adopting an entirely nomadic lifestyle.^[62] Only three species are migratory – the orange-bellied, blue-winged and swift parrots.^[63]

Behaviour

 

Macaw parrot sitting on a tree branch

 

A rose-ringed parakeet

Numerous challenges are found in studying wild parrots, as they are difficult to catch and once caught, they are difficult to mark. Most wild bird studies rely on banding or wing tagging, but parrots chew off such attachments.^[62] Parrots also tend to range widely, and consequently many gaps occur in knowledge of their behaviour. Some parrots have a strong, direct flight. Most species spend much of their time perched or climbing in tree canopies. They often use their bills for climbing by gripping or hooking on branches and other supports. On the ground, parrots often walk with a rolling gait.^[38]

Diet

A yellow-tailed black cockatoo using its strong bill to search for grubs

The diet of parrots consists of seeds, fruit, nectar, pollen, buds, and sometimes arthropods and other animal prey. The most important of these for most true parrots and cockatoos are seeds; the large and powerful bill has evolved to open and consume tough seeds. All true parrots, except the Pesquet's parrot, employ the same method to obtain the seed from the husk; the seed is held between the mandibles and the lower mandible crushes the husk, whereupon the seed is rotated in the bill and the remaining husk is removed.^[62] They may use their foot sometimes to hold large seeds in place. Parrots are granivores rather than seed dispersers, and in many cases where they are seen consuming fruit, they are only eating the fruit to get at the seed. As seeds often have poisons that protect them, parrots carefully remove seed coats and other chemically defended fruit parts prior to ingestion. Many species in the Americas, Africa, and Papua New Guinea consume clay, which releases minerals and absorbs toxic compounds from the gut.^[64]

 

Chestnut-fronted macaws, yellow-crowned amazons, and dusky-headed parakeets at a clay lick in Ecuador

Geographical range and body size predominantly explains diet composition of Neotropical parrots rather than phylogeny.^[65]

Lories, lorikeets, hanging parrots, and swift parrots are primarily nectar and pollen consumers, and have tongues with brush tips to collect it, as well as some specialised gut adaptations. Many other species also consume nectar when it becomes available.^[66][67]

Some parrot species prey on animals, especially invertebrate larvae. Golden-winged parakeets prey on water snails,^[68] the New Zealand kea can, though uncommonly, hunt adult sheep,^[69] and the Antipodes parakeet, another New Zealand parrot, enters the burrows of nesting grey-backed storm petrels and kills the incubating adults.^[70] Some cockatoos and the New Zealand kaka excavate branches and wood to feed on grubs; the bulk of the yellow-tailed black cockatoo's diet is made up of insects.^[71]

Some extinct parrots had carnivorous diets. Pseudasturids were probably cuckoo- or puffbird-like insectivores, while messelasturids were raptor-like carnivores.^[20]

Breeding

With few exceptions, parrots are monogamous breeders who nest in cavities and hold no territories other than their nesting sites.^[62][72] The pair bonds of the parrots and cockatoos are strong and a pair remains close during the nonbreeding season, even if they join larger flocks. As with many birds, pair bond formation is preceded by courtship displays; these are relatively simple in the case of cockatoos. In Psittacidae parrots' common breeding displays, usually undertaken by the male, include slow, deliberate steps known as a "parade" or "stately walk" and the "eye-blaze", where the pupil of the eye constricts to reveal the edge of the iris.^[62] Allopreening is used by the pair to help maintain the bond. Cooperative breeding, where birds other than the breeding pair help raise the young and is common in some bird families, is extremely rare in parrots, and has only unambiguously been demonstrated in the El Oro parakeet and the golden parakeet (which may also exhibit polygamous, or group breeding, behaviour with multiple females contributing to the clutch).^[73]

 

The vast majority of parrots are, like this rose-ringed parakeet, cavity nesters.

Only the monk parakeet and five species of lovebirds build nests in trees,^[74] and three Australian and New Zealand ground parrots nest on the ground. All other parrots and cockatoos nest in cavities, either tree hollows or cavities dug into cliffs, banks, or the ground. The use of holes in cliffs is more common in the Americas. Many species use termite nests, possibly to reduce the conspicuousness of the nesting site or to create a favourable microclimate.^[75] In most cases, both parents participate in the nest excavation. The length of the burrow varies with species, but is usually between 0.5 and 2 m (1.6 and 6.6 ft) in length. The nests of cockatoos are often lined with sticks, wood chips, and other plant material. In the larger species of parrots and cockatoos, the availability of nesting hollows may be limited, leading to intense competition for them both within the species and between species, as well as with other bird families. The intensity of this competition can limit breeding success in some cases.^[76][77] Hollows created artificially by arborists have proven successful in boosting breeding rates in these areas.^[78] Some species are colonial, with the burrowing parrot nesting in colonies up to 70,000 strong.^[79] Coloniality is not as common in parrots as might be expected, possibly because most species adopt old cavities rather than excavate their own.^[80]

The eggs of parrots are white. In most species, the female undertakes all the incubation, although incubation is shared in cockatoos, the blue lorikeet, and the vernal hanging parrot. The female remains in the nest for almost all of the incubation period and is fed both by the male and during short breaks. Incubation varies from 17 to 35 days, with larger species having longer incubation periods. The newly born young are altricial, either lacking feathers or with sparse white down. The young spend three weeks to four months in the nest, depending on species, and may receive parental care for several months thereafter.^[81]

As typical of K-selected species, the macaws and other larger parrot species have low reproductive rates. They require several years to reach maturity, produce one or very few young per year, and do not necessarily breed every year.^[82]: 125

Intelligence and learning

 

Sun conure demonstrating parrots' puzzle-solving skills

See also: Bird intelligence

Some grey parrots have shown an ability to associate words with their meanings and form simple sentences. Along with crows, ravens, and jays (family Corvidae), parrots are considered the most intelligent of birds. The brain-to-body size ratio of psittacines and corvines is comparable to that of higher primates.^[83] Instead of using the cerebral cortex like mammals, birds use the mediorostral HVC for cognition.^{[84][failed verification]} Not only have parrots demonstrated intelligence through scientific testing of their language-using ability, but also some species of parrots, such as the kea, are also highly skilled at using tools and solving puzzles.^[85]

Learning in early life is apparently important to all parrots, and much of that learning is social learning. Social interactions are often practised with siblings, and in several species, crèches are formed with several broods. Foraging behaviour is generally learnt from parents, and can be a very protracted affair. Generalists and specialists generally become independent of their parents much quicker than partly specialised species who may have to learn skills over long periods as various resources become seasonally available. Play forms a large part of learning in parrots; play can be solitary or social. Species may engage in play fights or wild flights to practice predator evasion. An absence of stimuli can delay the development of young birds, as demonstrated by a group of vasa parrots kept in tiny cages with domesticated chickens from the age of 3 months; at 9 months, these birds still behaved in the same way as 3-month-olds, but had adopted some chicken behaviour.^[62] In a similar fashion, captive birds in zoo collections or pets can, if deprived of stimuli, develop stereotyped and harmful behaviours like self-plucking. Aviculturists working with parrots have identified the need for environmental enrichment to keep parrots stimulated.^[86]

Sound imitation and speech

Main article: Talking bird

See also: Animal language

Video of an orange-winged amazon saying "hello" having been prompted by some humans

Many parrots can imitate human speech or other sounds. A study by scientist Irene Pepperberg suggested a high learning ability in a grey parrot named Alex. Alex was trained to use words to identify objects, describe them, count them, and even answer complex questions such as "How many red squares?" with over 80% accuracy.^[87] N'kisi, another grey parrot, has been shown to have a vocabulary around a thousand words, and has displayed an ability to invent and use words in context in correct tenses.^[88]

Parrots do not have vocal cords, so sound is accomplished by expelling air across the mouth of the trachea in the organ called the syrinx. Different sounds are produced by changing the depth and shape of the trachea.^[89] Grey parrots are known for their superior ability to imitate sounds and human speech, which has made them popular pets since ancient times.^[90]

Although most parrot species are able to imitate, some of the amazon parrots are generally regarded as the next-best imitators and speakers of the parrot world. The question of why birds imitate remains open, but those that do often score very high on tests designed to measure problem-solving ability. Wild grey parrots have been observed imitating other birds.^[91]

Song

Parrots are unusual among birds due to their learned vocalizations, a trait they share with only hummingbirds and songbirds.^[92] The syrinx (vocal organ) of parrots, which aids in their ability to produce song, is located at the base of the trachea and consists of two complex syringeal muscles that allow for the production of sound vibrations, and a pair of lateral tympaniform membranes that control sound frequency.^[93] The position of the syrinx in birds allows for directed air flow into the interclavicular air sacs according to air sac pressure, which in turn creates a higher and louder tone in birds’ singing.^[92]

Cooperation

A 2011 study stated that some African grey parrots preferred to work alone, while others like to work together.^[94] With two parrots, they know the order of tasks or when they should do something together at once, but they have trouble exchanging roles. With three parrots, one parrot usually prefers to cooperate with one of the other two, but all of them are cooperating to solve the task.^[95]

Relationship with humans

Pets

Further information: Companion parrot

 

Pet Cuban amazons in Cuba

Parrots may not make good pets for most people because of their natural wild instincts such as screaming and chewing. Although parrots can be very affectionate and cute when immature, they often become aggressive when mature (partly due to mishandling and poor training) and may bite, causing serious injury.^[96] For this reason, parrot rescue groups estimate that most parrots are surrendered and rehomed through at least five homes before reaching their permanent destinations or before dying prematurely from unintentional or intentional neglect and abuse. The parrots' ability to mimic human words and their bright colours and beauty prompt impulse buying from unsuspecting consumers. The domesticated budgerigar, a small parrot, is the most popular of all pet bird species.^[97] In 1992, the newspaper USA Today published that 11 million pet birds were in the United States alone,^[98] many of them parrots. Europeans kept birds matching the description of the rose-ringed parakeet (or called the ring-necked parrot), documented particularly in a first-century account by Pliny the Elder.^[99] As they have been prized for thousands of years for their beauty and ability to talk, they have also often been misunderstood. For example, author Wolfgang de Grahl says in his 1987 book The Grey Parrot that some importers had parrots drink only coffee while they were shipped by boat, believing that pure water was detrimental and that their actions would increase survival rates during shipping.^[100] Nowadays, it is commonly accepted that the caffeine in coffee is toxic to birds.^[101]

Pet parrots may be kept in a cage or aviary; though generally, tame parrots should be allowed out regularly on a stand or gym. Depending on locality, parrots may be either wild-caught or be captive-bred, though in most areas without native parrots, pet parrots are captive-bred. Parrot species that are commonly kept as pets include conures, macaws, amazon parrots, cockatoos, greys, lovebirds, cockatiels, budgerigars, caiques, parakeets, and Eclectus, Pionus, and Poicephalus species. Temperaments and personalities vary even within a species, just as with dog breeds. Grey parrots are thought to be excellent talkers, but not all grey parrots want to talk, though they have the capability to do so. Noise level, talking ability, cuddliness with people, and care needs can sometimes depend on how the bird is cared for and the attention he/she regularly receives.^[102]

 

Scarlet macaw riding a tricycle at a show in Spain

Parrots invariably require an enormous amount of attention, care, and intellectual stimulation to thrive, akin to that required by a three-year-old child, which many people find themselves unable to provide in the long term.^[103] Parrots that are bred for pets may be hand fed or otherwise accustomed to interacting with people from a young age to help ensure they become tame and trusting. However, even when hand fed, parrots revert to biting and aggression during hormonal surges and if mishandled or neglected.^[104] Parrots are not low-maintenance pets; they require feeding, grooming, veterinary care, training, environmental enrichment through the provision of toys, exercise, and social interaction (with other parrots or humans) for good health.^[105]

Some large parrot species, including large cockatoos, amazons, and macaws, have very long lifespans, with 80 years being reported,^[106] and record ages of over 100.^[107] Small parrots, such as lovebirds, hanging parrots, and budgies, have shorter lifespans up to 15–20 years.^[108] Some parrot species can be quite loud, and many of the larger parrots can be destructive and require a very large cage, and a regular supply of new toys, branches, or other items to chew up.^[102] The intelligence of parrots means they are quick to learn tricks and other behaviours—both good and bad—that get them what they want, such as attention or treats.^[105]

The popularity, longevity, and intelligence of many of the larger kinds of pet parrots and their wild traits such as screaming, has led to many birds needing to be rehomed during the course of their long lifespans. A common problem is that large parrots that are cuddly and gentle as juveniles mature into intelligent, complex, often demanding adults who can outlive their owners, and can also become aggressive or even dangerous. Due to an increasing number of homeless parrots, they are being euthanised like dogs and cats, and parrot adoption centres and sanctuaries are becoming more common.^{[82]: 77–78} Parrots do not often do well in captivity, causing some parrots to go insane and develop repetitive behaviours, such as swaying and screaming, or they become riddled with intense fear. Feather destruction and self-mutilation, although not commonly seen in the wild, occur frequently in captivity.^[109][110]

Trade

 

Hyacinth macaws were taken from the wild for the pet trade in the 1980s.^[111] As a result, Brazil now has only a very small number of breeding pairs left in the wild.^[112]

Main article: International parrot trade

The popularity of parrots as pets has led to a thriving—and often illegal—trade in the birds, and some species are now threatened with extinction. A combination of trapping of wild birds and damage to parrot habitats makes survival difficult or even impossible for some species of parrot. Importation of wild-caught parrots into the US and Europe is illegal after the Wild Bird Population Act was passed in 1992.^[113]

The scale of the problem can be seen in the Tony Silva case of 1996, in which a parrot expert and former director at Tenerife's Loro Parque (Europe's largest parrot park) was jailed in the United States for 82 months and fined $100,000 for smuggling hyacinth macaws (such birds command a very high price.)^[114]

Different nations have different methods of handling internal and international trade. Australia has banned the export of its native birds since 1960.^[115] In July 2007, following years of campaigning by NGOs and outbreaks of avian flu, the European Union (EU) halted the importation of all wild birds with a permanent ban on their import.^[116] Prior to an earlier temporary ban started in late October 2005, the EU was importing about two million live birds a year, about 90% of the international market: hundreds of thousands of these were parrots.^[117] No national laws protect feral parrot populations in the U.S.^[118]

Mexico has a licensing system for capturing and selling native birds.^[119] According to a 2007 report, 65,000 to 78,500 parrots are captured annually, but the mortality rate before reaching a buyer is over 75%, meaning around 50,000 to 60,000 will die.^[120]

Culture

 

Moche parrot, 200 CE Larco Museum Collection Lima, Peru

Parrots have featured in human writings, story, art, humor, religion, and music for thousands of years, such as Aesop's fable "The parrot and the cat"^[121] the Masnavi by Rumi of Persia in 1250 "The Merchant and the Parrot".^[122] Recent books about parrots in human culture include Parrot Culture.^[123]

In ancient times and current, parrot feathers have been used in ceremonies and for decoration.^[124] They also have a long history as pets, stretching back thousands of years, and were often kept as a symbol of royalty or wealth.^[125]

Parrots are used as symbols of nations and nationalism. A parrot is found on the flag of Dominica and two parrots on their coat of arms.^[126] The St. Vincent parrot is the national bird of St. Vincent and the Grenadines, a Caribbean nation.^[127]

Sayings about parrots colour the modern English language. The verb "parrot" in the dictionary means "to repeat by rote". Also clichés such as the British expression "sick as a parrot" are given; although this refers to extreme disappointment rather than illness, it may originate from the disease of psittacosis, which can be passed to humans.^[128][129] The first occurrence of a related expression is in Aphra Behn's 1681 play The False Count.^[130] Fans of Jimmy Buffett are known as parrotheads.^[131] Parrots feature in many media. Magazines are devoted to parrots as pets, and to the conservation of parrots.^[132] Fictional media include Monty Python's "Dead Parrot sketch",^[133] Home Alone 3^[134] and Rio;^[135] and documentaries include The Wild Parrots of Telegraph Hill.^[136]

Mythology

In Polynesian legend as current in the Marquesas Islands, the hero Laka/Aka is mentioned as having undertaken a long and dangerous voyage to Aotona in what are now the Cook Islands, to obtain the highly prized feathers of a red parrot as gifts for his son and daughter. On the voyage, 100 of his 140 rowers died of hunger on their way, but the survivors reached Aotona and captured enough parrots to fill 140 bags with their feathers.^[137][138]

Parrots have also been considered sacred. The Moche people of ancient Peru worshipped birds and often depicted parrots in their art.^[139] Parrots are popular in Buddhist scripture and many writings about them exist. For example, Amitābha once changed himself into a parrot to aid in converting people. Another old story tells how after a forest caught fire, the parrot was so concerned, it carried water to try to put out the flames. The ruler of heaven was so moved upon seeing the parrot's act, he sent rain to put out the fire.^[140] In Chinese Buddhist iconography, a parrot is sometimes depicted hovering on the upper right side Guan Yin clasping a pearl or prayer beads in its beak.^[141]

Feral populations

 

Feral red-masked parakeets in San Francisco

Main article: Feral parrot

Escaped parrots of several species have become established in the wild outside their natural ranges and in some cases outside the natural range of parrots. Among the earliest instances were pet red shining-parrots from Fiji, which established a population on the islands of southern Tonga. These introductions were prehistoric and red-shining parrots were recorded in Tonga by Captain Cook in the 1770s.^[52] Escapees first began breeding in cities in California, Texas, and Florida in the 1950s (with unproven earlier claims dating back to the 1920s in Texas and Florida).^[57] They have proved surprisingly hardy in adapting to conditions in Europe and North America. They sometimes even multiply to the point of becoming a nuisance or pest, and a threat to local ecosystems, and control measures have been used on some feral populations.^[142]

Feral parrot flocks can be formed after mass escapes of newly imported, wild-caught parrots from airports or quarantine facilities. Large groups of escapees have the protection of a flock and possess the skills to survive and breed in the wild.^[143] Some feral parakeets may have descended from escaped zoo birds. Escaped or released pets rarely contribute to establishing feral populations, as they usually result in only a few escapees, and most captive-born birds do not possess the necessary survival skills to find food or avoid predators and often do not survive long without human caretakers. However, in areas where there are existing feral parrot populations, escaped pets may sometimes successfully join these flocks.^[143][144] The most common years that feral parrots were released to non-native environments was from the 1890s to the 1940s, during the wild-caught parrot era.^[144] In the "parrot fever" panic of 1930, a city health commissioner urged everyone who owned a parrot to put them down, but some owners abandoned their parrots on the streets.^[145]

Threats and conservation

 

The Norfolk kaka went extinct in the mid-1800s due to overhunting and habitat loss.^[146]

The principal threats of parrots are habitat loss and degradation, hunting, and, for certain species, the wild-bird trade.^[4] Parrots are persecuted because, in some areas, they are (or have been) hunted for food and feathers, and as agricultural pests. For a time, Argentina offered a bounty on monk parakeets for that reason, resulting in hundreds of thousands of birds being killed, though apparently this did not greatly affect the overall population.^[147]

Parrots, being cavity nesters, are vulnerable to the loss of nesting sites and to competition with introduced species for those sites. The loss of old trees is a particular problem in some areas, particularly in Australia, where suitable nesting trees must be centuries old. Many parrots occur only on islands and are vulnerable to introduced species such as rats and feral cat, as they lack the appropriate antipredator behaviours needed to deal with predators.^[148] Island species, such as the Puerto Rican amazon, which have small populations in restricted habitats, are also vulnerable to natural events, such as hurricanes.^[149] Due to deforestation, the Puerto Rican amazon is one of the world's rarest birds despite conservation efforts.^[150]

 

A mounted specimen of the Carolina parakeet, which was hunted to extinction

One of the largest parrot conservation groups is the World Parrot Trust,^[151] an international organisation. The group gives assistance to worthwhile projects, as well as producing a magazine (PsittaScene)^[152] and raising funds through donations and memberships, often from pet parrot owners. On a smaller scale, local parrot clubs raise money to donate to a conservation cause. Zoo and wildlife centres usually provide public education, to change habits that cause damage to wild populations. Conservation measures to conserve the habitats of some of the high-profile charismatic parrot species has also protected many of the less charismatic species living in the ecosystem.^[153]: 12 A popular attraction that many zoos employ is a feeding station for lories and lorikeets, where visitors feed them with cups of liquid food. This is usually done in association with educational signs and lectures.^[154] Birdwatching-based ecotourism can be beneficial to economies.^[155]

Several projects aimed specifically at parrot conservation have met with success. Translocation of vulnerable kakapo, followed by intensive management and supplementary feeding, has increased the population from 50 individuals to 123 in 2010.^[156] In New Caledonia, the Ouvea parakeet was threatened by trapping for the pet trade and loss of habitat. Community-based conservation, which eliminated the threat of poaching, has allowed the population to increase from around 600 birds in 1993 to over 2000 birds in 2009.^[157]

As of 2009, the IUCN recognises 19 species of parrot as extinct since 1500 (the date used to denote modern extinctions).^[158] This does not include species like the New Caledonian lorikeet, which has not been officially seen for 100 years, yet is still listed as critically endangered.^[159]

Trade, export, and import of all wild-caught parrots is regulated and only permitted under special licensed circumstances in countries party to the Convention on the International Trade in Endangered Species (CITES) which came into force in 1975 to regulate the international trade of all endangered, wild-caught animal and plant species. In 1975, 24 parrot species were included on Appendix I, thus prohibiting commercial international trade in these birds. Since that initial listing, continuing threats from international trade led it to add an additional 32 parrot varieties to Appendix I.^[160] All other parrot species, aside from the rosy-faced lovebird, budgerigar, cockatiel and rose-ringed parakeet (which are not included in the appendices) are protected on Appendix II of CITES.^[161][162] In addition, individual countries may have laws to regulate trade in certain species; for example, the EU has banned parrot trade,^[117] whereas Mexico has a licensing system for capturing parrots.^[119]

World Parrot Day

Every year on 31 May, World Parrot Day is celebrated.^[163]

See also

•  Birds portal

• List of parrots
• Parrots of New Zealand
• Parrots of New Guinea

References

1. . American Heritage Dictionary of the English Language, Fourth Edition. Houghton Mifflin Company. 2000. Archived from the original on 27 August 2007. Retrieved 9 September 2007.
2. "Psittacine". Merriam-Webster Online Dictionary. Merriam-Webster, Inc. Retrieved 9 September 2007.
3. Gill, F.; Donsker, D.; Rasmussen, P. (eds.). "Order Psittaciformes". IOC World Bird List. International Ornithological Congress. Retrieved 19 November 2020.
4. Olah, George; Butchart, Stuart H. M.; Symes, Andy; Guzmán, Iliana Medina; Cunningham, Ross; Brightsmith, Donald J.; Heinsohn, Robert (2016). "Ecological and socio-economic factors affecting extinction risk in parrots" (PDF). Biodiversity and Conservation. 25 (2): 205–223. doi:10.1007/s10531-015-1036-z. ISSN 0960-3115. S2CID 17282072.
5. Berkunsky, I.; Quillfeldt, P.; Brightsmith, D.J.; Abbud, M.C.; Aguilar, J.M.R.E.; Alemán-Zelaya, U.; Aramburú, R.M.; Arce Arias, A.; Balas McNab, R.; Balsby, T.J.S.; Barredo Barberena, J.M.; Beissinger, S.R.; Rosales, M.; Berg, K.S.; Bianchi, C.A. (2017). "Current threats faced by Neotropical parrot populations". Biological Conservation. 214: 278–287. doi:10.1016/j.biocon.2017.08.016.
6. Olah, George; Theuerkauf, Jörn; Legault, Andrew; Gula, Roman; Stein, John; Butchart, Stuart; O’Brien, Mark; Heinsohn, Robert (2018-01-02). "Parrots of Oceania – a comparative study of extinction risk" (PDF). Emu - Austral Ornithology. 118 (1): 94–112. doi:10.1080/01584197.2017.1410066. ISSN 0158-4197. S2CID 135275510.
7. Mellor, Emma L.; McDonald Kinkaid, Heather K.; Mendl, Michael T.; Cuthill, Innes C.; van Zeeland, Yvonne R. A.; Mason, Georgia J. (6 October 2021). "Nature calls: intelligence and natural foraging style predict poor welfare in captive parrots". Proceedings of the Royal Society B: Biological Sciences. 288 (1960). doi:10.1098/rspb.2021.1952. PMC 8493207. PMID 34610768.
8. Melody W. Young, Edwin Dickinson, Nicholas D. Flaim and Michael C. Granatosky (2022). Overcoming a ‘forbidden phenotype’: the parrot’s head supports, propels and powers tripedal locomotion, Proceedings of the Royal Society B: Biological Sciences, 20220245, https://doi.org/10.1098/rspb.2022.0245
9. Wright, T.F.; Schirtzinger E. E.; Matsumoto T.; Eberhard J. R.; Graves G. R.; Sanchez J. J.; Capelli S.; Muller H.; Scharpegge J.; Chambers G. K.; Fleischer R. C. (2008). "A Multilocus Molecular Phylogeny of the Parrots (Psittaciformes): Support for a Gondwanan Origin during the Cretaceous". Molecular Biology and Evolution. 25 (10): 2141–2156. doi:10.1093/molbev/msn160. PMC 2727385. PMID 18653733.
10. Mayr, G. (2009). Paleogene fossil birds. Springer.
11. Tavares ES, Baker AJ, Pereira SL, Miyaki CY (2006). "Phylogenetic relationships and historical biogeography of neotropical parrots (Psittaciformes: Psittacidae: Arini) inferred from mitochondrial and nuclear DNA sequences". Systematic Biology. 55 (3): 454–470. doi:10.1080/10635150600697390. PMID 16861209.
12. Stidham, T. (1998). "A lower jaw from a Cretaceous parrot" (PDF). Nature. 396 (6706): 29–30. Bibcode:1998Natur.396...29S. doi:10.1038/23841. S2CID 204995638.
13. Dyke, G. J.; Mayr, G. (1999). "Did parrots exist in the Cretaceous period?". Nature. 399 (6734): 317–318. Bibcode:1999Natur.399..317D. doi:10.1038/20583. S2CID 204993284.
14. Waterhouse, D. M. (2006). "Parrots in a nutshell: The fossil record of Psittaciformes (Aves)". Historical Biology. 18 (2): 227–238. doi:10.1080/08912960600641224. S2CID 83664072.
15. Ksepka, Daniel T.; Clarke, Julia A.; Grande, Lance (2011). "Stem Parrots (Aves, Halcyornithidae) from the Green River Formation and a Combined Phylogeny of Pan-Psittaciformes". Journal of Paleontology. 85 (5): 835–852. doi:10.1666/10-108.1. ISSN 1937-2337. S2CID 86618579.
16. Suh A, Paus M, Kiefmann M; Paus; Kiefmann; Churakov; Franke; Brosius; Kriegs; Schmitz (2011). "Mesozoic retroposons reveal parrots as the closest living relatives of passerine birds". Nature Communications. 2 (8): 443–8. Bibcode:2011NatCo...2..443S. doi:10.1038/ncomms1448. PMC 3265382. PMID 21863010.{{cite journal}}: CS1 maint: multiple names: authors list (link)
17. Waterhouse, D.M.; Lindow, B.E.K.; Zelenkov, N.; Dyke, G.J. (2008). "Two new fossil parrots (Psittaciformes) from the Lower Eocene Fur Formation of Denmark" (PDF). Palaeontology. 51 (3): 575–582. doi:10.1111/j.1475-4983.2008.00777.x.
18. Dyke, GJ; Cooper, JH (2000). "A new psittaciform bird from the London clay (Lower Eocene) of England". Palaeontology. 43 (2): 271–285. doi:10.1111/1475-4983.00126.
19. Ksepka, Daniel T.; Clarke, Julia A.; Grande, Lance (2011). "Stem Parrots (Aves, Halcyornithidae) from the Green River Formation and a Combined Phylogeny of Pan-Psittaciformes". Journal of Paleontology. 85 (5): 835–852. doi:10.1666/10-108.1. ISSN 1937-2337. S2CID 86618579.
20. Mayr, Gerald (2011). "Well-preserved new skeleton of the Middle Eocene Messelastursubstantiates sister group relationship between Messelasturidae and Halcyornithidae (Aves, ?Pan-Psittaciformes)". Journal of Systematic Palaeontology. 9: 159–171. doi:10.1080/14772019.2010.505252. S2CID 84569222.
21. Cameron 2007, p. 43.
22. Toft, Catherine A.; Wright, Timothy F. (2015). Parrots of the Wild: A Natural History of the World's Most Captivating Birds. Oakland, California: University of California Press. p. 14. ISBN 978-0-520-23925-8.
23. Ball, V. (1889). "Further Notes on the Identification of the Animals and Plants of India, Which Were Known to Early Greek Authors". Proceedings of the Royal Irish Academy. 1: 1–9. JSTOR 20503820.
24. Schrader, O. (1890). Prehistoric antiquities of the Aryan Peoples: A manual of comparative philology and the earliest culture. Being the "Sprachvergleichung und Urgeschichte" translated by Frank Byron Jevons. London: Charles Griffin and Co. p. 270.
25. de Kloet, RS; de Kloet SR (2005). "The evolution of the spindlin gene in birds: Sequence analysis of an intron of the spindlin W and Z gene reveals four major divisions of the Psittaciformes". Molecular Phylogenetics and Evolution. 36 (3): 706–721. doi:10.1016/j.ympev.2005.03.013. PMID 16099384.
26. Tokita, M; Kiyoshi T; Armstrong KN (2007). "Evolution of craniofacial novelty in parrots through developmental modularity and heterochrony". Evolution and Development. 9 (6): 590–601. doi:10.1111/j.1525-142X.2007.00199.x. PMID 17976055. S2CID 46659963.
27. Joseph, Leo; et al. (2012). "A revised nomenclature and classification for family-group taxa of parrots (Psittaciformes)" (PDF). Zootaxa. 3205: 26–40. doi:10.11646/zootaxa.3205.1.2.
28. Burtt, E. H.; Schroeder, M. R.; Smith, L. A.; Sroka, J. E.; McGraw, K. J. (2010). "Colourful parrot feathers resist bacterial degradation". Biology Letters. 7 (2): 214–216. doi:10.1098/rsbl.2010.0716. PMC 3061162. PMID 20926430.
29. Forshaw, Joseph M.; Cooper, William T. (1978) [1973]. Parrots of the World (2nd ed.). Melbourne, Australia: Landsdowne Editions. ISBN 978-0-7018-0690-3.
30. "Zoological Nomenclature Resource: Psittaciformes (Version 9.013)". zoonomen.net. 29 December 2008.
31. White, Nicole E.; et al. (2011). "The evolutionary history of cockatoos (Aves: Psittaciformes: Cacatuidae)". Molecular Phylogenetics and Evolution. 59 (3): 615–622. doi:10.1016/j.ympev.2011.03.011. PMID 21419232.
32. Schweizer, Manuel; Seehausen, Ole & Hertwig, Stefan T. (2011). "Macroevolutionary patterns in the diversification of parrots: effects of climate change, geological events and key innovations". Journal of Biogeography. 38 (11): 2176–2194. doi:10.1111/j.1365-2699.2011.02555.x. S2CID 85625053.
33. Joseph, Leo; Toon, Alicia; Schirtzinger, Erin E. & Wright, Timothy F. (2011). "Molecular systematics of two enigmatic genera Psittacella and Pezoporus illuminate the ecological radiation of Australo-Papuan parrots (Aves: Psittaciformes)". Molecular Phylogenetics and Evolution. 59 (3): 675–684. doi:10.1016/j.ympev.2011.03.017. PMID 21453777.
34. Schweizer, M.; Seehausen O; Güntert M; Hertwig ST (2009). "The evolutionary diversification of parrots supports a taxon pulse model with multiple trans-oceanic dispersal events and local radiations". Molecular Phylogenetics and Evolution. 54 (3): 984–94. doi:10.1016/j.ympev.2009.08.021. PMID 19699808. S2CID 1831016.
35. Forshaw, Joseph M. (2006). Parrots of the World; an Identification Guide. Princeton University Press. p. 70. ISBN 978-0-691-09251-5.
36. Dunning, John B. Jr., ed. (2008). CRC Handbook of Avian Body Masses (2 ed.). CRC Press. ISBN 978-1-4200-6444-5.
37. Sweat, Rebecca. . Archived from the original on 15 July 2016. Retrieved 8 August 2016.{{cite web}}: CS1 maint: unfit URL (link)
38. Demery, Zoe P.; Chappell, J.; Martin, G. R. (2011). "Vision, touch and object manipulation in Senegal parrots Poicephalus senegalus". Proceedings of the Royal Society B. 278 (1725): 3687–3693. doi:10.1098/rspb.2011.0374. PMC 3203496. PMID 21525059.
39. Ödeen, A.; Håstad, O. (2013). "The phylogenetic distribution of ultraviolet sensitivity in birds". BMC Evol Biol. 13 (36): 36. doi:10.1186/1471-2148-13-36. PMC 3637589. PMID 23394614.
40. Brennand, Emma (2 February 2011). "Parrots prefer 'left handedness'". BBC Earth News. Retrieved 5 February 2011.
41. Cameron 2007, p. 57.
42. Cameron 2007, p. 1.
43. Santos, Susana I. C O.; Elward, Brian; Lumeij, Johannes T. (1 March 2006). "Sexual Dichromatism in the Blue-fronted Amazon Parrot (Amazona aestiva) Revealed by Multiple-angle Spectrometry". Journal of Avian Medicine and Surgery. 20 (1): 8–14. doi:10.1647/1082-6742(2006)20[8:SDITBA]2.0.CO;2. S2CID 17124101.
44. Masello, J. F.; Lubjuhn, T.; Quillfeldt, P. (2009). . Hornero. 24 (1): 47–55. ISSN 1850-4884. Archived from the original on 20 June 2018. Retrieved 8 September 2019.
45. Martin, Rowan O; Perrin, Michael R; Boyes, Rutledge S; Abebe, Yilma D; Annorbah, Nathaniel D; Asamoah, Augustus; Bizimana, Dieudonné; Bobo, Kadiri S; Bunbury, Nancy; Brouwer, Joost; Diop, Moussa S; Ewnetu, Mihret; Fotso, Roger C; Garteh, Jerry; Hall, Philip (2014-09-02). "Research and conservation of the larger parrots of Africa and Madagascar: a review of knowledge gaps and opportunities". Ostrich. 85 (3): 205–233. doi:10.2989/00306525.2014.948943. ISSN 0030-6525. S2CID 55821856.
46. Cooke, Fred; Bruce, Jenni (2004). The Encyclopedia of Animals: a complete visual guide (1 ed.). Berkeley, California: University of California Press. p. 296. ISBN 978-0-520-24406-1.
47. Bradford, Alina (2014). "Parrot Facts: Habits, Habitats and Species". LiveScience. Retrieved 7 August 2016.
48. Forshaw, Joseph M. (1989). Parrots of the World (3 ed.). London: Blandford Press. ISBN 978-0-7137-2134-8.
49. . BBC Nature. 2014. Archived from the original on 25 December 2017. Retrieved 7 August 2016.
50. Bonaparte, C. L. (1850). Conspectus Generum Avium (in Latin). Lugduni Batavorum.
51. Heads, Michael (2012). Molecular Panbiogeography of the Tropics. Vol. 4. Berkeley, California: University of California Press. p. 296. ISBN 978-0-520-27196-8.
52. Steadman, D. (2006). Extinction and Biogeography in Tropical Pacific Birds. University of Chicago Press. pp. 342–351. ISBN 978-0-226-77142-7.
53. Cameron 2007, p. 86.
54. Steve Baldwin. . BrooklynParrots.com. Archived from the original on 24 February 2013. Retrieved 27 February 2013.
55. Coughlan, Sean (6 July 2004). "Wild parrots settle in suburbs". BBC News.
56. Mayer-Hohdahl, Alexandra (22 August 2015). "Exotic parakeets set hearts, dread aflutter in Brussels". Gulf Times. Retrieved 23 March 2018.
57. Butler, C (2005). "Feral Parrots in the Continental United States and United Kingdom: Past, Present, and Future". Journal of Avian Medicine and Surgery. 19 (2): 142–149. doi:10.1647/183. S2CID 86275800.
58. Sol, Daniel; Santos, David M.; Feria, Elías; Clavell, Jordi (1997). "Habitat Selection by the Monk Parakeet during Colonization of a New Area in Spain". Condor. 99 (1): 39–46. doi:10.2307/1370222. JSTOR 1370222. S2CID 88376216.
59. Kalodimos, Nicholas P. (2013). "First Account of a Nesting Population of Monk Parakeets, Myiopsitta monachus With Nodule-shaped Bill Lesions in Katehaki, Athens, Greece" (PDF). Bird Populations. 12 (1–6).
60. "Red-crowned Parrot". American Bird Conservancy. Retrieved 6 January 2019.
61. Benham, W. B. (1906). "Notes on the Flesh-eating Propensity of the Kea (Nestor notabilis)". Transactions of the Royal Society of New Zealand. 39: 71–89.
62. Collar, N. (1997). "Family Psittacidae (Parrots)". In del Hoyo, J.; Elliott, A.; Sargatal, J. (eds.). Handbook of the Birds of the World. Vol. Sandgrouse to Cuckoos. Barcelona: Lynx Editions. ISBN 978-84-87334-22-1.
63. Juniper, Tony; Mike Parr (2010). Parrots: A Guide to Parrots of the World. A & C Black. p. 21. ISBN 978-1-4081-3575-4.
64. Diamond, J (1999). "Evolutionary biology: Dirty eating for healthy living". Nature. 400 (6740): 120–121. Bibcode:1999Natur.400..120D. doi:10.1038/22014. PMID 10408435. S2CID 4404459.
65. Bendavidez, A.; Palacio, F.X.; Rivera, L.O.; Echevarria, A.L.; Politi, N. (2018). "Diet of Neotropical parrots is independent of phylogeny but correlates with body size and geographical range". Ibis. 160 (4): 742–754. doi:10.1111/ibi.12630.
66. Gartrell, B; Jones, S; Brereton, R; Astheimer, L (2000). "Morphological Adaptations to Nectarivory of the Alimentary Tract of the Swift Parrot Lathamus discolor". Emu. 100 (4): 274–279. doi:10.1071/MU9916. S2CID 86013058.
67. Schweizer, Manuel; Güntert, Marcel; Seehausen, Ole; Leuenberger, Christoph; Hertwig, Stefan T. (2014). "Parallel adaptations to nectarivory in parrots, key innovations and the diversification of the Loriinae". Ecology and Evolution. 4 (14): 2867–2883. doi:10.1002/ece3.1131. PMC 4130445. PMID 25165525.
68. "Golden-winged Parakeet (Brotogeris chrysoptera)". World Parrot Trust. Retrieved 8 August 2016.
69. Jackson, J. R. (1962). "Do Keas Attack Sheep?" (PDF). Notornis. 10 (1).
70. Greene, Terry (November–December 1999). (PDF). Notornis. Ornithological Society of New Zealand. 46 (2): 301–310. Archived from the original (PDF) on 2021-03-03. Retrieved 2011-09-29.
71. Cameron 2007, p. 114–116.
72. Rowley, I. (1997). "Family Cacatuidae (Cockatoos)". In del Hoyo, J.; Elliott, A.; Sargatal, J. (eds.). Handbook of the Birds of the World. Vol. 4. Barcelona: Lynx Editions. ISBN 978-84-87334-22-1.
73. Oren, David C.; Novaes, Fernando (1986). "Observations on the golden parakeet Aratinga guarouba in Northern Brazil". Biological Conservation. 36 (4): 329–337. doi:10.1016/0006-3207(86)90008-X.
74. Eberhard, J (1998). "Evolution of nest-building behavior in Agapornis parrots" (PDF). Auk. 115 (2): 455–464. doi:10.2307/4089204. JSTOR 4089204.
75. Sanchez-Martinez, Tania; Katherine Renton (2009). "Availability and selection of arboreal termitaria as nest-sites by Orange-fronted Parakeets Aratinga canicularis in conserved and modified landscapes in Mexico". Ibis. 151 (2): 311–320. doi:10.1111/j.1474-919X.2009.00911.x.
76. Heinsohn, Robert; Murphy, Stephen; Legge, Sarah (2003). "Overlap and competition for nest holes among eclectus parrots, palm cockatoos and sulphur-crested cockatoos". Australian Journal of Zoology. 51 (1): 81–94. doi:10.1071/ZO02003. S2CID 83711585.
77. Pell, A; Tidemann, C (1997). "The impact of two exotic hollow-nesting birds on two native parrots in savannah and woodland in eastern Australia". Biological Conservation. 79 (2): 145–153. doi:10.1016/S0006-3207(96)00112-7.
78. "Man-made nesting hollows big hit with endangered swift parrots". ABC News. 21 October 2016.
79. Masello, J; Pagnossin, M; Sommer, C; Quillfeldt, P (2006). "Population size, provisioning frequency, flock size and foraging range at the largest known colony of Psittaciformes: the Burrowing Parrots of the north-eastern Patagonian coastal cliffs" (PDF). Emu. 106 (1): 69–79. doi:10.1071/MU04047. S2CID 59270301.
80. Eberhard, Jessica (2002). "Cavity adoption and the evolution of coloniality in cavity-nesting birds". Condor. 104 (2): 240–247. doi:10.1650/0010-5422(2002)104[0240:CAATEO]2.0.CO;2. ISSN 0010-5422. S2CID 59455644.
81. Forshaw, Joseph (1991). Forshaw, Joseph (ed.). Encyclopaedia of Animals: Birds. London: Merehurst Press. pp. 118–124. ISBN 978-1-85391-186-6.
82. Tweti, Mira (2008). Of Parrots and People (1 ed.). New York: Viking Penguin. ISBN 978-0-14-311575-5.
83. Iwaniuk, A. N.; Nelson, J. E. (2003). "Developmental differences are correlated with relative brain size in birds: a comparative analysis". Canadian Journal of Zoology. 81 (12): 1913–1928. doi:10.1139/z03-190.
84. Emery, Nathan J. (2006). "Cognitive ornithology: the evolution of avian intelligence". Philosophical Transactions of the Royal Society B: Biological Sciences. 361 (1465): 23–43. doi:10.1098/rstb.2005.1736. PMC 1626540. PMID 16553307.
85. Beynon, Mike (April 2000). . BBC News. Archived from the original on 1 September 2007. Retrieved 9 September 2007.
86. Cussen, Victoria A.; Mench, Joy A. (2015). Wicker-Thomas, Claude (ed.). "The Relationship between Personality Dimensions and Resiliency to Environmental Stress in Orange-Winged Amazon Parrots (Amazona amazonica), as Indicated by the Development of Abnormal Behaviors". PLOS ONE. 10 (6): e0126170. Bibcode:2015PLoSO..1026170C. doi:10.1371/journal.pone.0126170. PMC 4482636. PMID 26114423.
87. Pepperberg, Irene M. (1988). "Comprehension of "Absence" by an African Grey Parrot: Learning with Respect to Questions of Same/Different". Journal of the Experimental Analysis of Behavior. 50 (3): 553–564. doi:10.1901/jeab.1988.50-553. PMC 1338917. PMID 16812572.
88. Dunayer, Joan (2013). Corbey, Raymond; Lanjouw, Annette (eds.). The Politics of Species: Reshaping our Relationships with Other Animals. New York: Cambridge University Press. p. 33. ISBN 978-1-107-03260-6.
89. Patterson, Dianne K.; Pepperberg, Irene M.; Story, Brad H.; Hoffman, Eric A. (1997). "How parrots talk: Insights based on CT scans, image processing and mathematical models". Proceedings of the International Society for Optical Engineering. Medical Imaging 1997: Physiology and Function from Multidimensional Images. 3033: 14–24. Bibcode:1997SPIE.3033...14P. doi:10.1117/12.274039. S2CID 108771414.
90. BirdLife International (2018). "Psittacus erithacus". IUCN Red List of Threatened Species. 2018: e.T22724813A129879439. doi:10.2305/IUCN.UK.2018-2.RLTS.T22724813A129879439.en. Retrieved 12 November 2021.
91. Cruickshank, A; Gautier, J; Chappuis, C (1993). "Vocal mimicry in wild African Grey Parrots Psittacus erithacus". Ibis. 135 (3): 293–299. doi:10.1111/j.1474-919X.1993.tb02846.x.
92. Gaunt, Abbot S. (1 October 1983). "An Hypothesis concerning the Relationship of Syringeal Structure to Vocal Abilities". The Auk. 100 (4): 853–862. doi:10.1093/auk/100.4.853. ISSN 0004-8038.
93. SUTHERS, RODERICK A.; ZOLLINGER, SUE ANNE (June 2004). "Producing Song: The Vocal Apparatus". Annals of the New York Academy of Sciences. 1016 (1): 109–129. Bibcode:2004NYASA1016..109S. doi:10.1196/annals.1298.041. ISSN 0077-8923. PMID 15313772. S2CID 45809019.
94. Péron, F.; Rat-Fischer, L.; Lalot, M.; Nagle, L.; Bovet, D. (2011). "Cooperative problem solving in African grey parrots (Psittacus erithacus)". Animal Cognition. 14 (4): 545–553. doi:10.1007/s10071-011-0389-2. PMID 21384141. S2CID 5616569.
95. Gill, Victoria (18 May 2011). . BBC Nature. Archived from the original on 20 May 2011. Retrieved 3 July 2016.
96. "Choosing a New Pet Bird". Petco. Retrieved 1 August 2016.
97. Grindol, Diane (2013). "Budgie Vs. Cockatiel – Which Right For You?". Retrieved 1 August 2016.
98. Ward, Sam (21 December 1992). . USA Today. Archived from the original on 28 July 2013. Retrieved 6 September 2009.
99. "Parrot". The Medieval Bestiary. 2011. Retrieved 1 August 2016.
100. de Grahl, Wolfgang (1989). The Grey Parrot. Tfh Publications Incorporated. ISBN 978-0-86622-094-1.
101. Sweat, Rebecca (March 2015). "Foods such as alcohol, chocolate, avocado, uncooked meats and shellfish can be bad for your pet bird". BirdChannel.com. Retrieved 12 July 2016.
102. "Pet Birds, Parrots, Cockatiels, Macaws, Conures, Parakeets Care". BirdChannel. Retrieved 1 August 2016.
103. The National Parrot Sanctuary. . parrotsanctuary.co.uk. Archived from the original on 19 June 2008.
104. Noeth, Gay; Johnson, Sibylle. "Biting: The Sharp Beak". Avian Web: Beauty of Birds. Retrieved 1 August 2016.
105. Patt, Jill M. "Bird Behavior and Training". Avian Web: Beauty of Birds. Retrieved 1 August 2016.
106. Leisure, Susan. "How Long can a Parrot Live?". The nest. Retrieved 5 August 2016.
107. "Winston's obscene parrot lives on". BBC News. 2004. Retrieved 5 August 2016.
108. Coila, Bridget. "The Average Lifespan of Parrots". The nest. Retrieved 1 August 2016.
109. "The Truth About Parrots as Pets". In Defense of Animals. Retrieved 1 August 2016.
110. Kelly, Denise; Rae, Joan; Menzel, Krista (2007). "The True Nature of Parrots". Avian Welfare Coalition. Retrieved 1 August 2016.
111. BirdLife International (2016). "Anodorhynchus hyacinthinus". IUCN Red List of Threatened Species. 2016: e.T22685516A93077457. doi:10.2305/IUCN.UK.2016-3.RLTS.T22685516A93077457.en. Retrieved 12 November 2021.
112. Munn, Charles A.; Thomsen, Jorgen B.; Yamashita, Carlos (2013). Chandler, William J.; Wille, Chris; Labate, Lillian (eds.). Audubon Wildlife Report, 1988–1989. Academic Press. pp. 405–419. ISBN 978-0-12-041001-9.
113. Vallery, Anna (January 2015). "The Truth About the Exotic Bird Trade Will Make You Rethink Buying a Parrot in the Pet Shop". One Green Planet. Retrieved 5 August 2016.
114. Lowther, Jason; Cook, Dee; Roberts, Martin (5 August 2002). (PDF). World Wildlife Federation. Archived from the original (PDF) on 10 September 2008. Retrieved 9 September 2007.
115. Soucek, Gayle (2008). Gouldian Finches. Barron's Educational Series. p. 7. ISBN 978-0-7641-3850-8.
116. "New rules for captive bird imports to protect animal health in the EU and improve the welfare of imported birds". European Commission: Press Release Database. 2015. Retrieved 6 August 2016.
117. . Defenders of Wildlife. January 2007. Archived from the original on 13 April 2019. Retrieved 5 August 2016.
118. "Naturalized Parrots in the U.S." Avian Welfare Coalition. Retrieved 9 August 2016.
119. Guzman, Juan C. C.; Sanchez, Saldana M. E.; Grosselet, Manuel; Gomez, Jesus S. (2007). The Illegal Parrot Trade in Mexico (Report). Mexico: Defenders of Wildlife. pp. 9–10.
120. "Stopping the Illegal Mexican Parrot Trade" (PDF). Defenders of Wildlife. Retrieved 23 December 2007.
121. "The Cat and the Parrot". Fairy Tales and Other Traditional Stories. 2012. Retrieved 9 August 2016.
122. Gamard, Ibrahim. "The Merchant and the Parrot". Dar-Al Masnavi. Retrieved 9 August 2016.
123. Boehrer, Bruce (2004). Parrot Culture. Philadelphia: University of Pennsylvania Press. ISBN 978-0-8122-3793-1.
124. . BirdLife International. 2008. Archived from the original on 3 May 2016. Retrieved 9 August 2016.
125. Chamberlain, Susan (2013). "Parrot History: Yesterday and Today". BirdChannel. Retrieved 9 August 2016.
126. "Dominica Flags and Symbols and National Anthem". World Atlas. Retrieved 9 August 2016.
127. "The National Bird". The Government of St. Vincent and the Grenadines Ministry of Foreign Affairs, Trade and Commerce. Retrieved 5 August 2016.
128. Quinion, Michael (10 June 2000). "Sick as a." World Wide Words. Retrieved 13 October 2011.
129. Swithenbank, G. . Archived from the original on 30 June 2018. Retrieved 13 October 2011.
130. Behn, Aphra (1681). "The False Count". Project Gutenberg. Retrieved 13 January 2018. 'Lord, Madam, you are as melancholy as a sick Parrot.' 'And can you blame me, Jacinta? have I not many Reasons to be sad?'
131. . Parrot Heads of North Carolina. Archived from the original on 21 June 2016. Retrieved 3 July 2016.
132. "PsittaScene Magazine". World Parrot Trust. Retrieved 3 July 2016.
133. "The Dead Parrot sketch". Monty Python. Season 1. Episode 8. United Kingdom. December 1969. 5:33 minutes in. BBC1.
134. Gosnell, Raja (Director) (1997). Home Alone 3 (Motion picture). 20th Century Fox. Archived from the original on 30 October 2021.
135. "Rio the Movie Synopsis". Fandango. 2011. Retrieved 9 August 2016.
136. "The Wild Parrots of Telegraph Hill". Internet Movie Database. 2005. Retrieved 9 August 2016.
137. Craig, Robert D. (1989). Dictionary of Polynesian mythology. Greenwood Publishing Group. p. 6. ISBN 978-0-313-25890-9.
138. Handy, E. S. C. (1930). Marquesan Legends. Honolulu: Bernice P. Bishop Museum Press. pp. 130–1.
139. Katherine, Berrin; Museum, Larco (1997). The Spirit of Ancient Peru: Treasures from the Museo Arqueológico Rafael Larco Herrera. New York: Thames and Hudson. ISBN 978-0-500-01802-6.
140. Chung Tai World (2009). "The Parrot who Extinguished a Forest Fire". Chung Tai Monastery. Retrieved 6 August 2016.
141. Idema, Wilt L. (2008). Personal salvation and filial piety: two precious scroll narratives of Guanyin and her acolytes. University of Hawaii Press. p. 33. ISBN 978-0-8248-3215-5.
142. Department of Conservation (2008). . Archived from the original on 2 December 2013. Retrieved 14 July 2008.
143. Toft, Catherine A.; Wright, Timothy F. (2015). "Parrots as Invasive Species". Parrots of the Wild: A Natural History of the World's Most Captivating Birds (1 ed.). Oakland, California: University of California Press. p. 248. ISBN 978-0-520-96264-4.
144. Butler, Christopher J. (2005). "Feral Parrots in the Continental United States and United Kingdom: Past, Present, and Future". Journal of Avian Medicine and Surgery. 19 (2): 142–149. doi:10.1647/183. S2CID 86275800.
145. Lepore, Jill (1 June 2009). "It's Spreading". The New Yorker. pp. 27–29. Retrieved 13 September 2011.
146. BirdLife International (2016). "Nestor productus". IUCN Red List of Threatened Species. 2016: e.T22684834A93049105. doi:10.2305/IUCN.UK.2016-3.RLTS.T22684834A93049105.en. Retrieved 12 November 2021.
147. Campbell, T. S. (December 2000). . The Institute for Biological Invasions. Archived from the original on 1 July 2007.
148. Moorhouse, Ron; Greene, Terry; Dilks, Peter; Powlesland, Ralph; Moran, Les; Taylor, Genevieve; Jones, Alan; Knegtmans, Jaap; Wills, Dave; Pryde, Moira; Fraser, Ian; August, Andrew; August, Claude (2002). "Control of introduced mammalian predators improves kaka Nestor meridionalis breeding success: reversing the decline of a threatened New Zealand parrot". Biological Conservation. 110 (1): 33–44. doi:10.1016/S0006-3207(02)00173-8.
149. White, Thomas H. Jr.; Collazo, Jaime A.; Vilella, Francisco J.; Guerrero, Simón A. (2005). "Effects of Hurricane Georges on Habitat Used by Captive-Reared Hispaniola Parrots (Amazona ventralis) Released in the Dominican Republic" (PDF). Ornitologia Neotropical. 16: 405–417.
150. "Natural Resources – Endangered and Threatened Species". USDA. Retrieved 21 August 2013.
151. "History & Accomplishments". The World Parrot Trust. Retrieved 6 January 2019.
152. "Our publications: PsittaScene Magazine". World Parrot Trust.
153. Snyder, N.; McGowan, P.; Gilardi, J.; Grajal, A. (2000). Parrots: Status Survey and Conservation Action Plan, 2000–2004. International Union for Conservation of Nature. ISBN 978-2-8317-0504-0.
154. "Rainbow Lorikeet". Beauty of Birds. 2011. Retrieved 9 August 2016.
155. Christian, C.; Potts, T.; Burnett, G.; Lacher, T. (1999). "Parrot Conservation and Ecotourism in the Windward Islands". Journal of Biogeography. 23 (3): 387–393. doi:10.1046/j.1365-2699.1996.00041.x. S2CID 59582818.
156. Kakapo Recovery Programme (2010). "Then and Now". Kakapo Recovery Programme. Retrieved 1 April 2010.
157. Barré, Nicholas; Theuerkauf, Jörn; Verfaille, Ludovic; Primot, Pierre; Saoumoé, Maurice (2010). "Exponential population increase in the endangered Ouvéa Parakeet (Eunymphicus uvaeensis) after community-based protection from nest poaching" (PDF). Journal of Ornithology. 151 (3): 695–701. doi:10.1007/s10336-010-0499-7. S2CID 20411641.
158. Cooney, Stuart J. N. (2009). Ecological Association of the Hooded Parrot (Psephotus dissimilis) (PDF) (Thesis). Australian National University. p. 13.
159. BirdLife International (2019). "Charmosyna diadema". IUCN Red List of Threatened Species. 2019: e.T22684689A156512185. doi:10.2305/IUCN.UK.2019-3.RLTS.T22684689A156512185.en. Retrieved 12 November 2021.
160. A group of 226 non-governmental organisations (19 May 2005). (PDF). birdsareforwatching.org. Archived from the original (PDF) on 11 April 2016. Retrieved 14 September 2007.
161. "Appendices I, II and III valid from 22 June 2021". CITES. Retrieved 7 March 2022.
162. . American Federation of Aviculture. Archived from the original on 14 April 2017. Retrieved 6 August 2016.
163. "World Parrot Day". Northern Parrots. 22 April 2017.

Cited sources

• Cameron, Matt (2007). Cockatoos. Collingwood, VIC, Australia: CSIRO Publishing. ISBN 978-0-643-09232-7.

External links

•   Media related to Psittaciformes at Wikimedia Commons
• Parrot videos on the Internet Bird Collection