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Wikipedia

Triassic

The Triassic (/trˈæsɪk/ try-ASS-ik)[8] is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.36 Mya.[9] The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events.[10] The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.

Triassic
251.902 ± 0.024 – 201.4 ± 0.2 Ma
Chronology
Etymology
Name formalityFormal
Usage information
Celestial bodyEarth
Regional usageGlobal (ICS)
Time scale(s) usedICS Time Scale
Definition
Chronological unitPeriod
Stratigraphic unitSystem
Time span formalityFormal
Lower boundary definitionFirst appearance of the conodont Hindeodus parvus
Lower boundary GSSPMeishan, Zhejiang, China
31°04′47″N 119°42′21″E / 31.0798°N 119.7058°E / 31.0798; 119.7058
Lower GSSP ratified2001[6]
Upper boundary definitionFirst appearance of the ammonite Psiloceras spelae tirolicum
Upper boundary GSSPKuhjoch section, Karwendel mountains, Northern Calcareous Alps, Austria
47°29′02″N 11°31′50″E / 47.4839°N 11.5306°E / 47.4839; 11.5306
Upper GSSP ratified2010[7]

The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period.[11] Archosaurs that became dominant in this period were primarily pseudosuchians, ancestors of modern crocodilians, while some archosaurs specialized in flight, the first time among vertebrates, becoming the pterosaurs.

Therapsids, the dominant vertebrates of the preceding Permian period, declined throughout the period. The first true mammals, themselves a specialized subgroup of therapsids, also evolved during this period. The vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to gradually rift into two separate landmasses, Laurasia to the north and Gondwana to the south.

The global climate during the Triassic was mostly hot and dry,[12] with deserts spanning much of Pangaea's interior. However, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another major mass extinction, the Triassic–Jurassic extinction event, that wiped out many groups, including most pseudosuchians, and allowed dinosaurs to assume dominance in the Jurassic.

Etymology

The Triassic was named in 1834 by Friedrich August von Alberti, after a succession of three distinct rock layers (Greek triás meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone), the middle Muschelkalk (shell-bearing limestone) and the upper Keuper (coloured clay).[13]

Dating and subdivisions

On the geologic time scale, the Triassic is usually divided into Early, Middle, and Late Triassic Epochs, and the corresponding rocks are referred to as Lower, Middle, or Upper Triassic. The faunal stages from the youngest to oldest are:

Series/Epoch Faunal stage Time span
Upper/Late Triassic (Tr3) Rhaetian (208.5 – 201.4 ± 0.2 Mya)
Norian (227 – 208.5 Mya)
Carnian (237 – 227 Mya)
Middle Triassic (Tr2) Ladinian (242 – 237 Mya)
Anisian (247.2 – 242 Mya)
Lower/Early Triassic (Scythian) Olenekian (251.2 – 247.2 Mya)
Induan (251.902 ± 0.024 – 251.2 Mya)

Paleogeography

 
View of the Tethys area during the Ladinian stage
 
230 Ma continental reconstruction

During the Triassic, almost all the Earth's land mass was concentrated into a single supercontinent, Pangaea (lit.'entire land').[14] This supercontinent was more-or-less centered on the equator and extended between the poles, though it did drift northwards as the period progressed. Southern Pangea, also known as Gondwana, was made up by closely-appressed cratons corresponding to modern South America, Africa, Madagascar, India, Antarctica, and Australia. North Pangea, also known as Laurussia or Laurasia, corresponds to modern-day North America and the fragmented predecessors of Eurasia.

The western edge of Pangea lay at the margin of an enormous ocean, Panthalassa (lit. 'entire sea'), which roughly corresponds to the modern Pacific Ocean. Practically all deep-ocean crust present during the Triassic has been recycled through the subduction of oceanic plates, so very little is known about the open ocean from this time period. Most information on Panthalassan geology and marine life is derived from island arcs and rare seafloor sediments accreted onto surrounding land masses, such as present-day Japan and western North America.

The eastern edge of Pangea was encroached upon by a pair of extensive oceanic basins: The Neo-Tethys (or simply Tethys) and Paleo-Tethys Oceans. These extended from China to Iberia, hosting abundant marine life along their shallow tropical peripheries. They were divided from each other by a long string of microcontinents known as the Cimmerian terranes. Cimmerian crust had detached from Gondwana in the early Permian and drifted northwards during the Triassic, enlarging the Neo-Tethys Ocean which formed in their wake. At the same time, they forced the Paleo-Tethys Ocean to shrink as it was being subducted under Asia. By the end of the Triassic, the Paleo-Tethys Ocean occupied a small area and the Cimmerian terranes began to collide with southern Asia. This collision, known as the Cimmerian Orogeny, continued into the Jurassic and Cretaceous to produce a chain of mountain ranges stretching from Turkey to Malaysia.[15][16]

 
Sydney, Australia lies on Triassic shales and sandstones. Almost all of the exposed rocks around Sydney belong to the Triassic Sydney sandstone.[17]

Pangaea was fractured by widespread faulting and rift basins during the Triassic—especially late in that period—but had not yet separated. The first nonmarine sediments in the rift that marks the initial break-up of Pangaea, which separated eastern North America from Morocco, are of Late Triassic age; in the United States, these thick sediments comprise the Newark Supergroup.[18] Rift basins are also common in South America, Europe, and Africa. Terrestrial environments are particularly well-represented in the South Africa,[19] Russia, central Europe, and the southwest United States. Terrestrial Triassic biostratigraphy is mostly based on terrestrial and freshwater tetrapods, as well as conchostracans ("clam shrimps"), a type of fast-breeding crustacean which lived in lakes and hypersaline environments.

Because a supercontinent has less shoreline compared to a series of smaller continents, Triassic marine deposits are relatively uncommon on a global scale. A major exception is in Western Europe, where the Triassic was first studied. The northeastern margin of Gondwana was a stable passive margin along the Neo-Tethys Ocean, and marine sediments have been preserved in parts of northern India and Arabia.[16] In North America, marine deposits are limited to a few exposures in the west.

Scandinavia

During the Triassic peneplains are thought to have formed in what is now Norway and southern Sweden.[20][21][22] Remnants of this peneplain can be traced as a tilted summit accordance in the Swedish West Coast.[20] In northern Norway Triassic peneplains may have been buried in sediments to be then re-exposed as coastal plains called strandflats.[21] Dating of illite clay from a strandflat of Bømlo, southern Norway, have shown that landscape there became weathered in Late Triassic times (c. 210 million years ago) with the landscape likely also being shaped during that time.[23]

Paleooceanography

Eustatic sea level in the Triassic was consistently low compared to the other geological periods. The beginning of the Triassic was around present sea level, rising to about 10–20 metres (33–66 ft) above present-day sea level during the Early and Middle Triassic. Sea level rise accelerated in the Ladinian, culminating with a sea level up to 50 metres (164 ft) above present-day levels during the Carnian. Sea level began to decline in the Norian, reaching a low of 50 metres (164 ft) below present sea level during the mid-Rhaetian. Low global sea levels persisted into the earliest Jurassic. The long-term sea level trend is superimposed by 22 sea level drop events widespread in the geologic record, mostly of minor (less than 25-metre (82 ft)) and medium (25–75-metre (82–246 ft)) magnitudes. A lack of evidence for Triassic continental ice sheets suggest that glacial eustasy is unlikely to be the cause of these changes.[24]

Climate

The Triassic continental interior climate was generally hot and dry, so that typical deposits are red bed sandstones and evaporites. There is no evidence of glaciation at or near either pole; in fact, the polar regions were apparently moist and temperate, providing a climate suitable for forests and vertebrates, including reptiles. Pangaea's large size limited the moderating effect of the global ocean; its continental climate was highly seasonal, with very hot summers and cold winters.[25] The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons,[25] sometimes referred to as megamonsoons.[26]

The Triassic may have mostly been a dry period, but evidence exists that it was punctuated by several episodes of increased rainfall in tropical and subtropical latitudes of the Tethys Sea and its surrounding land.[27] Sediments and fossils suggestive of a more humid climate are known from the Anisian to Ladinian of the Tethysian domain, and from the Carnian and Rhaetian of a larger area that includes also the Boreal domain (e.g., Svalbard Islands), the North American continent, the South China block and Argentina.

The best-studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event. A 2020 study found bubbles of carbon dioxide in basaltic rocks dating back to the end of the Triassic, and concluded that volcanic activity helped trigger climate change in that period.[28]

Flora

 
Triassic flora as depicted in Meyers Konversations-Lexikon (1885–90)

Land plants

On land, the surviving vascular plants included the lycophytes, the dominant cycadophytes, ginkgophyta (represented in modern times by Ginkgo biloba), ferns, horsetails and glossopterids. The spermatophytes, or seed plants, came to dominate the terrestrial flora: in the northern hemisphere, conifers, ferns and bennettitales flourished. The seed fern genus Dicroidium would dominate Gondwana throughout the period.

Coal

 
Immediately above the Permian–Triassic boundary the glossopteris flora was suddenly[29] largely displaced by an Australia-wide coniferous flora.

No known coal deposits date from the start of the Triassic Period. This is known as the Early Triassic "coal gap" and can be seen as part of the Permian–Triassic extinction event.[30] Possible explanations for the coal gap include sharp drops in sea level at the time of the Permo-Triassic boundary;[31] acid rain from the Siberian Traps eruptions or from an impact event that overwhelmed acidic swamps; climate shift to a greenhouse climate that was too hot and dry for peat accumulation; evolution of fungi or herbivores that were more destructive of wetlands; the extinction of all plants adapted to peat swamps, with a hiatus of several million years before new plant species evolved that were adapted to peat swamps;[30] or soil anoxia as oxygen levels plummeted.[32]

Phytoplankton

Before the Permian extinction, Archaeplastida (red and green algae) had been the major marine phytoplanktons since about 659–645 million years ago,[33] when they replaced marine planktonic cyanobacteria, which first appeared about 800 million years ago, as the dominant phytoplankton in the oceans.[34] In the Triassic, secondary endosymbiotic algae became the most important plankton.[35]

Fauna

 
Middle Triassic marginal marine sequence, southwestern Utah
 
Reconstruction of the Triassic amphibian Mastodonsaurus

Marine invertebrates

In marine environments, new modern types of corals appeared in the Early Triassic, forming small patches of reefs of modest extent compared to the great reef systems of Devonian or modern times. Serpulids appeared in the Middle Triassic.[36] Microconchids were abundant. The shelled cephalopods called ammonites recovered, diversifying from a single line that survived the Permian extinction.

Fish

The fish fauna was remarkably uniform, with many families and genera exhibiting a global distribution in the wake of the Permian-Triassic mass extinction event.[37] Ray-finned fishes (actinopterygians) went through a remarkable diversification during the Triassic, leading to peak diversity during the Middle Triassic; however, the pattern of this diversification is still not well understood due to a taphonomic megabias.[38] Large predatory actinopterygians such as saurichthyids and birgeriids appeared in the Early Triassic and became widespread and successful during the period as a whole.[39] Lakes and rivers were populated by lungfish (Dipnoi), such as Ceratodus, which are mainly known from the dental plates, abundant in the fossils record.[40] Hybodonts, a group of shark-like cartilaginous fish, were dominant in both freshwater and marine environments throughout the Triassic.[41]

Amphibians

Temnospondyl amphibians were among those groups that survived the Permian–Triassic extinction. Once abundant in both terrestrial and aquatic environments, the terrestrial species had mostly died out during the extinction event. The Triassic survivors were aquatic or semi-aquatic, and were represented by Tupilakosaurus, Thabanchuia, Branchiosauridae and Micropholis, all of which died out in Early Triassic, and the successful Stereospondyli, with survivors into the Cretaceous Period. The largest Triassic stereospondyls, such as Mastodonsaurus, were up to 4 to 6 metres (13 to 20 ft) in length.[42][43] Some lineages (e.g. trematosaurs) flourished briefly in the Early Triassic, while others (e.g. capitosaurs) remained successful throughout the whole period, or only came to prominence in the Late Triassic (e.g. Plagiosaurus, metoposaurs).

The first Lissamphibians (modern amphibians) appear in the Triassic, with the progenitors of the first frogs already present by the Early Triassic. However, the group as a whole did not become common until the Jurassic, when the temnospondyls had become very rare.

Most of the Reptiliomorpha, stem-amniotes that gave rise to the amniotes, disappeared in the Triassic, but two water-dwelling groups survived: Embolomeri that only survived into the early part of the period, and the Chroniosuchia, which survived until the end of the Triassic.

Reptiles

Archosauromorphs

The Permian–Triassic extinction devastated terrestrial life. Biodiversity rebounded as the surviving species repopulated empty terrain, but these were short-lived. Diverse communities with complex food-web structures took 30 million years to reestablish.[10][44] Archosauromorph reptiles, which had already appeared and diversified to an extent in the Permian Period, exploding in diversity as an adaptive radiation in response to the Permian-Triassic mass extinction. By the Early Triassic, several major archosauromorph groups had appeared. Long-necked, lizard-like early archosauromorphs were known as protorosaurs, which is likely a paraphyletic group rather than a true clade. Tanystropheids were a family of protorosaurs which elevated their neck size to extremes, with the largest genus Tanystropheus having a neck longer than its body. The protorosaur family Sharovipterygidae used their elongated hindlimbs for gliding. Other archosauromorphs, such as rhynchosaurs and allokotosaurs, were mostly stocky-bodied herbivores with specialized jaw structures.

Rhynchosaurs, barrel-gutted herbivores, thrived for only a short period of time, becoming extinct about 220 million years ago. They were exceptionally abundant in the middle of the Triassic, as the primary large herbivores in many Carnian-age ecosystems. They sheared plants with premaxillary beaks and plates along the upper jaw with multiple rows of teeth. Allokotosaurs were iguana-like reptiles, including Trilophosaurus (a common Late Triassic reptile with three-crowned teeth), Teraterpeton (which had a long beak-like snout), and Shringasaurus (a horned herbivore which reached a body length of 3–4 metres (9.8–13.1 ft).

One group of archosauromorphs, the archosauriforms, were distinguished by their active predatory lifestyle, with serrated teeth and upright limb postures. Archosauriforms were diverse in the Triassic, including various terrestrial and semiaquatic predators of all shapes and sizes. The large-headed and robust erythrosuchids were among the dominant carnivores in the early Triassic. Phytosaurs were a particularly common group which prospered during the Late Triassic. These long-snouted and semiaquatic predators resemble living crocodiles and probably had a similar lifestyle, hunting for fish and small reptiles around the water's edge. However, this resemblance is only superficial and is a prime-case of convergent evolution.

True archosaurs appeared in the early Triassic, splitting into two branches: Avemetatarsalia (the ancestors to birds) and Pseudosuchia (the ancestors to crocodilians). Avemetatarsalians were a minor component of their ecosystems, but eventually produced the earliest pterosaurs and dinosaurs in the Late Triassic. Early long-tailed pterosaurs appeared in the Norian and quickly spread worldwide. Triassic dinosaurs evolved in the Carnian and include early sauropodomorphs and theropods. Most Triassic dinosaurs were small predators and only a few were common, such as Coelophysis, which was 1 to 2 metres (3.3 to 6.6 ft) long.

The large predator Smok was most likely also an archosaur, but it is uncertain if it was a primitive dinosaur or a pseudosuchian.

Pseudosuchians were far more ecologically dominant in the Triassic, including large herbivores (such as aetosaurs), large carnivores ("rauisuchians"), and the first crocodylomorphs ("sphenosuchians"). Aetosaurs were heavily-armored reptiles that were common during the last 30 million years of the Late Triassic until they died out at the Triassic-Jurassic extinction. Most aetosaurs were herbivorous and fed on low-growing plants, but some may have eaten meat. "rauisuchians" (formally known as paracrocodylomorphs) were the keystone predators of most Triassic terrestrial ecosystems. Over 25 species have been found, including giant quadrupedal hunters, sleek bipedal omnivores, and lumbering beasts with deep sails on their backs. They probably occupied the large-predator niche later filled by theropods. "Rauisuchians" were ancestral to small, lightly-built crocodylomorphs, the only pseudosuchians which survived into the Jurassic.

Marine reptiles

 
Marine vertebrate apex predators of the Early Triassic and Anisian (Middle Triassic)[45]

There were many types of marine reptiles. These included the Sauropterygia, which featured pachypleurosaurus and nothosaurs (both common during the Middle Triassic, especially in the Tethys region), placodonts, the earliest known herbivorous marine reptile Atopodentatus, and the first plesiosaurs. The first of the lizardlike Thalattosauria (askeptosaurs) and the highly successful ichthyosaurs, which appeared in Early Triassic seas soon diversified, and some eventually developed to huge size during the Late Triassic.

Other reptiles

Among other reptiles, the earliest turtles, like Proganochelys and Proterochersis, appeared during the Norian Age (Stage) of the Late Triassic Period. The Lepidosauromorpha, specifically the Sphenodontia, are first found in the fossil record of the earlier Carnian Age, though the earliest lepidosauromorphs likely occurred in the Permian. The Procolophonidae, the last surviving parareptiles, were an important group of small lizard-like herbivores. The drepanosaurs were a clade of unusual, chameleon-like arboreal reptiles with birdlike heads and specialised claws.

Synapsids

Three therapsid groups survived into the Triassic: dicynodonts, therocephalians, and cynodonts. The cynodont Cynognathus was a characteristic top predator in the Olenekian and Anisian of Gondwana. Both kannemeyeriiform dicynodonts and gomphodont cynodonts remained important herbivores during much of the period. Therocephalians included both large predators (Moschorhinus) and herbivorous forms (bauriids) until their extinction midway through the period. Ecteniniid cynodonts played a role as large-sized, cursorial predators in the Late Triassic. During the Carnian (early part of the Late Triassic), some advanced cynodonts gave rise to the first mammals.

During the Triassic, archosaurs displaced therapsids as the largest and most ecologically prolific terrestrial amniotes. This "Triassic Takeover" may have contributed to the evolution of mammals by forcing the surviving therapsids and their mammaliaform successors to live as small, mainly nocturnal insectivores. Nocturnal life may have forced the mammaliaforms to develop fur and a higher metabolic rate.[46]

Lagerstätten

The Monte San Giorgio lagerstätte, now in the Lake Lugano region of northern Italy and Switzerland, was in Triassic times a lagoon behind reefs with an anoxic bottom layer, so there were no scavengers and little turbulence to disturb fossilization, a situation that can be compared to the better-known Jurassic Solnhofen Limestone lagerstätte.

The remains of fish and various marine reptiles (including the common pachypleurosaur Neusticosaurus, and the bizarre long-necked archosauromorph Tanystropheus), along with some terrestrial forms like Ticinosuchus and Macrocnemus, have been recovered from this locality. All these fossils date from the Anisian/Ladinian transition (about 237 million years ago).

Triassic–Jurassic extinction event

 
The mass extinction event is marked by 'End Tr'

The Triassic Period ended with a mass extinction, which was particularly severe in the oceans; the conodonts disappeared, as did all the marine reptiles except ichthyosaurs and plesiosaurs. Invertebrates like brachiopods and molluscs (such as gastropods) were severely affected. In the oceans, 22% of marine families and possibly about half of marine genera went missing.

Though the end-Triassic extinction event was not equally devastating in all terrestrial ecosystems, several important clades of crurotarsans (large archosaurian reptiles previously grouped together as the thecodonts) disappeared, as did most of the large labyrinthodont amphibians, groups of small reptiles, and most synapsids. Some of the early, primitive dinosaurs also became extinct, but more adaptive ones survived to evolve into the Jurassic. Surviving plants that went on to dominate the Mesozoic world included modern conifers and cycadeoids.

The cause of the Late Triassic extinction is uncertain. It was accompanied by huge volcanic eruptions that occurred as the supercontinent Pangaea began to break apart about 202 to 191 million years ago (40Ar/39Ar dates),[47] forming the Central Atlantic Magmatic Province (CAMP),[48] one of the largest known inland volcanic events since the planet had first cooled and stabilized. Other possible but less likely causes for the extinction events include global cooling or even a bolide impact, for which an impact crater containing Manicouagan Reservoir in Quebec, Canada, has been singled out. However, the Manicouagan impact melt has been dated to 214±1 Mya. The date of the Triassic-Jurassic boundary has also been more accurately fixed recently, at 201.4 Mya. Both dates are gaining accuracy by using more accurate forms of radiometric dating, in particular the decay of uranium to lead in zircons formed at time of the impact. So, the evidence suggests the Manicouagan impact preceded the end of the Triassic by approximately 10±2 Ma. It could not therefore be the immediate cause of the observed mass extinction.[49]

 
Skull of a Triassic Period phytosaur found in the Petrified Forest National Park

The number of Late Triassic extinctions is disputed. Some studies suggest that there are at least two periods of extinction towards the end of the Triassic, separated by 12 to 17 million years. But arguing against this is a recent study of North American faunas. In the Petrified Forest of northeast Arizona there is a unique sequence of late Carnian-early Norian terrestrial sediments. An analysis in 2002 found no significant change in the paleoenvironment.[50] Phytosaurs, the most common fossils there, experienced a change-over only at the genus level, and the number of species remained the same. Some aetosaurs, the next most common tetrapods, and early dinosaurs, passed through unchanged. However, both phytosaurs and aetosaurs were among the groups of archosaur reptiles completely wiped out by the end-Triassic extinction event.

It seems likely then that there was some sort of end-Carnian extinction, when several herbivorous archosauromorph groups died out, while the large herbivorous therapsids—the kannemeyeriid dicynodonts and the traversodont cynodonts—were much reduced in the northern half of Pangaea (Laurasia).

These extinctions within the Triassic and at its end allowed the dinosaurs to expand into many niches that had become unoccupied. Dinosaurs became increasingly dominant, abundant and diverse, and remained that way for the next 150 million years. The true "Age of Dinosaurs" is during the following Jurassic and Cretaceous periods, rather than the Triassic.

See also

Notes

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  2. ^ McElwain, J. C.; Punyasena, S. W. (2007). "Mass extinction events and the plant fossil record". Trends in Ecology & Evolution. 22 (10): 548–557. doi:10.1016/j.tree.2007.09.003. PMID 17919771.
  3. ^ Retallack, G. J.; Veevers, J.; Morante, R. (1996). "Global coal gap between Permian–Triassic extinctions and middle Triassic recovery of peat forming plants". GSA Bulletin. 108 (2): 195–207. Bibcode:1996GSAB..108..195R. doi:10.1130/0016-7606(1996)108<0195:GCGBPT>2.3.CO;2. Retrieved 2007-09-29.
  4. ^ Payne, J. L.; Lehrmann, D. J.; Wei, J.; Orchard, M. J.; Schrag, D. P.; Knoll, A. H. (2004). "Large Perturbations of the Carbon Cycle During Recovery from the End-Permian Extinction". Science. 305 (5683): 506–9. Bibcode:2004Sci...305..506P. doi:10.1126/science.1097023. PMID 15273391. S2CID 35498132.
  5. ^ Ogg, James G.; Ogg, Gabi M.; Gradstein, Felix M. (2016). "Triassic". A Concise Geologic Time Scale: 2016. Elsevier. pp. 133–149. ISBN 978-0-444-63771-0.
  6. ^ Hongfu, Yin; Kexin, Zhang; Jinnan, Tong; Zunyi, Yang; Shunbao, Wu (June 2001). "The Global Stratotype Section and Point (GSSP) of the Permian-Triassic Boundary" (PDF). Episodes. 24 (2): 102–114. doi:10.18814/epiiugs/2001/v24i2/004. Retrieved 8 December 2020.
  7. ^ Hillebrandt, A.v.; Krystyn, L.; Kürschner, W. M.; et al. (September 2013). "The Global Stratotype Sections and Point (GSSP) for the base of the Jurassic System at Kuhjoch (Karwendel Mountains, Northern Calcareous Alps, Tyrol, Austria)". Episodes. 36 (3): 162–198. CiteSeerX 10.1.1.736.9905. doi:10.18814/epiiugs/2013/v36i3/001.
  8. ^ "Triassic". Dictionary.com Unabridged (Online). n.d.
  9. ^ Ogg, James G.; Ogg, Gabi M.; Gradstein, Felix M. (2016). "Triassic". A Concise Geologic Time Scale: 2016. Elsevier. pp. 133–149. ISBN 978-0444637710.
  10. ^ a b Sahney, S. & Benton, M.J. (2008). "Recovery from the most profound mass extinction of all time". Proceedings of the Royal Society B: Biological Sciences. 275 (1636): 759–765. doi:10.1098/rspb.2007.1370. PMC 2596898. PMID 18198148.
  11. ^ Brusatte, S. L.; Benton, M. J.; Ruta, M.; Lloyd, G. T. (2008-09-12). (PDF). Science. 321 (5895): 1485–1488. Bibcode:2008Sci...321.1485B. doi:10.1126/science.1161833. hdl:20.500.11820/00556baf-6575-44d9-af39-bdd0b072ad2b. PMID 18787166. S2CID 13393888. Archived from the original (PDF) on 2014-06-24. Retrieved 2012-01-14.
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  13. ^ Friedrich von Alberti, Beitrag zu einer Monographie des bunten Sandsteins, Muschelkalks und Keupers, und die Verbindung dieser Gebilde zu einer Formation [Contribution to a monograph on the colored sandstone, shell limestone and mudstone, and the joining of these structures into one formation] (Stuttgart and Tübingen, (Germany): J. G. Cotta, 1834). Alberti coined the term "Trias" on page 324 :
    "… bunter Sandstein, Muschelkalk und Keuper das Resultat einer Periode, ihre Versteinerungen, um mich der Worte E. de Beaumont’s zu bedeinen, die Thermometer einer geologischen Epoche seyen, … also die bis jezt beobachtete Trennung dieser Gebilde in 3 Formationen nicht angemessen, und es mehr dem Begriffe Formation entsprechend sey, sie zu einer Formation, welche ich vorläufig Trias nennen will, zu verbinden."
    ( … colored sandstone, shell limestone, and mudstone are the result of a period; their fossils are, to avail myself of the words of E. de Beaumont, the thermometer of a geologic epoch; … thus the separation of these structures into 3 formations, which has been maintained until now, isn't appropriate, and it is more consistent with the concept of "formation" to join them into one formation, which for now I will name "trias".)
  14. ^ Forte, Giuseppa; Kustatscher, Evelyn; Preto, Nereo (1 September 2022). "Carbon (δ13C) isotope variations indicate climate shifts and reflect plant habitats in the Middle Triassic (Anisian, Pelsonian) succession at Kühwiesenkopf/Monte Prà della Vacca (Dolomites, Northeast Italy)". Palaeogeography, Palaeoclimatology, Palaeoecology. 601: 111098. doi:10.1016/j.palaeo.2022.111098. S2CID 249483335. Retrieved 1 December 2022.
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References

  • Emiliani, Cesare. (1992). Planet Earth: Cosmology, Geology, & the Evolution of Life & the Environment. Cambridge University Press. (Paperback Edition ISBN 0-521-40949-7)
  • Ogg, Jim; June, 2004, Overview of Global Boundary Stratotype Sections and Points (GSSP's) Stratigraphy.org, Accessed April 30, 2006
  • Stanley, Steven M. Earth System History. New York: W.H. Freeman and Company, 1999. ISBN 0-7167-2882-6
  • Sues, Hans-Dieter & Fraser, Nicholas C. Triassic Life on Land: The Great Transition New York: Columbia University Press, 2010. Series: Critical Moments and Perspectives in Earth History and Paleobiology. ISBN 978-0-231-13522-1
  • van Andel, Tjeerd, (1985) 1994, New Views on an Old Planet: A History of Global Change, Cambridge University Press

External links

  • Douglas Henderson's illustrations of Triassic animals[permanent dead link]
  • Examples of Triassic Fossils

triassic, geologic, period, system, which, spans, million, years, from, permian, period, million, years, beginning, jurassic, period, first, shortest, period, mesozoic, both, start, period, marked, major, extinction, events, period, subdivided, into, three, ep. The Triassic t r aɪ ˈ ae s ɪ k try ASS ik 8 is a geologic period and system which spans 50 6 million years from the end of the Permian Period 251 902 million years ago Mya to the beginning of the Jurassic Period 201 36 Mya 9 The Triassic is the first and shortest period of the Mesozoic Era Both the start and end of the period are marked by major extinction events 10 The Triassic Period is subdivided into three epochs Early Triassic Middle Triassic and Late Triassic Triassic251 902 0 024 201 4 0 2 Ma PreꞒ Ꞓ O S D C P T J K Pg NChronology 255 250 245 240 235 230 225 220 215 210 205 200 PzMesozoicPermianTriassicJurassicEarMiddleLateOlenekianInduanAnisianLadinianCarnianNorianRhaetian Permian Triassic extinction event Smithian Spathian boundary event 1 Carnian pluvial episode Full recovery of woody trees 2 Coals return 3 Scleractiniancorals amp calcified sponges 4 Triassic Jurassic extinction event Manicouagan impactSubdivision of the Triassic according to the ICS as of 2021 5 Vertical axis scale millions of years ago EtymologyName formalityFormalUsage informationCelestial bodyEarthRegional usageGlobal ICS Time scale s usedICS Time ScaleDefinitionChronological unitPeriodStratigraphic unitSystemTime span formalityFormalLower boundary definitionFirst appearance of the conodont Hindeodus parvusLower boundary GSSPMeishan Zhejiang China31 04 47 N 119 42 21 E 31 0798 N 119 7058 E 31 0798 119 7058Lower GSSP ratified2001 6 Upper boundary definitionFirst appearance of the ammonite Psiloceras spelae tirolicumUpper boundary GSSPKuhjoch section Karwendel mountains Northern Calcareous Alps Austria47 29 02 N 11 31 50 E 47 4839 N 11 5306 E 47 4839 11 5306Upper GSSP ratified2010 7 The Triassic began in the wake of the Permian Triassic extinction event which left the Earth s biosphere impoverished it was well into the middle of the Triassic before life recovered its former diversity Three categories of organisms can be distinguished in the Triassic record survivors from the extinction event new groups that flourished briefly and other new groups that went on to dominate the Mesozoic Era Reptiles especially archosaurs were the chief terrestrial vertebrates during this time A specialized subgroup of archosaurs called dinosaurs first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period 11 Archosaurs that became dominant in this period were primarily pseudosuchians ancestors of modern crocodilians while some archosaurs specialized in flight the first time among vertebrates becoming the pterosaurs Therapsids the dominant vertebrates of the preceding Permian period declined throughout the period The first true mammals themselves a specialized subgroup of therapsids also evolved during this period The vast supercontinent of Pangaea existed until the mid Triassic after which it began to gradually rift into two separate landmasses Laurasia to the north and Gondwana to the south The global climate during the Triassic was mostly hot and dry 12 with deserts spanning much of Pangaea s interior However the climate shifted and became more humid as Pangaea began to drift apart The end of the period was marked by yet another major mass extinction the Triassic Jurassic extinction event that wiped out many groups including most pseudosuchians and allowed dinosaurs to assume dominance in the Jurassic Contents 1 Etymology 2 Dating and subdivisions 3 Paleogeography 3 1 Scandinavia 3 2 Paleooceanography 4 Climate 5 Flora 5 1 Land plants 5 1 1 Coal 5 2 Phytoplankton 6 Fauna 6 1 Marine invertebrates 6 2 Fish 6 3 Amphibians 6 4 Reptiles 6 4 1 Archosauromorphs 6 4 2 Marine reptiles 6 4 3 Other reptiles 6 5 Synapsids 7 Lagerstatten 8 Triassic Jurassic extinction event 9 See also 10 Notes 11 References 12 External linksEtymology EditThe Triassic was named in 1834 by Friedrich August von Alberti after a succession of three distinct rock layers Greek trias meaning triad that are widespread in southern Germany the lower Buntsandstein colourful sandstone the middle Muschelkalk shell bearing limestone and the upper Keuper coloured clay 13 Early Triassic sandstone Buntsandstein near Stadtroda Germany Middle Triassic Muschelkalk shell bearing limestone near Dorzbach Germany Late Triassic Steigerwald Formation and overlying Hassberge Formation in Schonbuch GermanyDating and subdivisions EditOn the geologic time scale the Triassic is usually divided into Early Middle and Late Triassic Epochs and the corresponding rocks are referred to as Lower Middle or Upper Triassic The faunal stages from the youngest to oldest are Series Epoch Faunal stage Time spanUpper Late Triassic Tr3 Rhaetian 208 5 201 4 0 2 Mya Norian 227 208 5 Mya Carnian 237 227 Mya Middle Triassic Tr2 Ladinian 242 237 Mya Anisian 247 2 242 Mya Lower Early Triassic Scythian Olenekian 251 2 247 2 Mya Induan 251 902 0 024 251 2 Mya Paleogeography Edit View of the Tethys area during the Ladinian stage 230 Ma continental reconstructionDuring the Triassic almost all the Earth s land mass was concentrated into a single supercontinent Pangaea lit entire land 14 This supercontinent was more or less centered on the equator and extended between the poles though it did drift northwards as the period progressed Southern Pangea also known as Gondwana was made up by closely appressed cratons corresponding to modern South America Africa Madagascar India Antarctica and Australia North Pangea also known as Laurussia or Laurasia corresponds to modern day North America and the fragmented predecessors of Eurasia The western edge of Pangea lay at the margin of an enormous ocean Panthalassa lit entire sea which roughly corresponds to the modern Pacific Ocean Practically all deep ocean crust present during the Triassic has been recycled through the subduction of oceanic plates so very little is known about the open ocean from this time period Most information on Panthalassan geology and marine life is derived from island arcs and rare seafloor sediments accreted onto surrounding land masses such as present day Japan and western North America The eastern edge of Pangea was encroached upon by a pair of extensive oceanic basins The Neo Tethys or simply Tethys and Paleo Tethys Oceans These extended from China to Iberia hosting abundant marine life along their shallow tropical peripheries They were divided from each other by a long string of microcontinents known as the Cimmerian terranes Cimmerian crust had detached from Gondwana in the early Permian and drifted northwards during the Triassic enlarging the Neo Tethys Ocean which formed in their wake At the same time they forced the Paleo Tethys Ocean to shrink as it was being subducted under Asia By the end of the Triassic the Paleo Tethys Ocean occupied a small area and the Cimmerian terranes began to collide with southern Asia This collision known as the Cimmerian Orogeny continued into the Jurassic and Cretaceous to produce a chain of mountain ranges stretching from Turkey to Malaysia 15 16 Sydney Australia lies on Triassic shales and sandstones Almost all of the exposed rocks around Sydney belong to the Triassic Sydney sandstone 17 Pangaea was fractured by widespread faulting and rift basins during the Triassic especially late in that period but had not yet separated The first nonmarine sediments in the rift that marks the initial break up of Pangaea which separated eastern North America from Morocco are of Late Triassic age in the United States these thick sediments comprise the Newark Supergroup 18 Rift basins are also common in South America Europe and Africa Terrestrial environments are particularly well represented in the South Africa 19 Russia central Europe and the southwest United States Terrestrial Triassic biostratigraphy is mostly based on terrestrial and freshwater tetrapods as well as conchostracans clam shrimps a type of fast breeding crustacean which lived in lakes and hypersaline environments Because a supercontinent has less shoreline compared to a series of smaller continents Triassic marine deposits are relatively uncommon on a global scale A major exception is in Western Europe where the Triassic was first studied The northeastern margin of Gondwana was a stable passive margin along the Neo Tethys Ocean and marine sediments have been preserved in parts of northern India and Arabia 16 In North America marine deposits are limited to a few exposures in the west Scandinavia Edit During the Triassic peneplains are thought to have formed in what is now Norway and southern Sweden 20 21 22 Remnants of this peneplain can be traced as a tilted summit accordance in the Swedish West Coast 20 In northern Norway Triassic peneplains may have been buried in sediments to be then re exposed as coastal plains called strandflats 21 Dating of illite clay from a strandflat of Bomlo southern Norway have shown that landscape there became weathered in Late Triassic times c 210 million years ago with the landscape likely also being shaped during that time 23 Paleooceanography Edit Eustatic sea level in the Triassic was consistently low compared to the other geological periods The beginning of the Triassic was around present sea level rising to about 10 20 metres 33 66 ft above present day sea level during the Early and Middle Triassic Sea level rise accelerated in the Ladinian culminating with a sea level up to 50 metres 164 ft above present day levels during the Carnian Sea level began to decline in the Norian reaching a low of 50 metres 164 ft below present sea level during the mid Rhaetian Low global sea levels persisted into the earliest Jurassic The long term sea level trend is superimposed by 22 sea level drop events widespread in the geologic record mostly of minor less than 25 metre 82 ft and medium 25 75 metre 82 246 ft magnitudes A lack of evidence for Triassic continental ice sheets suggest that glacial eustasy is unlikely to be the cause of these changes 24 Climate EditThe Triassic continental interior climate was generally hot and dry so that typical deposits are red bed sandstones and evaporites There is no evidence of glaciation at or near either pole in fact the polar regions were apparently moist and temperate providing a climate suitable for forests and vertebrates including reptiles Pangaea s large size limited the moderating effect of the global ocean its continental climate was highly seasonal with very hot summers and cold winters 25 The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross equatorial monsoons 25 sometimes referred to as megamonsoons 26 The Triassic may have mostly been a dry period but evidence exists that it was punctuated by several episodes of increased rainfall in tropical and subtropical latitudes of the Tethys Sea and its surrounding land 27 Sediments and fossils suggestive of a more humid climate are known from the Anisian to Ladinian of the Tethysian domain and from the Carnian and Rhaetian of a larger area that includes also the Boreal domain e g Svalbard Islands the North American continent the South China block and Argentina The best studied of such episodes of humid climate and probably the most intense and widespread was the Carnian Pluvial Event A 2020 study found bubbles of carbon dioxide in basaltic rocks dating back to the end of the Triassic and concluded that volcanic activity helped trigger climate change in that period 28 Flora Edit Triassic flora as depicted in Meyers Konversations Lexikon 1885 90 Land plants Edit On land the surviving vascular plants included the lycophytes the dominant cycadophytes ginkgophyta represented in modern times by Ginkgo biloba ferns horsetails and glossopterids The spermatophytes or seed plants came to dominate the terrestrial flora in the northern hemisphere conifers ferns and bennettitales flourished The seed fern genus Dicroidium would dominate Gondwana throughout the period Coal Edit Immediately above the Permian Triassic boundary the glossopteris flora was suddenly 29 largely displaced by an Australia wide coniferous flora No known coal deposits date from the start of the Triassic Period This is known as the Early Triassic coal gap and can be seen as part of the Permian Triassic extinction event 30 Possible explanations for the coal gap include sharp drops in sea level at the time of the Permo Triassic boundary 31 acid rain from the Siberian Traps eruptions or from an impact event that overwhelmed acidic swamps climate shift to a greenhouse climate that was too hot and dry for peat accumulation evolution of fungi or herbivores that were more destructive of wetlands the extinction of all plants adapted to peat swamps with a hiatus of several million years before new plant species evolved that were adapted to peat swamps 30 or soil anoxia as oxygen levels plummeted 32 Phytoplankton Edit Before the Permian extinction Archaeplastida red and green algae had been the major marine phytoplanktons since about 659 645 million years ago 33 when they replaced marine planktonic cyanobacteria which first appeared about 800 million years ago as the dominant phytoplankton in the oceans 34 In the Triassic secondary endosymbiotic algae became the most important plankton 35 Fauna Edit Middle Triassic marginal marine sequence southwestern Utah Reconstruction of the Triassic amphibian Mastodonsaurus Marine invertebrates Edit In marine environments new modern types of corals appeared in the Early Triassic forming small patches of reefs of modest extent compared to the great reef systems of Devonian or modern times Serpulids appeared in the Middle Triassic 36 Microconchids were abundant The shelled cephalopods called ammonites recovered diversifying from a single line that survived the Permian extinction Fish Edit The fish fauna was remarkably uniform with many families and genera exhibiting a global distribution in the wake of the Permian Triassic mass extinction event 37 Ray finned fishes actinopterygians went through a remarkable diversification during the Triassic leading to peak diversity during the Middle Triassic however the pattern of this diversification is still not well understood due to a taphonomic megabias 38 Large predatory actinopterygians such as saurichthyids and birgeriids appeared in the Early Triassic and became widespread and successful during the period as a whole 39 Lakes and rivers were populated by lungfish Dipnoi such as Ceratodus which are mainly known from the dental plates abundant in the fossils record 40 Hybodonts a group of shark like cartilaginous fish were dominant in both freshwater and marine environments throughout the Triassic 41 Amphibians Edit Temnospondyl amphibians were among those groups that survived the Permian Triassic extinction Once abundant in both terrestrial and aquatic environments the terrestrial species had mostly died out during the extinction event The Triassic survivors were aquatic or semi aquatic and were represented by Tupilakosaurus Thabanchuia Branchiosauridae and Micropholis all of which died out in Early Triassic and the successful Stereospondyli with survivors into the Cretaceous Period The largest Triassic stereospondyls such as Mastodonsaurus were up to 4 to 6 metres 13 to 20 ft in length 42 43 Some lineages e g trematosaurs flourished briefly in the Early Triassic while others e g capitosaurs remained successful throughout the whole period or only came to prominence in the Late Triassic e g Plagiosaurus metoposaurs The first Lissamphibians modern amphibians appear in the Triassic with the progenitors of the first frogs already present by the Early Triassic However the group as a whole did not become common until the Jurassic when the temnospondyls had become very rare Most of the Reptiliomorpha stem amniotes that gave rise to the amniotes disappeared in the Triassic but two water dwelling groups survived Embolomeri that only survived into the early part of the period and the Chroniosuchia which survived until the end of the Triassic Reptiles Edit Archosauromorphs Edit The Permian Triassic extinction devastated terrestrial life Biodiversity rebounded as the surviving species repopulated empty terrain but these were short lived Diverse communities with complex food web structures took 30 million years to reestablish 10 44 Archosauromorph reptiles which had already appeared and diversified to an extent in the Permian Period exploding in diversity as an adaptive radiation in response to the Permian Triassic mass extinction By the Early Triassic several major archosauromorph groups had appeared Long necked lizard like early archosauromorphs were known as protorosaurs which is likely a paraphyletic group rather than a true clade Tanystropheids were a family of protorosaurs which elevated their neck size to extremes with the largest genus Tanystropheus having a neck longer than its body The protorosaur family Sharovipterygidae used their elongated hindlimbs for gliding Other archosauromorphs such as rhynchosaurs and allokotosaurs were mostly stocky bodied herbivores with specialized jaw structures Rhynchosaurs barrel gutted herbivores thrived for only a short period of time becoming extinct about 220 million years ago They were exceptionally abundant in the middle of the Triassic as the primary large herbivores in many Carnian age ecosystems They sheared plants with premaxillary beaks and plates along the upper jaw with multiple rows of teeth Allokotosaurs were iguana like reptiles including Trilophosaurus a common Late Triassic reptile with three crowned teeth Teraterpeton which had a long beak like snout and Shringasaurus a horned herbivore which reached a body length of 3 4 metres 9 8 13 1 ft One group of archosauromorphs the archosauriforms were distinguished by their active predatory lifestyle with serrated teeth and upright limb postures Archosauriforms were diverse in the Triassic including various terrestrial and semiaquatic predators of all shapes and sizes The large headed and robust erythrosuchids were among the dominant carnivores in the early Triassic Phytosaurs were a particularly common group which prospered during the Late Triassic These long snouted and semiaquatic predators resemble living crocodiles and probably had a similar lifestyle hunting for fish and small reptiles around the water s edge However this resemblance is only superficial and is a prime case of convergent evolution True archosaurs appeared in the early Triassic splitting into two branches Avemetatarsalia the ancestors to birds and Pseudosuchia the ancestors to crocodilians Avemetatarsalians were a minor component of their ecosystems but eventually produced the earliest pterosaurs and dinosaurs in the Late Triassic Early long tailed pterosaurs appeared in the Norian and quickly spread worldwide Triassic dinosaurs evolved in the Carnian and include early sauropodomorphs and theropods Most Triassic dinosaurs were small predators and only a few were common such as Coelophysis which was 1 to 2 metres 3 3 to 6 6 ft long The large predator Smok was most likely also an archosaur but it is uncertain if it was a primitive dinosaur or a pseudosuchian Pseudosuchians were far more ecologically dominant in the Triassic including large herbivores such as aetosaurs large carnivores rauisuchians and the first crocodylomorphs sphenosuchians Aetosaurs were heavily armored reptiles that were common during the last 30 million years of the Late Triassic until they died out at the Triassic Jurassic extinction Most aetosaurs were herbivorous and fed on low growing plants but some may have eaten meat rauisuchians formally known as paracrocodylomorphs were the keystone predators of most Triassic terrestrial ecosystems Over 25 species have been found including giant quadrupedal hunters sleek bipedal omnivores and lumbering beasts with deep sails on their backs They probably occupied the large predator niche later filled by theropods Rauisuchians were ancestral to small lightly built crocodylomorphs the only pseudosuchians which survived into the Jurassic Tanystropheus a long necked tanystropheid Proterosuchus a crocodile like early archosauriform from the Early Triassic Staurikosaurus one of the earliest dinosaurs a member of the Triassic family Herrerasauridae Postosuchus a rauisuchid which was an apex predator in parts of Late Triassic North America Plateosaurus was one of the largest of early sauropodomorphs or prosauropods of the Late Triassic Coelophysis was one of the most abundant theropod dinosaurs in the Late TriassicMarine reptiles Edit Marine vertebrate apex predators of the Early Triassic and Anisian Middle Triassic 45 There were many types of marine reptiles These included the Sauropterygia which featured pachypleurosaurus and nothosaurs both common during the Middle Triassic especially in the Tethys region placodonts the earliest known herbivorous marine reptile Atopodentatus and the first plesiosaurs The first of the lizardlike Thalattosauria askeptosaurs and the highly successful ichthyosaurs which appeared in Early Triassic seas soon diversified and some eventually developed to huge size during the Late Triassic Other reptiles Edit Among other reptiles the earliest turtles like Proganochelys and Proterochersis appeared during the Norian Age Stage of the Late Triassic Period The Lepidosauromorpha specifically the Sphenodontia are first found in the fossil record of the earlier Carnian Age though the earliest lepidosauromorphs likely occurred in the Permian The Procolophonidae the last surviving parareptiles were an important group of small lizard like herbivores The drepanosaurs were a clade of unusual chameleon like arboreal reptiles with birdlike heads and specialised claws Synapsids Edit Three therapsid groups survived into the Triassic dicynodonts therocephalians and cynodonts The cynodont Cynognathus was a characteristic top predator in the Olenekian and Anisian of Gondwana Both kannemeyeriiform dicynodonts and gomphodont cynodonts remained important herbivores during much of the period Therocephalians included both large predators Moschorhinus and herbivorous forms bauriids until their extinction midway through the period Ecteniniid cynodonts played a role as large sized cursorial predators in the Late Triassic During the Carnian early part of the Late Triassic some advanced cynodonts gave rise to the first mammals During the Triassic archosaurs displaced therapsids as the largest and most ecologically prolific terrestrial amniotes This Triassic Takeover may have contributed to the evolution of mammals by forcing the surviving therapsids and their mammaliaform successors to live as small mainly nocturnal insectivores Nocturnal life may have forced the mammaliaforms to develop fur and a higher metabolic rate 46 Lystrosaurus was a widespread dicynodont and the most common land vertebrate during the Early Triassic after animal life had been greatly diminished Cynognathus was a carnivorous mammal like cynodont from the Early Triassic Lagerstatten EditThe Monte San Giorgio lagerstatte now in the Lake Lugano region of northern Italy and Switzerland was in Triassic times a lagoon behind reefs with an anoxic bottom layer so there were no scavengers and little turbulence to disturb fossilization a situation that can be compared to the better known Jurassic Solnhofen Limestone lagerstatte The remains of fish and various marine reptiles including the common pachypleurosaur Neusticosaurus and the bizarre long necked archosauromorph Tanystropheus along with some terrestrial forms like Ticinosuchus and Macrocnemus have been recovered from this locality All these fossils date from the Anisian Ladinian transition about 237 million years ago Triassic Jurassic extinction event EditMain article Triassic Jurassic extinction event The mass extinction event is marked by End Tr The Triassic Period ended with a mass extinction which was particularly severe in the oceans the conodonts disappeared as did all the marine reptiles except ichthyosaurs and plesiosaurs Invertebrates like brachiopods and molluscs such as gastropods were severely affected In the oceans 22 of marine families and possibly about half of marine genera went missing Though the end Triassic extinction event was not equally devastating in all terrestrial ecosystems several important clades of crurotarsans large archosaurian reptiles previously grouped together as the thecodonts disappeared as did most of the large labyrinthodont amphibians groups of small reptiles and most synapsids Some of the early primitive dinosaurs also became extinct but more adaptive ones survived to evolve into the Jurassic Surviving plants that went on to dominate the Mesozoic world included modern conifers and cycadeoids The cause of the Late Triassic extinction is uncertain It was accompanied by huge volcanic eruptions that occurred as the supercontinent Pangaea began to break apart about 202 to 191 million years ago 40Ar 39Ar dates 47 forming the Central Atlantic Magmatic Province CAMP 48 one of the largest known inland volcanic events since the planet had first cooled and stabilized Other possible but less likely causes for the extinction events include global cooling or even a bolide impact for which an impact crater containing Manicouagan Reservoir in Quebec Canada has been singled out However the Manicouagan impact melt has been dated to 214 1 Mya The date of the Triassic Jurassic boundary has also been more accurately fixed recently at 201 4 Mya Both dates are gaining accuracy by using more accurate forms of radiometric dating in particular the decay of uranium to lead in zircons formed at time of the impact So the evidence suggests the Manicouagan impact preceded the end of the Triassic by approximately 10 2 Ma It could not therefore be the immediate cause of the observed mass extinction 49 Skull of a Triassic Period phytosaur found in the Petrified Forest National Park The number of Late Triassic extinctions is disputed Some studies suggest that there are at least two periods of extinction towards the end of the Triassic separated by 12 to 17 million years But arguing against this is a recent study of North American faunas In the Petrified Forest of northeast Arizona there is a unique sequence of late Carnian early Norian terrestrial sediments An analysis in 2002 found no significant change in the paleoenvironment 50 Phytosaurs the most common fossils there experienced a change over only at the genus level and the number of species remained the same Some aetosaurs the next most common tetrapods and early dinosaurs passed through unchanged However both phytosaurs and aetosaurs were among the groups of archosaur reptiles completely wiped out by the end Triassic extinction event It seems likely then that there was some sort of end Carnian extinction when several herbivorous archosauromorph groups died out while the large herbivorous therapsids the kannemeyeriid dicynodonts and the traversodont cynodonts were much reduced in the northern half of Pangaea Laurasia These extinctions within the Triassic and at its end allowed the dinosaurs to expand into many niches that had become unoccupied Dinosaurs became increasingly dominant abundant and diverse and remained that way for the next 150 million years The true Age of Dinosaurs is during the following Jurassic and Cretaceous periods rather than the Triassic See also Edit Mesozoic portal Geology portal Paleontology portalGeologic time scale List of fossil sites with link directory Triassic land vertebrate faunachrons Phylloceratina DinosaursNotes Edit Widmann Philipp Bucher Hugo Leu Marc et al 2020 Dynamics of the Largest Carbon Isotope Excursion During the Early Triassic Biotic Recovery Frontiers in Earth Science 8 196 196 Bibcode 2020FrEaS 8 196W doi 10 3389 feart 2020 00196 McElwain J C Punyasena S W 2007 Mass extinction events and the plant fossil record Trends in Ecology amp Evolution 22 10 548 557 doi 10 1016 j tree 2007 09 003 PMID 17919771 Retallack G J Veevers J Morante R 1996 Global coal gap between Permian Triassic extinctions and middle Triassic recovery of peat forming plants GSA Bulletin 108 2 195 207 Bibcode 1996GSAB 108 195R doi 10 1130 0016 7606 1996 108 lt 0195 GCGBPT gt 2 3 CO 2 Retrieved 2007 09 29 Payne J L Lehrmann D J Wei J Orchard M J Schrag D P Knoll A H 2004 Large Perturbations of the Carbon Cycle During Recovery from the End Permian Extinction Science 305 5683 506 9 Bibcode 2004Sci 305 506P doi 10 1126 science 1097023 PMID 15273391 S2CID 35498132 Ogg James G Ogg Gabi M Gradstein Felix M 2016 Triassic A Concise Geologic Time Scale 2016 Elsevier pp 133 149 ISBN 978 0 444 63771 0 Hongfu Yin Kexin Zhang Jinnan Tong Zunyi Yang Shunbao Wu June 2001 The Global Stratotype Section and Point GSSP of the Permian Triassic Boundary PDF Episodes 24 2 102 114 doi 10 18814 epiiugs 2001 v24i2 004 Retrieved 8 December 2020 Hillebrandt A v Krystyn L Kurschner W M et al September 2013 The Global Stratotype Sections and Point GSSP for the base of the Jurassic System at Kuhjoch Karwendel Mountains Northern Calcareous Alps Tyrol Austria Episodes 36 3 162 198 CiteSeerX 10 1 1 736 9905 doi 10 18814 epiiugs 2013 v36i3 001 Triassic Dictionary com Unabridged Online n d Ogg James G Ogg Gabi M Gradstein Felix M 2016 Triassic A Concise Geologic Time Scale 2016 Elsevier pp 133 149 ISBN 978 0444637710 a b Sahney S amp Benton M J 2008 Recovery from the most profound mass extinction of all time Proceedings of the Royal Society B Biological Sciences 275 1636 759 765 doi 10 1098 rspb 2007 1370 PMC 2596898 PMID 18198148 Brusatte S L Benton M J Ruta M Lloyd G T 2008 09 12 Superiority Competition and Opportunism in the Evolutionary Radiation of Dinosaurs PDF Science 321 5895 1485 1488 Bibcode 2008Sci 321 1485B doi 10 1126 science 1161833 hdl 20 500 11820 00556baf 6575 44d9 af39 bdd0b072ad2b PMID 18787166 S2CID 13393888 Archived from the original PDF on 2014 06 24 Retrieved 2012 01 14 Lethally Hot Earth Was Devoid of Life Could It Happen Again nationalgeographic com 19 October 2012 Friedrich von Alberti Beitrag zu einer Monographie des bunten Sandsteins Muschelkalks und Keupers und die Verbindung dieser Gebilde zu einer Formation Contribution to a monograph on the colored sandstone shell limestone and mudstone and the joining of these structures into one formation Stuttgart and Tubingen Germany J G Cotta 1834 Alberti coined the term Trias on page 324 bunter Sandstein Muschelkalk und Keuper das Resultat einer Periode ihre Versteinerungen um mich der Worte E de Beaumont s zu bedeinen die Thermometer einer geologischen Epoche seyen also die bis jezt beobachtete Trennung dieser Gebilde in 3 Formationen nicht angemessen und es mehr dem Begriffe Formation entsprechend sey sie zu einer Formation welche ich vorlaufig Trias nennen will zu verbinden colored sandstone shell limestone and mudstone are the result of a period their fossils are to avail myself of the words of E de Beaumont the thermometer of a geologic epoch thus the separation of these structures into 3 formations which has been maintained until now isn t appropriate and it is more consistent with the concept of formation to join them into one formation which for now I will name trias Forte Giuseppa Kustatscher Evelyn Preto Nereo 1 September 2022 Carbon d13C isotope variations indicate climate shifts and reflect plant habitats in the Middle Triassic Anisian Pelsonian succession at Kuhwiesenkopf Monte Pra della Vacca Dolomites Northeast Italy Palaeogeography Palaeoclimatology Palaeoecology 601 111098 doi 10 1016 j palaeo 2022 111098 S2CID 249483335 Retrieved 1 December 2022 Mazaheri Johari Mina Roghi Guido Caggiati Marcello Kustatscher Evelyn Ghasemi Nejad Ebrahim Zanchi Andrea Gianolla Piero 15 January 2022 Disentangling climate signal from tectonic forcing The Triassic Aghdarband Basin Turan Domain Iran Palaeogeography Palaeoclimatology Palaeoecology 586 doi 10 1016 j palaeo 2021 110777 Retrieved 10 January 2023 a b Cocks L Robin M Torsvik Trond H eds 2016 Triassic Earth History and Palaeogeography Cambridge Cambridge University Press pp 195 207 doi 10 1017 9781316225523 012 ISBN 978 1 316 22552 3 retrieved 2022 05 16 Herbert Chris Helby Robin 1980 A Guide to the Sydney basin Maitland NSW Geological Survey of NSW p 582 ISBN 978 0 7240 1250 3 Lecture 10 Triassic Newark Chinle rainbow ldeo columbia edu Jacobs Louis L 1997 African Dinosaurs Encyclopedia of Dinosaurs Edited by Phillip J Currie and Kevin Padian Academic Press pp 2 4 a b Lidmar Bergstrom Karna 1993 Denudation surfaces and tectonics in the southernmost part of the Baltic Shield Precambrian Research 64 1 4 337 345 Bibcode 1993PreR 64 337L doi 10 1016 0301 9268 93 90086 h a b Olesen Odleiv Kierulf Halfdan Pascal Bronner Marco Dalsegg Einar Fredin Ola Solbakk Terje 2013 Deep weathering neotectonics and strandflat formation in Nordland northern Norway Norwegian Journal of Geology 93 189 213 Japsen Peter Green Paul F Bonow Johan M Erlstrom Mikael 2016 Episodic burial and exhumation of the southern Baltic Shield Epeirogenic uplifts during and after break up of Pangaea Gondwana Research 35 357 77 Bibcode 2016GondR 35 357J doi 10 1016 j gr 2015 06 005 Fredin Ola Viola Giulio Zwingmann Horst Sorlie Ronald Bronner Marco Lie Jan Erik Margrethe Grandal Else Muller Axel Margeth Annina Vogt Christoph Knies Jochen 2017 The inheritance of a Mesozoic landscape in western Scandinavia Nature 8 14879 Bibcode 2017NatCo 814879F doi 10 1038 ncomms14879 PMC 5477494 PMID 28452366 Haq Bilal U December 2018 Triassic Eustatic Variations Reexamined GSA Today Geological Society of America 28 12 4 9 doi 10 1130 GSATG381A 1 S2CID 134477691 a b Stanley 452 53 Zeng Zhiwei Zhu Hongtao Yang Xianghua Zeng Hongliu Hu Xiaolin Xia Chenchen May 2019 The Pangaea Megamonsoon records Evidence from the Triassic Mungaroo Formation Northwest Shelf of Australia Gondwana Research 69 1 24 doi 10 1016 j gr 2018 11 015 Retrieved 9 January 2023 Preto N Kustatscher E Wignall P B 2010 Triassic climates State of the art and perspectives Palaeogeography Palaeoclimatology Palaeoecology 290 1 4 1 10 Bibcode 2010PPP 290 1P doi 10 1016 j palaeo 2010 03 015 Manfredo Capriolo et al 2020 Deep CO2 in the end Triassic Central Atlantic Magmatic Province Vol 11 no 1670 Nature Communications doi 10 1038 s41467 020 15325 6 Hosher WT Magaritz M Clark D 1987 Events near the time of the Permian Triassic boundary Mod Geol 11 155 80 173 74 a b Retallack G J Veevers J J Morante R 1996 Global coal gap between Permian Triassic extinction and Middle Triassic recovery of peat forming plants Bulletin of the Geological Society of America 108 2 195 207 Bibcode 1996GSAB 108 195R doi 10 1130 0016 7606 1996 108 lt 0195 GCGBPT gt 2 3 CO 2 Holser WT Schoenlaub H P Klein P Attrep M Boeckelmann Klaus et al 1989 A unique geochemical record at the Permian Triassic boundary Nature 337 6202 39 42 Bibcode 1989Natur 337 39H doi 10 1038 337039a0 S2CID 8035040 Retallack G J Krull E S 2006 Carbon isotopic evidence for terminal Permian methane outbursts and their role in extinctions of animals plants coral reefs and peat swamps PDF Geological Society of America Special Paper 399 249 doi 10 1130 2006 2399 12 ISBN 9780813723990 Retrieved 14 December 2020 How snowball Earth gave rise to complex life Cosmos Magazine 16 August 2017 December Phytoplankton News University of Bristol The rise of algae in Cryogenian oceans and the emergence of animals ResearchGate Vinn O Mutvei H 2009 Calcareous tubeworms of the Phanerozoic PDF Estonian Journal of Earth Sciences 58 4 286 96 doi 10 3176 earth 2009 4 07 Retrieved 2012 09 16 Romano Carlo Koot Martha B Kogan Ilja Brayard Arnaud Minikh Alla V Brinkmann Winand Bucher Hugo Kriwet Jurgen February 2016 Permian Triassic Osteichthyes bony fishes diversity dynamics and body size evolution Biological Reviews 91 1 106 47 doi 10 1111 brv 12161 PMID 25431138 S2CID 5332637 Romano Carlo January 2021 A Hiatus Obscures the Early Evolution of Modern Lineages of Bony Fishes Frontiers in Earth Science 8 618853 doi 10 3389 feart 2020 618853 Romano Carlo Jenks James F Jattiot Romain Scheyer Torsten M 2017 Marine Early Triassic Actinopterygii from Elko County Nevada USA implications for the Smithian equatorial vertebrate eclipse Journal of Paleontology 91 5 1 22 doi 10 1017 jpa 2017 36 Agnolin F L Mateus O Milan J Marzola M Wings O Adolfssen J S amp Clemmensen L B 2018 Ceratodus tunuensis sp nov a new lungfish Sarcopterygii Dipnoi from the Upper Triassic of central East Greenland Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology e1439834 Kumar Krishna Bajpai Sunil Pandey Pragya Ghosh Triparna Bhattacharya Debasish 2021 08 04 Hybodont sharks from the Jurassic of Jaisalmer western India Historical Biology 34 6 953 963 doi 10 1080 08912963 2021 1954920 ISSN 0891 2963 S2CID 238781606 Wells Kentwood D 2010 The Ecology and Behavior of Amphibians University of Chicago Press ISBN 978 0226893334 via Google Books Benton Michael 2009 Vertebrate Palaeontology John Wiley amp Sons ISBN 978 1405144490 via Google Books Douglas Palmer amp Peter Barrett 2009 Evolution The Story of Life London The Natural History Museum ISBN 978 1845333393 a href Template Cite book html title Template Cite book cite book a CS1 maint uses authors parameter link Scheyer et al 2014 Early Triassic Marine Biotic Recovery The Predators Perspective PLoS ONE https doi org 10 1371 journal pone 0088987 Ruben J A amp Jones T D 2000 Selective Factors Associated with the Origin of Fur and Feathers American Zoologist 40 4 585 96 doi 10 1093 icb 40 4 585 Nomade et al 2007 Palaeogeography Palaeoclimatology Palaeoecology 244 326 44 Marzoli et al 1999 Science 284 Extensive 200 million year old continental flood basalts of the Central Atlantic Magmatic Province pp 618 620 Hodych amp Dunning 1992 NO SIGNIFICANT NONMARINE CARNIAN NORIAN LATE TRIASSIC EXTINCTION EVENT EVIDENCE FROM PETRIFIED FOREST NATIONAL PARK gsa confex com Archived from the original on 2003 11 06 Retrieved 2003 12 12 References EditEmiliani Cesare 1992 Planet Earth Cosmology Geology amp the Evolution of Life amp the Environment Cambridge University Press Paperback Edition ISBN 0 521 40949 7 Ogg Jim June 2004 Overview of Global Boundary Stratotype Sections and Points GSSP s Stratigraphy org Accessed April 30 2006 Stanley Steven M Earth System History New York W H Freeman and Company 1999 ISBN 0 7167 2882 6 Sues Hans Dieter amp Fraser Nicholas C Triassic Life on Land The Great Transition New York Columbia University Press 2010 Series Critical Moments and Perspectives in Earth History and Paleobiology ISBN 978 0 231 13522 1 van Andel Tjeerd 1985 1994 New Views on an Old Planet A History of Global Change Cambridge University PressExternal links Edit Wikimedia Commons has media related to Triassic Look up triassic in Wiktionary the free dictionary Wikisource has the text of the 1911 Encyclopaedia Britannica article Triassic System Overall introduction The Triassic world Douglas Henderson s illustrations of Triassic animals permanent dead link Paleofiles page on the Triassic extinctions Examples of Triassic Fossils Triassic chronostratigraphy scale Retrieved from https en wikipedia org w index php title Triassic amp oldid 1135533500, wikipedia, wiki, book, books, library,

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