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List of African animals extinct in the Holocene

This list of African species extinct in the Holocene covers extinctions from the Holocene epoch, a geologic epoch that began about 11,650 years before present (about 9700 BCE)[a] and continues to the present.[1]

Map of Africa
The quagga (Equus quagga quagga), extinct since 1883, was zebra-like in the front but more horse-like in the rear. A breeding program aims to create similar-looking animals, but these are not true quaggas.

Africa is highly biodiverse; it is the continent with the largest number of megafauna species, as it was least affected by the extinction of the Pleistocene megafauna. However, a few species have disappeared from Africa as part of the ongoing Holocene extinction, driven by human activity.

Madagascar and the Indian Ocean islands, Macaronesia, and Saint Helena, Ascension and Tristan da Cunha are biogeographically distinct from mainland Africa and have a much greater number of Holocene extinctions. Recently extinct species from these regions are listed in separate articles.

Many extinction dates are unknown due to a lack of relevant information.

Mammals (class Mammalia) Edit

Golden moles, otter shrews, and tenrecs (order Afrosoricida) Edit

Golden moles (family Chrysochloridae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments
De Winton's golden mole Cryptochloris wintoni Port Nolloth, South Africa Not recorded since 1937. Its only known location has suffered some habitat alteration due to port construction and diamond mining. However, the difficulty of detecting golden moles and identifying the different species makes difficult to have an accurate idea of its real presence.[2]

Elephant-like mammals (order Proboscidea) Edit

Elephants and mammoths (family Elephantidae) Edit

Common name Scientific name Range Comments Pictures
North African elephant Loxodonta africana pharaoensis North Africa Neolithic rock paintings reveal that the African bush elephant inhabited much of the Sahara desert and North Africa at the beginning of the Holocene, and Ancient authors indicate that it was present in the Atlas Mountains, the Red Sea coast, and Nubia until the first few centuries AD.[3] It was also present in much of Egypt, except for the Sinai Peninsula, during the Late Paleolithic or early Holocene.[4] However the validity of separate subspecies in Loxodonta africana has been called into question, including the purported North African subspecies L. a. pharaoensis.[3]

Ptolemy II Philadelphus (r. 284-246 BC) founded the cities of Berenice Troglodytica and Ptolemais Theron as hunting bases to provide himself with African raw ivory and war elephants, replacing the costlier imports from India. This caused the depletion of elephant populations along the Red Sea coast and northern Somalia in less than two or three decades. These "Eritrean" elephants were at one point identified as African forest elephants due to Polybius's claim that they were smaller than the Asian elephants they faced at the Battle of Raphia,[5] but genetic analyses show they were bush elephants.[6] In the Roman Empire, the Maghreb and possibly Western Africa through the Garamantes became additional sources of ivory and live elephants for the circus games. The last clear mention of wild elephants in the former is a speech of the orator Themistius delivered in 370 AD,[6] where he mentions that "elephants have been removed from Libya".[b]

 

Rodents (order Rodentia) Edit

Old World rats and mice (family Muridae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments
Ethiopian amphibious rat Nilopegamys plumbeus Little Abbai river, Northwestern Ethiopia Known from, and described from a single specimen captured on March 20, 1927. The species is believed to be semiaquatic due to adaptations shared with aquatic rodents from South America that are not known in other African rodents. If this is correct, this is probably an extremely solitary species as similarly adapted rodents are, which increases the difficulty of detection. However the area where the original individual was captured has been also altered by extensive overgrazing by livestock, which may have caused its decline and extinction.[8][9]

Primates (order Primates) Edit

Lorises, pottos, and angwantibos (family Lorisidae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments
Mount Kenya potto Perodicticus ibeanus stockleyi Mount Kenya, Kenya Known from a single specimen collected from montane forest in 1938.[10]

True insectivores (order Eulipotyphla) Edit

True shrews (family Soricidae) Edit

Common name Scientific name Range Comments
Balsam shrew Crocidura balsamifera Lower Nile, Egypt Described from mummified remains from Ancient Egypt dating to 821-171 BCE. Presumed to have been a swamp or gallery forest specialist whose natural habitat was cleared for agriculture.[11]
Locally extinct Edit
Common name Scientific name Range Comments
Güldenstädt's shrew Crocidura gueldenstaedtii Southern Europe and Western Asia Mummified remains from the beginning of the Ptolemaic Period at Quesna, Egypt indicate that it once occurred in the Nile Delta, where it no longer does, supporting a moister regional environment at the time.[12]

Carnivorans (order Carnivora) Edit

Cats (family Felidae) Edit

Locally extinct Edit
Common name Scientific name Range Comments Pictures
Barbary lion Population of the northern lion (Panthera leo leo) North Africa Lions existed throughout Egypt in ancient times.[4] The last lion in Libya was killed in 1700,[13] in Tunisia in 1891, in Morocco in 1942 on the Tizi-N'Tichka pass of the High Atlas, and in Algeria in 1943. There was an unconfirmed sighting of a lion by the passengers of a bus in a remote wooded area of the Béni Ourtilane District of Algeria in 1956.[14] Despite being the first subspecies named by Linnaeus in the 18th century, modern molecular studies indicate that there is not enough difference with the extant lions of India, western and central Africa to warrant separate subspecies status, and as a result the taxon P. l. leo is not extinct.[15]  
Cape lion Population of the southern lion (Panthera leo melanochaita) South Africa Last individual was killed in KwaZulu-Natal in 1865.[14] Though widely recognized as a subspecies since being named in 1842, modern molecular studies indicate that there is not enough difference with extant lions of southern and eastern Africa to warrant separate subspecies status, and as a result the taxon P. l. melanochaita is not extinct.[15]  
Barbary leopard Population of the African leopard (Panthera pardus pardus) Atlas Mountains Last recorded in 1996.[14] Though named as the subspecies P. p. panthera in 1777, it was later included in P. p. pardus on morphological and molecular grounds.[16]  
Zanzibar leopard Population of the African leopard (Panthera pardus pardus) Unguja, Tanzania Only African insular population of leopards, it was subjected to a extermination campaign after the Zanzibar Revolution of 1964, and the last confirmed sighting happened in 1986.[14] Though named as the subspecies P. p. adersi in 1932, it was included in the African leopard P. p. pardus in 1996 on morphological grounds.[16] There was an unconfirmed recording of a leopard in Unguja in 2018.[14]  

Dogs (family Canidae) Edit

Locally extinct Edit
Common name Scientific name Range Comments Pictures
Gray wolf Canis lupus Eurasia and North America Lived in the Nile Delta in prehistoric times.[4] The African wolf has considerable genetic admixture from the gray wolf.[17]  

Bears (family Ursidae) Edit

Common name Scientific name Range Comments Pictures
Atlas bear Ursus arctos crowtheri Northern Maghreb This subspecies was named after the second-hand description of a female killed in the Rif near Tétouan in 1834 and its pelt, which is now lost.[18] The presence of brown bears in Morocco and Algeria was confirmed with the finding of several bones ranging from the Pleistocene to 662-778 AD,[19] mostly in high mountains around 1200-2000 meters above sea level.[18] Bears were of similar size to the small southern populations of Spain, Italy, and the Middle East. Native knowledge of bears was also documented in Algeria in the 18th century and the beginning of the 19th century.[18] mtDNA studies revealed that two highly distinct lineages of bears existed in North Africa through the Holocene: one identical to Cantabrian brown bears from Spain, and another that was basal to all European brown bears.[c] The North African bear could have disappeared due to increased habitat fragmentation.[18]  
Locally extinct Edit
Common name Scientific name Range Comments Pictures
Syrian bear Ursus arctos syriacus Near East Lived in the Nile Delta in prehistoric times and possibly in northeastern Egypt in early historical times.[4]  

Odd-toed ungulates (order Perissodactyla) Edit

Horses and allies (family Equidae) Edit

Common name Scientific name Range Comments Pictures
Atlas wild ass Equus africanus atlanticus North Africa Disappeared around 300 AD.[21] This subspecies is attributed a distribution in the Atlas region of northern Morocco, Algeria, and Tunisia, although E. africanus is also known from the Holocene of the Sahara,[22] Egypt,[4] and Arabia,[23] leaving aside the extant subspecies E. a. africanus and E. a. somaliensis in Sudan and the Horn of Africa.[22] North African rock art and Roman mosaics show animals with characteristic leg stripes and a shoulder stripe, often doubled, different from the extant subspecies. However, it's been claimed that the name E. a. atlanticus would be unavailable due to improper description of a type specimen.[24][25] Domestic donkeys have two different haplotypes, one shared with the Nubian wild ass, and another of unknown origin that is not found in the Somali wild ass. The presence of the Atlas wild ass in the Ancient world makes it a plausible source for the second haplotype.[22][26]  
North African horse Equus algericus North Africa Most recent remains dated to 4855-4733 BCE at El Harhoura 1, Morocco.[27]  
Giant Cape zebra Equus capensis Southern Africa Most recent remains at Wonderwerk Cave, South Africa were dated to 8120-7980 BCE. Described as the largest equid of the African Quaternary and an extreme hypsodont, its extinction is speculated to be related to the decline in the availability or productivity of grassland habitats since the end of the Last Glacial Maximum.[27] However, ancient DNA studies indicate that the giant Cape zebra is not a separate species, but a distinct lineage of the plains zebra (E. quagga).[28]  
North African zebra Equus mauritanicus North Africa Related to the plains zebra. Disappeared c. 4000 BCE.[27]  
Equus melkiensis Northern Algeria and Morocco Disappeared c. 4000 BCE. Related to the African wild ass[27] and sometimes considered the same as E. a. atlanticus.[21]
Quagga Equus quagga quagga Cape Province, South Africa Last seen in the wild between 1860 and 1865. The last individual died in captivity in Europe in 1883. It was hunted to extinction.[29]  
Possibly extinct Edit
Common name Scientific name Range Comments Pictures
Nubian wild ass Equus africanus africanus Nubian Desert Considered possibly extinct as it has only been infrequently seen since it was sighted in Ethiopia's Barka Valley and Eritrea during the 1970s. The subspecies is threatened by hunting for food and traditional medicine, competition with livestock for vegetation and water, and possibly interbreeding with domestic donkeys.[21] Some haplotypes in domestic donkeys are also found in the Nubian wild ass, either suggesting that domestic donkeys are partially descended from the Nubian wild ass, or that there has been interbreeding between Nubian wild asses and feral donkeys.[22]  

Rhinoceroses (family Rhinocerotidae) Edit

Common name Scientific name Range Comments Pictures
Ceratotherium mauritanicum Northern and eastern Africa Though more known from the Pliocene and Pleistocene, it survived into the early Holocene of Morocco and Tunisia and is commonly depicted in North African rock art hunting scenes up to the Bronze Age. It was extremely similar to the northern white rhinoceros C. simum cottoni in size, proportions, and dentition, and has been treated as its direct ancestor, a subspecies (though cottoni is now recognized as a subspecies of C. simum itself), or synonymous with it.[19][30][31]  
Southern black rhinoceros Diceros bicornis bicornis Southwestern Africa Disappeared from the Cape Colony in the mid-19th century.[32] The IUCN considers the south-western black rhinoceros (D. b. occidentalis) from Namibia and Angola, used to re-stock South Africa, to be the same subspecies. If this is followed, the taxon D. b. bicornis is not extinct.[33]  
Western black rhinoceros Diceros bicornis longipes Burkina Faso to South Sudan An investigation into the last known location in Cameroon in 2006 found abundant evidence of wildlife poaching and no sign of rhinoceroses except that faked by local rhinoceros monitors. There have been no sightings or other evidence afterward.[34]  
Possibly extinct Edit
Common name Scientific name Range Comments Pictures
North-eastern black rhinoceros Diceros bicornis brucii Horn of Africa to eastern Sudan and Bahr el Ghazal Considered probably extinct by 2011.[35]  
Extinct in the wild Edit
Common name Scientific name Range Comments Pictures
Northern white rhinoceros Ceratotherium simum cottoni Upper Chari, Ubangi, and White Nile river basins The last four wild animals were sighted in 2006 and the last indirect sign of their presence was detected in 2007, both under an uptick of poaching in the region.[36] In 2009,[37] the last four captive rhinos were moved from the Safari Park Dvůr Králové in the Czech Republic to a private reserve in Kenya, outside of the subspecies's recent range, but the two males died without breeding. The last remaining individuals are a mother and a daughter and attempts at artificial insemination have been unsuccessful.[36]  

Even-toed ungulates (order Artiodactyla) Edit

Pigs (family Suidae) Edit

Common name Scientific name Range Comments Pictures
Cape warthog Phacochoerus aethiopicus aethiopicus Cape Province, South Africa Last known individual killed in 1871.[38]  

True deer (family Cervidae) Edit

Scientific name Range Comments Pictures
Megaceroides algericus Northern Maghreb Most recent remains dated to 4691-4059 BCE in Bizmoune, Morocco.[39]  
Locally extinct Edit
Common name Scientific name Range Comments Pictures
Persian fallow deer Dama mesopotamica Middle East Deer, known as hnn in the Egyptian language, are depicted in art from the Predynastic to the Ptolemaic period, and remains of Persian fallow deer have been found in archaeological sites of the eastern Nile Delta dating mostly to the 14th-10th centuries BCE. However, the autochthonous nature of these animals is controversial, as is the presence of other deer species like red deer or chital in Ancient Egypt.[4][40][41]  

Cattle, goats, antelopes, and others (family Bovidae) Edit

Common name Scientific name Range Comments Pictures
Bubal hartebeest Alcelaphus buselaphus buselaphus North Africa and southern Levant[42] Last animal in Tunisia was killed in 1902 near Tataouine, in Algeria south of the Chott Ech Chergui in the 1920s, and in Morocco in Missour in 1925.[43] The subspecies was also present in Egypt along the Nile and in the oases of the western desert[4] until the early Middle Ages.[43]  
Southern springbok Antidorcas australis Southern coast of South Africa Most recent remains at Nelson Bay Cave dated to 16810-9850 BCE.[27]
Bond's springbok Antidorcas bondi Southern Africa Most recent remains at Kruger Cave, South Africa dated to 5680-5560 BCE.[27]
North African aurochs Bos primigenius mauritanicus North Africa Wild populations are assumed to have disappeared c. 4000 BCE, though genetic evidence suggests that North African aurochs underwent indigenous domestication near the onset of the Holocene, and that some races of African cattle are descended from it.[27] Of these, the N'Dama, Kuri, and some varieties of West African Shorthorn descend exclusively from the African aurochs, without admixture from Eurasian cattle.[44] The aurochs possibly survived for longer in Egypt, disappearing from the upper Nile in the Predynastic period but surviving in the Delta (Buto) until the Roman era. Hunting, habitat modification for agriculture, and competition with domestic cattle may have caused its decrease in numbers and ultimate disappearance.[45]  
Caprinae indet.
(Makapania?)
South Africa mountains Most recent remains at Colwinton Shelter, South Africa dated to 4360-4280 BCE. The extinction coincided with changes in vegetation leading to the replacement of grazing ungulates for browsers.[27]  
Damaliscus hypsodon Kenya and Tanzania Most recent remains dated to less than 8902-8638 BCE in Kisese II, Tanzania.[27]
Bluebuck Hippotragus leucophaeus Overberg, South Africa Fossil evidence and rock art suggests that the species was more broadly spread around southern Africa in the Pleistocene and early Holocene, but its range contracted because of climate-driven vegetation change until it was reduced to just 4300 km2 east of Cape Town. It finally disappeared around 1800 AD as a result of hunting, competition with livestock, and habitat loss and fragmentation caused by agriculture.[27]  
Roberts' lechwe Kobus leche robertsi Luongo and Kalungwishi drainage systems, Luapula, Zambia Last seen between 1980 and 1985.[46]
Giant wildebeest Megalotragus priscus Southern and possibly eastern Africa Most recent remains at Wonderwerk Cave, South Africa dated to 6442-6210 BCE.[27]  
Kenya oribi Ourebia ourebi kenyae Lower slopes of Mount Kenya, Kenya[10]
African giant buffalo Syncerus antiquus Africa Widespread through the continent in the Pleistocene, it became restricted to North Africa in the Holocene and survived until 3060-2470 BCE. Increased aridification and competition with domestic cattle have both been suggested as causes of its extinction.[11]  
Extinct in the wild Edit
Common name Scientific name Range Comments Pictures
Mohrr gazelle Nanger dama mohrr Northwestern Sahara Disappeared from the wild in 1968, being last seen in Western Sahara.[47] The first reintroduction program began in Senegal in 1984[48] and was followed by others in Morocco and Tunisia. The Tunisian project ended in failure with the death of the last animal in 2020.[47]  
Scimitar oryx Oryx dammah Fringes of the Sahara The last wild population disappeared in Chad between 1988 and 1990.[49] A reintroduction program began in Tunisia in 1985.[50]  
Locally extinct Edit
Common name Scientific name Range Comments Pictures
Arabian oryx Oryx leucoryx Arabian Peninsula Probably lived in the north of Egypt's eastern desert during historical times.[4]  

Right and bowhead whales (family Balaenidae) Edit

Locally extinct Edit
Common name Scientific name Range Comments Pictures
North Atlantic right whale Eubalaena glacialis North Atlantic and western Mediterranean Sea Possibly calved in the Mediterranean in ancient times. Probable remains were found in Roman archaeological sites at Tetouan and Ceuta dated to 180-396 and 226-440 AD, respectively, and an individual was sighted off Algiers in 1888.[51] A calving area existed in Western Sahara in recent times, but was declared extinct in 1998.[52] The species is still present sporadically in Macaronesia, where it visits and possibly calves near the Açores[53] and Canary Islands.[54]  

Gray whales (family Eschrichtiidae) Edit

Locally extinct Edit
Common name Scientific name Range Comments Pictures
Gray whale Eschrichtius robustus North Atlantic, Mediterranean, and northern Pacific Ocean Possibly calved in the Mediterranean in ancient times. Remains were found in Tetouan dating to 71–245 AD.[51] A vagrant from the North Pacific population was seen off the coast of Namibia in May 2013.[55][56]  

Birds (class Aves) Edit

Landfowl (order Galliformes) Edit

Guineafowl (family Numididae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments Pictures
Moroccan guineafowl Numida meleagris sabyi Between the Oum er Rbia and Sebou rivers of Morocco Last recorded with certainty in the wild in the 1950s. It succumbed to habitat destruction and over-hunting. Reports of a captive population in the 1980s are unsubstantiated.[57]  

Bustards (order Otidiformes) Edit

Bustards (family Otididae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments Pictures
Moroccan bustard Ardeotis arabs lynesi Western Morocco Last recorded at Lakes Merzouga and Tamezguidat between 1987 and 1993. All Arabian bustard subspecies declined due to hunting and habitat destruction.[57]  

Shorebirds (order Charadriiformes) Edit

Oystercatchers (family Haematopodidae) Edit

Common name Scientific name Range Comments Pictures
Canary Islands oystercatcher Haematopus meadewaldoi Canary Islands to the coast of Senegal Last recorded in Senegal between 1968 and 1981. Its decline was probably a result of overharvesting of intertidal invertebrates and disturbance by people, although predation by rats and cats has also been implicated.[58]  

Sandpipers (order Scolopacidae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments Pictures
Slender-billed curlew Numenius tenuirostris North Africa and Western Eurasia The species breeds in Central Asia (the steppes of northern Kazakhstan and southern central Russia) and winters in the Mediterranean area and south Arabia,[59] but has declined due to intense hunting in the wintering grounds and habitat destruction in the breeding grounds. Slender-billed curlews were regular visitors to Merja Zerga, Morocco until 1995.[57]  

Auks (family Alcidae) Edit

Common name Scientific name Range Comments Pictures
Great auk Pinguinus impennis Northern Atlantic and western Mediterranean A bone found in El Harhoura 2, Morocco was dated to 5050-3850 BCE.[60] This is the second southernmost record of this species in the eastern Atlantic, after another bone from Madeira.[61] The species became extinct globally in 1852.[62]  

Hawks and relatives (order Accipitriformes) Edit

Hawks, eagles, kites, harriers and Old World vultures (family Accipitridae) Edit

Locally extinct Edit
Common name Scientific name Range Comments Pictures
Spanish imperial eagle Aquila adalberti Southwestern Iberia and northwestern Morocco[63] Could have disappeared as a breeder from Morocco before 1950,[64] though two adult pairs were seen in Tassaoti, Oued Laou and the mouth of the Moulouya river in 1977. Vagrant juveniles still visit the northern part of the country from the Guadalquivir marshes[65] and are sometimes killed in unprotected power lines.[64]  

Passerines (order Passeriformes) Edit

Cisticolas and allies (family Cisticolidae) Edit

Common name Scientific name Range Comments
Northern white-winged apalis Apalis chariessa chariessa Lower Tana River, Kenya Last recorded in 1961, when the forests of Mitole were cleared.[57]

Reptiles (class Reptilia) Edit

Squamates (order Squamata) Edit

Plated lizards (family Gerrhosauridae) Edit

Common name Scientific name Range Comments
Eastwood's long-tailed seps Tetradactylus eastwoodae Limpopo, South Africa Last seen in 1928. Its natural habitat was destroyed by afforestation.[66]

Amphibians (class Amphibia) Edit

Frogs (order Anura) Edit

African torrent frogs (family Petropedetidae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments
Du Toit's torrent frog Arthroleptides dutoiti Kenya-Uganda border Last recorded in 1962. It might have disappeared due to chytridiomycosis.[67]

True toads (family Bufonidae) Edit

Possibly extinct Edit
Common name Scientific name Range Comments
Osgood's Ethiopian toad Altiphrynoides osgoodi Mountains of south-central Ethiopia Last recorded in 2003.[68]
Extinct in the wild Edit
Common name Scientific name Range Comments Pictures
Kihansi spray toad Nectophrynoides asperginis Kihansi Falls, Udzungwa Mountains, Tanzania Last recorded in the wild in 2004, with an unconfirmed report in 2005. The species declined due to drought, chytridiomycosis, pesticide use in maize agriculture, and possibly other causes. Nevertheless, thousands exist in captivity and a reintroduction program began with large numbers in 2012.[69]  

Ray-finned fish (class Actinopterygii) Edit

Minnows and allies (order Cypriniformes) Edit

Carps, minnows, and relatives (family Cyprinidae) Edit

Common name Scientific name Range Comments Pictures
Labeobarbus microbarbis Lake Luhondo, Rwanda Known from a single individual collected c. 1937, it is presumed to have become extinct in the 1950s after the introduction of Tilapia and Haplochromis to the lake. However the validity of the species is doubtful and could be a hybrid of Barbus and Varicorhinus instead.[70]
Giant Atlas barbel Labeobarbus reinii Northwestern Morocco Last recorded in 2001. The rivers it inhabited have been affected by pollution and damming, but the precise causes of extinction are poorly understood.[71]  
Tunisian barb Luciobarbus antinorii Chott el Djerid, Tunisia Last collected in 1989. It could have disappeared due to excessive water substraction.[72]
Luciobarbus nasus Ksob river drainage, Morocco Last recorded in 1874. The river has been affected by pollution and damming, but the precise causes of extinction are poorly understood.[73]  

Salmon, trout and relatives (order Salmoniformes) Edit

Salmon, trout and relatives (family Salmonidae) Edit

Common name Scientific name Range Comments
Lake Sidi Ali trout Salmo pallaryi Lake Aguelmame Sidi Ali, Morocco Disappeared in 1934 after the introduction of the Eurasian carp.[74]

Toothcarps (order Cyprinodontiformes) Edit

Livebearers and relatives (family Poeciliidae) Edit

Scientific name Range Comments
Aplocheilichthys sp. nov. 'Naivasha' Lake Naivasha, Kenya Disappeared in the 1970s or 1980s due to competition with introduced fishes.[75]

Insects (class Insecta) Edit

Butterflies and moths (order Lepidoptera) Edit

Gossamer-winged butterflies (family Lycaenidae) Edit

Common name Scientific name Range Comments
Mbashe River buff Deloneura immaculata Mbhashe River, Eastern Cape, South Africa Only known from three individuals collected "at the end of December 1863".[76]
Morant's blue Lepidochrysops hypopolia Eastern South Africa Only recorded in the 1870s.[77]

Ostracods (class Ostracoda) Edit

Order Podocopida Edit

Family Candonidae Edit

Scientific name Range Comments Pictures
Namibcypris costata Southern Kaokoveld, Namibia Last recorded in 1987.[78]  

See also Edit

Notes Edit

  1. ^ The source gives "11,700 calendar yr b2k (before AD 2000)". But "BP" means "before AD 1950". Therefore, the Holocene began 11,650 BP. Doing the math, that is c. 9700 BCE.
  2. ^ "...and we are displeased because elephants have been removed from Libya, because lions have disappeared from Thessaly, because hippopotamoi have been gotten rid from the marshes of the Nile."[7]
  3. ^ This study did not use Syrian brown bears for comparison.[20]

References Edit

  1. ^ Walker, Mike; Johnsen, Sigfus; Rasmussen, Sune Olander; Popp, Trevor; Steffensen, Jorgen-Peder; Gibrard, Phil; Hoek, Wim; Lowe, John; Andrews, John; Bjo Rck, Svante; Cwynar, Les C.; Hughen, Konrad; Kersahw, Peter; Kromer, Bernd; Litt, Thomas; Lowe, David J.; Nakagawa, Takeshi; Newnham, Rewi; Schwander, Jakob (2009). "Formal definition and dating of the GSSP (Global Stratotype Section and Point) for the base of the Holocene using the Greenland NGRIP ice core, and selected auxiliary records" (PDF). Journal of Quaternary Science. 24 (1): 3–17. Bibcode:2009JQS....24....3W. doi:10.1002/jqs.1227. (PDF) from the original on 2013-11-04. Retrieved 2022-04-24.
  2. ^ Bronner, G. (2015). "Cryptochloris wintoni". IUCN Red List of Threatened Species. 2015: e.T5748A21287143. doi:10.2305/IUCN.UK.2015-2.RLTS.T5748A21287143.en. Retrieved 12 November 2021.
  3. ^ a b Happold, D.C.D. & Kalina, J., Ed. (2013) Mammals of Africa: Introductory chapters and Afrotheria. Bloomsbury, 352 pages.
  4. ^ a b c d e f g h Riemer, H. & Pöllath, N. (2007) Desert animals in the eastern Sahara: status, economic significance, and cultural reflection in antiquity. Proceedings of an interdisciplinary ACACIA workshop held at the University of Cologne December 14-15, 2007
  5. ^ Gowers, William (1948). "African Elephants and Ancient Authors". African Affairs. 47 (188): 173–180. doi:10.1093/oxfordjournals.afraf.a093647. JSTOR 718306.
  6. ^ a b BollóK, Á., & Koncz, I. (2020). Sixth- and Seventh-Century Elephant Ivory Finds from the Carpathian Basin. The Sources, Circulation and Value of Ivory in Late Antiquity and the Early Middle Ages. Archaeologiai Értesítő, Vol. 1: 39-68.
  7. ^ Braddock, A.C. (2023) Implication: An ecocritical dictionary of art history. Yale University Press, 256 pages.
  8. ^ "Scientific Illustration". Scientific Illustration. Retrieved 2019-04-18.
  9. ^ Kerbis Peterhans, J. & Lavrenchenko, L. (2008). "Nilopegamys plumbeus". IUCN Red List of Threatened Species. 2008: e.T40766A10363474. doi:10.2305/IUCN.UK.2008.RLTS.T40766A10363474.en.
  10. ^ a b Richardson, M. (2023) Threatened and recently extinct vertebrates of the world: A biogeographic approach. Cambridge University Press, 750 pages.
  11. ^ a b Turvey, Sam (2009). Holocene extinctions. Oxford University Press. ISBN 978-0-19-953509-5. Retrieved 29 February 2012.
  12. ^ Woodman, Neal; Ikram, Salima; Rowland, Joanne (2021-04-07). "A new addition to the embalmed fauna of ancient Egypt: Güldenstaedt's White-toothed Shrew, Crocidura gueldenstaedtii (Pallas, 1811) (Mammalia: Eulipotyphla: Soricidae)". PLOS ONE. 16 (4): e0249377. Bibcode:2021PLoSO..1649377W. doi:10.1371/journal.pone.0249377. ISSN 1932-6203. PMC 8026016. PMID 33826664.
  13. ^ Bartosiewicz, L. (2009) A lion’s share of attention: archaeozoology and the historical record. Acta Archaeologica, 60(1), 275-289.
  14. ^ a b c d e Rossi, L., Scuzzarella, C. M., & Angelici, F. M. (2020). "Extinct or Perhaps Surviving Relict Populations of Big Cats: Their Controversial Stories and Implications for Conservation". In Problematic Wildlife II (pp. 393-417). Springer, Cham.
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External links Edit

  • IUCN Red List of Threatened Species

list, african, animals, extinct, holocene, this, list, african, species, extinct, holocene, covers, extinctions, from, holocene, epoch, geologic, epoch, that, began, about, years, before, present, about, 9700, continues, present, africathe, quagga, equus, quag. This list of African species extinct in the Holocene covers extinctions from the Holocene epoch a geologic epoch that began about 11 650 years before present about 9700 BCE a and continues to the present 1 Map of AfricaThe quagga Equus quagga quagga extinct since 1883 was zebra like in the front but more horse like in the rear A breeding program aims to create similar looking animals but these are not true quaggas Africa is highly biodiverse it is the continent with the largest number of megafauna species as it was least affected by the extinction of the Pleistocene megafauna However a few species have disappeared from Africa as part of the ongoing Holocene extinction driven by human activity Madagascar and the Indian Ocean islands Macaronesia and Saint Helena Ascension and Tristan da Cunha are biogeographically distinct from mainland Africa and have a much greater number of Holocene extinctions Recently extinct species from these regions are listed in separate articles Many extinction dates are unknown due to a lack of relevant information Contents 1 Mammals class Mammalia 1 1 Golden moles otter shrews and tenrecs order Afrosoricida 1 1 1 Golden moles family Chrysochloridae 1 1 1 1 Possibly extinct 1 2 Elephant like mammals order Proboscidea 1 2 1 Elephants and mammoths family Elephantidae 1 3 Rodents order Rodentia 1 3 1 Old World rats and mice family Muridae 1 3 1 1 Possibly extinct 1 4 Primates order Primates 1 4 1 Lorises pottos and angwantibos family Lorisidae 1 4 1 1 Possibly extinct 1 5 True insectivores order Eulipotyphla 1 5 1 True shrews family Soricidae 1 5 1 1 Locally extinct 1 6 Carnivorans order Carnivora 1 6 1 Cats family Felidae 1 6 1 1 Locally extinct 1 6 2 Dogs family Canidae 1 6 2 1 Locally extinct 1 6 3 Bears family Ursidae 1 6 3 1 Locally extinct 1 7 Odd toed ungulates order Perissodactyla 1 7 1 Horses and allies family Equidae 1 7 1 1 Possibly extinct 1 7 2 Rhinoceroses family Rhinocerotidae 1 7 2 1 Possibly extinct 1 7 2 2 Extinct in the wild 1 8 Even toed ungulates order Artiodactyla 1 8 1 Pigs family Suidae 1 8 2 True deer family Cervidae 1 8 2 1 Locally extinct 1 8 3 Cattle goats antelopes and others family Bovidae 1 8 3 1 Extinct in the wild 1 8 3 2 Locally extinct 1 8 4 Right and bowhead whales family Balaenidae 1 8 4 1 Locally extinct 1 8 5 Gray whales family Eschrichtiidae 1 8 5 1 Locally extinct 2 Birds class Aves 2 1 Landfowl order Galliformes 2 1 1 Guineafowl family Numididae 2 1 1 1 Possibly extinct 2 2 Bustards order Otidiformes 2 2 1 Bustards family Otididae 2 2 1 1 Possibly extinct 2 3 Shorebirds order Charadriiformes 2 3 1 Oystercatchers family Haematopodidae 2 3 2 Sandpipers order Scolopacidae 2 3 2 1 Possibly extinct 2 3 3 Auks family Alcidae 2 4 Hawks and relatives order Accipitriformes 2 4 1 Hawks eagles kites harriers and Old World vultures family Accipitridae 2 4 1 1 Locally extinct 2 5 Passerines order Passeriformes 2 5 1 Cisticolas and allies family Cisticolidae 3 Reptiles class Reptilia 3 1 Squamates order Squamata 3 1 1 Plated lizards family Gerrhosauridae 4 Amphibians class Amphibia 4 1 Frogs order Anura 4 1 1 African torrent frogs family Petropedetidae 4 1 1 1 Possibly extinct 4 1 2 True toads family Bufonidae 4 1 2 1 Possibly extinct 4 1 2 2 Extinct in the wild 5 Ray finned fish class Actinopterygii 5 1 Minnows and allies order Cypriniformes 5 1 1 Carps minnows and relatives family Cyprinidae 5 2 Salmon trout and relatives order Salmoniformes 5 2 1 Salmon trout and relatives family Salmonidae 5 3 Toothcarps order Cyprinodontiformes 5 3 1 Livebearers and relatives family Poeciliidae 6 Insects class Insecta 6 1 Butterflies and moths order Lepidoptera 6 1 1 Gossamer winged butterflies family Lycaenidae 7 Ostracods class Ostracoda 7 1 Order Podocopida 7 1 1 Family Candonidae 8 See also 9 Notes 10 References 11 External linksMammals class Mammalia EditGolden moles otter shrews and tenrecs order Afrosoricida Edit Golden moles family Chrysochloridae Edit Possibly extinct Edit Common name Scientific name Range CommentsDe Winton s golden mole Cryptochloris wintoni Port Nolloth South Africa Not recorded since 1937 Its only known location has suffered some habitat alteration due to port construction and diamond mining However the difficulty of detecting golden moles and identifying the different species makes difficult to have an accurate idea of its real presence 2 Elephant like mammals order Proboscidea Edit Elephants and mammoths family Elephantidae Edit Common name Scientific name Range Comments PicturesNorth African elephant Loxodonta africana pharaoensis North Africa Neolithic rock paintings reveal that the African bush elephant inhabited much of the Sahara desert and North Africa at the beginning of the Holocene and Ancient authors indicate that it was present in the Atlas Mountains the Red Sea coast and Nubia until the first few centuries AD 3 It was also present in much of Egypt except for the Sinai Peninsula during the Late Paleolithic or early Holocene 4 However the validity of separate subspecies in Loxodonta africana has been called into question including the purported North African subspecies L a pharaoensis 3 Ptolemy II Philadelphus r 284 246 BC founded the cities of Berenice Troglodytica and Ptolemais Theron as hunting bases to provide himself with African raw ivory and war elephants replacing the costlier imports from India This caused the depletion of elephant populations along the Red Sea coast and northern Somalia in less than two or three decades These Eritrean elephants were at one point identified as African forest elephants due to Polybius s claim that they were smaller than the Asian elephants they faced at the Battle of Raphia 5 but genetic analyses show they were bush elephants 6 In the Roman Empire the Maghreb and possibly Western Africa through the Garamantes became additional sources of ivory and live elephants for the circus games The last clear mention of wild elephants in the former is a speech of the orator Themistius delivered in 370 AD 6 where he mentions that elephants have been removed from Libya b nbsp Rodents order Rodentia Edit Old World rats and mice family Muridae Edit Possibly extinct Edit Common name Scientific name Range CommentsEthiopian amphibious rat Nilopegamys plumbeus Little Abbai river Northwestern Ethiopia Known from and described from a single specimen captured on March 20 1927 The species is believed to be semiaquatic due to adaptations shared with aquatic rodents from South America that are not known in other African rodents If this is correct this is probably an extremely solitary species as similarly adapted rodents are which increases the difficulty of detection However the area where the original individual was captured has been also altered by extensive overgrazing by livestock which may have caused its decline and extinction 8 9 Primates order Primates Edit Lorises pottos and angwantibos family Lorisidae Edit Possibly extinct Edit Common name Scientific name Range CommentsMount Kenya potto Perodicticus ibeanus stockleyi Mount Kenya Kenya Known from a single specimen collected from montane forest in 1938 10 True insectivores order Eulipotyphla Edit True shrews family Soricidae Edit Common name Scientific name Range CommentsBalsam shrew Crocidura balsamifera Lower Nile Egypt Described from mummified remains from Ancient Egypt dating to 821 171 BCE Presumed to have been a swamp or gallery forest specialist whose natural habitat was cleared for agriculture 11 Locally extinct Edit Common name Scientific name Range CommentsGuldenstadt s shrew Crocidura gueldenstaedtii Southern Europe and Western Asia Mummified remains from the beginning of the Ptolemaic Period at Quesna Egypt indicate that it once occurred in the Nile Delta where it no longer does supporting a moister regional environment at the time 12 Carnivorans order Carnivora Edit Cats family Felidae Edit Locally extinct Edit Common name Scientific name Range Comments PicturesBarbary lion Population of the northern lion Panthera leo leo North Africa Lions existed throughout Egypt in ancient times 4 The last lion in Libya was killed in 1700 13 in Tunisia in 1891 in Morocco in 1942 on the Tizi N Tichka pass of the High Atlas and in Algeria in 1943 There was an unconfirmed sighting of a lion by the passengers of a bus in a remote wooded area of the Beni Ourtilane District of Algeria in 1956 14 Despite being the first subspecies named by Linnaeus in the 18th century modern molecular studies indicate that there is not enough difference with the extant lions of India western and central Africa to warrant separate subspecies status and as a result the taxon P l leo is not extinct 15 nbsp Cape lion Population of the southern lion Panthera leo melanochaita South Africa Last individual was killed in KwaZulu Natal in 1865 14 Though widely recognized as a subspecies since being named in 1842 modern molecular studies indicate that there is not enough difference with extant lions of southern and eastern Africa to warrant separate subspecies status and as a result the taxon P l melanochaita is not extinct 15 nbsp Barbary leopard Population of the African leopard Panthera pardus pardus Atlas Mountains Last recorded in 1996 14 Though named as the subspecies P p panthera in 1777 it was later included in P p pardus on morphological and molecular grounds 16 nbsp Zanzibar leopard Population of the African leopard Panthera pardus pardus Unguja Tanzania Only African insular population of leopards it was subjected to a extermination campaign after the Zanzibar Revolution of 1964 and the last confirmed sighting happened in 1986 14 Though named as the subspecies P p adersi in 1932 it was included in the African leopard P p pardus in 1996 on morphological grounds 16 There was an unconfirmed recording of a leopard in Unguja in 2018 14 nbsp Dogs family Canidae Edit Locally extinct Edit Common name Scientific name Range Comments PicturesGray wolf Canis lupus Eurasia and North America Lived in the Nile Delta in prehistoric times 4 The African wolf has considerable genetic admixture from the gray wolf 17 nbsp Bears family Ursidae Edit Common name Scientific name Range Comments PicturesAtlas bear Ursus arctos crowtheri Northern Maghreb This subspecies was named after the second hand description of a female killed in the Rif near Tetouan in 1834 and its pelt which is now lost 18 The presence of brown bears in Morocco and Algeria was confirmed with the finding of several bones ranging from the Pleistocene to 662 778 AD 19 mostly in high mountains around 1200 2000 meters above sea level 18 Bears were of similar size to the small southern populations of Spain Italy and the Middle East Native knowledge of bears was also documented in Algeria in the 18th century and the beginning of the 19th century 18 mtDNA studies revealed that two highly distinct lineages of bears existed in North Africa through the Holocene one identical to Cantabrian brown bears from Spain and another that was basal to all European brown bears c The North African bear could have disappeared due to increased habitat fragmentation 18 nbsp Locally extinct Edit Common name Scientific name Range Comments PicturesSyrian bear Ursus arctos syriacus Near East Lived in the Nile Delta in prehistoric times and possibly in northeastern Egypt in early historical times 4 nbsp Odd toed ungulates order Perissodactyla Edit Horses and allies family Equidae Edit Common name Scientific name Range Comments PicturesAtlas wild ass Equus africanus atlanticus North Africa Disappeared around 300 AD 21 This subspecies is attributed a distribution in the Atlas region of northern Morocco Algeria and Tunisia although E africanus is also known from the Holocene of the Sahara 22 Egypt 4 and Arabia 23 leaving aside the extant subspecies E a africanus and E a somaliensis in Sudan and the Horn of Africa 22 North African rock art and Roman mosaics show animals with characteristic leg stripes and a shoulder stripe often doubled different from the extant subspecies However it s been claimed that the name E a atlanticus would be unavailable due to improper description of a type specimen 24 25 Domestic donkeys have two different haplotypes one shared with the Nubian wild ass and another of unknown origin that is not found in the Somali wild ass The presence of the Atlas wild ass in the Ancient world makes it a plausible source for the second haplotype 22 26 nbsp North African horse Equus algericus North Africa Most recent remains dated to 4855 4733 BCE at El Harhoura 1 Morocco 27 nbsp Giant Cape zebra Equus capensis Southern Africa Most recent remains at Wonderwerk Cave South Africa were dated to 8120 7980 BCE Described as the largest equid of the African Quaternary and an extreme hypsodont its extinction is speculated to be related to the decline in the availability or productivity of grassland habitats since the end of the Last Glacial Maximum 27 However ancient DNA studies indicate that the giant Cape zebra is not a separate species but a distinct lineage of the plains zebra E quagga 28 nbsp North African zebra Equus mauritanicus North Africa Related to the plains zebra Disappeared c 4000 BCE 27 nbsp Equus melkiensis Northern Algeria and Morocco Disappeared c 4000 BCE Related to the African wild ass 27 and sometimes considered the same as E a atlanticus 21 Quagga Equus quagga quagga Cape Province South Africa Last seen in the wild between 1860 and 1865 The last individual died in captivity in Europe in 1883 It was hunted to extinction 29 nbsp Possibly extinct Edit Common name Scientific name Range Comments PicturesNubian wild ass Equus africanus africanus Nubian Desert Considered possibly extinct as it has only been infrequently seen since it was sighted in Ethiopia s Barka Valley and Eritrea during the 1970s The subspecies is threatened by hunting for food and traditional medicine competition with livestock for vegetation and water and possibly interbreeding with domestic donkeys 21 Some haplotypes in domestic donkeys are also found in the Nubian wild ass either suggesting that domestic donkeys are partially descended from the Nubian wild ass or that there has been interbreeding between Nubian wild asses and feral donkeys 22 nbsp Rhinoceroses family Rhinocerotidae Edit Common name Scientific name Range Comments PicturesCeratotherium mauritanicum Northern and eastern Africa Though more known from the Pliocene and Pleistocene it survived into the early Holocene of Morocco and Tunisia and is commonly depicted in North African rock art hunting scenes up to the Bronze Age It was extremely similar to the northern white rhinoceros C simum cottoni in size proportions and dentition and has been treated as its direct ancestor a subspecies though cottoni is now recognized as a subspecies of C simum itself or synonymous with it 19 30 31 nbsp Southern black rhinoceros Diceros bicornis bicornis Southwestern Africa Disappeared from the Cape Colony in the mid 19th century 32 The IUCN considers the south western black rhinoceros D b occidentalis from Namibia and Angola used to re stock South Africa to be the same subspecies If this is followed the taxon D b bicornis is not extinct 33 nbsp Western black rhinoceros Diceros bicornis longipes Burkina Faso to South Sudan An investigation into the last known location in Cameroon in 2006 found abundant evidence of wildlife poaching and no sign of rhinoceroses except that faked by local rhinoceros monitors There have been no sightings or other evidence afterward 34 nbsp Possibly extinct Edit Common name Scientific name Range Comments PicturesNorth eastern black rhinoceros Diceros bicornis brucii Horn of Africa to eastern Sudan and Bahr el Ghazal Considered probably extinct by 2011 35 nbsp Extinct in the wild Edit Common name Scientific name Range Comments PicturesNorthern white rhinoceros Ceratotherium simum cottoni Upper Chari Ubangi and White Nile river basins The last four wild animals were sighted in 2006 and the last indirect sign of their presence was detected in 2007 both under an uptick of poaching in the region 36 In 2009 37 the last four captive rhinos were moved from the Safari Park Dvur Kralove in the Czech Republic to a private reserve in Kenya outside of the subspecies s recent range but the two males died without breeding The last remaining individuals are a mother and a daughter and attempts at artificial insemination have been unsuccessful 36 nbsp Even toed ungulates order Artiodactyla Edit Pigs family Suidae Edit Common name Scientific name Range Comments PicturesCape warthog Phacochoerus aethiopicus aethiopicus Cape Province South Africa Last known individual killed in 1871 38 nbsp True deer family Cervidae Edit Scientific name Range Comments PicturesMegaceroides algericus Northern Maghreb Most recent remains dated to 4691 4059 BCE in Bizmoune Morocco 39 nbsp Locally extinct Edit Common name Scientific name Range Comments PicturesPersian fallow deer Dama mesopotamica Middle East Deer known as hnn in the Egyptian language are depicted in art from the Predynastic to the Ptolemaic period and remains of Persian fallow deer have been found in archaeological sites of the eastern Nile Delta dating mostly to the 14th 10th centuries BCE However the autochthonous nature of these animals is controversial as is the presence of other deer species like red deer or chital in Ancient Egypt 4 40 41 nbsp Cattle goats antelopes and others family Bovidae Edit Common name Scientific name Range Comments PicturesBubal hartebeest Alcelaphus buselaphus buselaphus North Africa and southern Levant 42 Last animal in Tunisia was killed in 1902 near Tataouine in Algeria south of the Chott Ech Chergui in the 1920s and in Morocco in Missour in 1925 43 The subspecies was also present in Egypt along the Nile and in the oases of the western desert 4 until the early Middle Ages 43 nbsp Southern springbok Antidorcas australis Southern coast of South Africa Most recent remains at Nelson Bay Cave dated to 16810 9850 BCE 27 Bond s springbok Antidorcas bondi Southern Africa Most recent remains at Kruger Cave South Africa dated to 5680 5560 BCE 27 North African aurochs Bos primigenius mauritanicus North Africa Wild populations are assumed to have disappeared c 4000 BCE though genetic evidence suggests that North African aurochs underwent indigenous domestication near the onset of the Holocene and that some races of African cattle are descended from it 27 Of these the N Dama Kuri and some varieties of West African Shorthorn descend exclusively from the African aurochs without admixture from Eurasian cattle 44 The aurochs possibly survived for longer in Egypt disappearing from the upper Nile in the Predynastic period but surviving in the Delta Buto until the Roman era Hunting habitat modification for agriculture and competition with domestic cattle may have caused its decrease in numbers and ultimate disappearance 45 nbsp Caprinae indet Makapania South Africa mountains Most recent remains at Colwinton Shelter South Africa dated to 4360 4280 BCE The extinction coincided with changes in vegetation leading to the replacement of grazing ungulates for browsers 27 nbsp Damaliscus hypsodon Kenya and Tanzania Most recent remains dated to less than 8902 8638 BCE in Kisese II Tanzania 27 Bluebuck Hippotragus leucophaeus Overberg South Africa Fossil evidence and rock art suggests that the species was more broadly spread around southern Africa in the Pleistocene and early Holocene but its range contracted because of climate driven vegetation change until it was reduced to just 4300 km2 east of Cape Town It finally disappeared around 1800 AD as a result of hunting competition with livestock and habitat loss and fragmentation caused by agriculture 27 nbsp Roberts lechwe Kobus leche robertsi Luongo and Kalungwishi drainage systems Luapula Zambia Last seen between 1980 and 1985 46 Giant wildebeest Megalotragus priscus Southern and possibly eastern Africa Most recent remains at Wonderwerk Cave South Africa dated to 6442 6210 BCE 27 nbsp Kenya oribi Ourebia ourebi kenyae Lower slopes of Mount Kenya Kenya 10 African giant buffalo Syncerus antiquus Africa Widespread through the continent in the Pleistocene it became restricted to North Africa in the Holocene and survived until 3060 2470 BCE Increased aridification and competition with domestic cattle have both been suggested as causes of its extinction 11 nbsp Extinct in the wild Edit Common name Scientific name Range Comments PicturesMohrr gazelle Nanger dama mohrr Northwestern Sahara Disappeared from the wild in 1968 being last seen in Western Sahara 47 The first reintroduction program began in Senegal in 1984 48 and was followed by others in Morocco and Tunisia The Tunisian project ended in failure with the death of the last animal in 2020 47 nbsp Scimitar oryx Oryx dammah Fringes of the Sahara The last wild population disappeared in Chad between 1988 and 1990 49 A reintroduction program began in Tunisia in 1985 50 nbsp Locally extinct Edit Common name Scientific name Range Comments PicturesArabian oryx Oryx leucoryx Arabian Peninsula Probably lived in the north of Egypt s eastern desert during historical times 4 nbsp Right and bowhead whales family Balaenidae Edit Locally extinct Edit Common name Scientific name Range Comments PicturesNorth Atlantic right whale Eubalaena glacialis North Atlantic and western Mediterranean Sea Possibly calved in the Mediterranean in ancient times Probable remains were found in Roman archaeological sites at Tetouan and Ceuta dated to 180 396 and 226 440 AD respectively and an individual was sighted off Algiers in 1888 51 A calving area existed in Western Sahara in recent times but was declared extinct in 1998 52 The species is still present sporadically in Macaronesia where it visits and possibly calves near the Acores 53 and Canary Islands 54 nbsp Gray whales family Eschrichtiidae Edit Locally extinct Edit Common name Scientific name Range Comments PicturesGray whale Eschrichtius robustus North Atlantic Mediterranean and northern Pacific Ocean Possibly calved in the Mediterranean in ancient times Remains were found in Tetouan dating to 71 245 AD 51 A vagrant from the North Pacific population was seen off the coast of Namibia in May 2013 55 56 nbsp Birds class Aves EditLandfowl order Galliformes Edit Guineafowl family Numididae Edit Possibly extinct Edit Common name Scientific name Range Comments PicturesMoroccan guineafowl Numida meleagris sabyi Between the Oum er Rbia and Sebou rivers of Morocco Last recorded with certainty in the wild in the 1950s It succumbed to habitat destruction and over hunting Reports of a captive population in the 1980s are unsubstantiated 57 nbsp Bustards order Otidiformes Edit Bustards family Otididae Edit Possibly extinct Edit Common name Scientific name Range Comments PicturesMoroccan bustard Ardeotis arabs lynesi Western Morocco Last recorded at Lakes Merzouga and Tamezguidat between 1987 and 1993 All Arabian bustard subspecies declined due to hunting and habitat destruction 57 nbsp Shorebirds order Charadriiformes Edit Oystercatchers family Haematopodidae Edit Common name Scientific name Range Comments PicturesCanary Islands oystercatcher Haematopus meadewaldoi Canary Islands to the coast of Senegal Last recorded in Senegal between 1968 and 1981 Its decline was probably a result of overharvesting of intertidal invertebrates and disturbance by people although predation by rats and cats has also been implicated 58 nbsp Sandpipers order Scolopacidae Edit Possibly extinct Edit Common name Scientific name Range Comments PicturesSlender billed curlew Numenius tenuirostris North Africa and Western Eurasia The species breeds in Central Asia the steppes of northern Kazakhstan and southern central Russia and winters in the Mediterranean area and south Arabia 59 but has declined due to intense hunting in the wintering grounds and habitat destruction in the breeding grounds Slender billed curlews were regular visitors to Merja Zerga Morocco until 1995 57 nbsp Auks family Alcidae Edit Common name Scientific name Range Comments PicturesGreat auk Pinguinus impennis Northern Atlantic and western Mediterranean A bone found in El Harhoura 2 Morocco was dated to 5050 3850 BCE 60 This is the second southernmost record of this species in the eastern Atlantic after another bone from Madeira 61 The species became extinct globally in 1852 62 nbsp Hawks and relatives order Accipitriformes Edit Hawks eagles kites harriers and Old World vultures family Accipitridae Edit Locally extinct Edit Common name Scientific name Range Comments PicturesSpanish imperial eagle Aquila adalberti Southwestern Iberia and northwestern Morocco 63 Could have disappeared as a breeder from Morocco before 1950 64 though two adult pairs were seen in Tassaoti Oued Laou and the mouth of the Moulouya river in 1977 Vagrant juveniles still visit the northern part of the country from the Guadalquivir marshes 65 and are sometimes killed in unprotected power lines 64 nbsp Passerines order Passeriformes Edit Cisticolas and allies family Cisticolidae Edit Common name Scientific name Range CommentsNorthern white winged apalis Apalis chariessa chariessa Lower Tana River Kenya Last recorded in 1961 when the forests of Mitole were cleared 57 Reptiles class Reptilia EditSquamates order Squamata Edit Plated lizards family Gerrhosauridae Edit Common name Scientific name Range CommentsEastwood s long tailed seps Tetradactylus eastwoodae Limpopo South Africa Last seen in 1928 Its natural habitat was destroyed by afforestation 66 Amphibians class Amphibia EditFrogs order Anura Edit African torrent frogs family Petropedetidae Edit Possibly extinct Edit Common name Scientific name Range CommentsDu Toit s torrent frog Arthroleptides dutoiti Kenya Uganda border Last recorded in 1962 It might have disappeared due to chytridiomycosis 67 True toads family Bufonidae Edit Possibly extinct Edit Common name Scientific name Range CommentsOsgood s Ethiopian toad Altiphrynoides osgoodi Mountains of south central Ethiopia Last recorded in 2003 68 Extinct in the wild Edit Common name Scientific name Range Comments PicturesKihansi spray toad Nectophrynoides asperginis Kihansi Falls Udzungwa Mountains Tanzania Last recorded in the wild in 2004 with an unconfirmed report in 2005 The species declined due to drought chytridiomycosis pesticide use in maize agriculture and possibly other causes Nevertheless thousands exist in captivity and a reintroduction program began with large numbers in 2012 69 nbsp Ray finned fish class Actinopterygii EditMinnows and allies order Cypriniformes Edit Carps minnows and relatives family Cyprinidae Edit Common name Scientific name Range Comments PicturesLabeobarbus microbarbis Lake Luhondo Rwanda Known from a single individual collected c 1937 it is presumed to have become extinct in the 1950s after the introduction of Tilapia and Haplochromis to the lake However the validity of the species is doubtful and could be a hybrid of Barbus and Varicorhinus instead 70 Giant Atlas barbel Labeobarbus reinii Northwestern Morocco Last recorded in 2001 The rivers it inhabited have been affected by pollution and damming but the precise causes of extinction are poorly understood 71 nbsp Tunisian barb Luciobarbus antinorii Chott el Djerid Tunisia Last collected in 1989 It could have disappeared due to excessive water substraction 72 Luciobarbus nasus Ksob river drainage Morocco Last recorded in 1874 The river has been affected by pollution and damming but the precise causes of extinction are poorly understood 73 nbsp Salmon trout and relatives order Salmoniformes Edit Salmon trout and relatives family Salmonidae Edit Common name Scientific name Range CommentsLake Sidi Ali trout Salmo pallaryi Lake Aguelmame Sidi Ali Morocco Disappeared in 1934 after the introduction of the Eurasian carp 74 Toothcarps order Cyprinodontiformes Edit Livebearers and relatives family Poeciliidae Edit Scientific name Range CommentsAplocheilichthys sp nov Naivasha Lake Naivasha Kenya Disappeared in the 1970s or 1980s due to competition with introduced fishes 75 Insects class Insecta EditButterflies and moths order Lepidoptera Edit Gossamer winged butterflies family Lycaenidae Edit Common name Scientific name Range CommentsMbashe River buff Deloneura immaculata Mbhashe River Eastern Cape South Africa Only known from three individuals collected at the end of December 1863 76 Morant s blue Lepidochrysops hypopolia Eastern South Africa Only recorded in the 1870s 77 Ostracods class Ostracoda EditOrder Podocopida Edit Family Candonidae Edit Scientific name Range Comments PicturesNamibcypris costata Southern Kaokoveld Namibia Last recorded in 1987 78 nbsp See also EditList of Madagascar and Indian Ocean Island animals extinct in the Holocene List of Macaronesian animals extinct in the Holocene List of Saint Helena Ascension and Tristan da Cunha animals extinct in the Holocene Lists of extinct species List of extinct bird species since 1500 Extinct in the wild Lazarus taxonNotes Edit The source gives 11 700 calendar yr b2k before AD 2000 But BP means before AD 1950 Therefore the Holocene began 11 650 BP Doing the math that is c 9700 BCE and we are displeased because elephants have been removed from Libya because lions have disappeared from Thessaly because hippopotamoi have been gotten rid from the marshes of the Nile 7 This study did not use Syrian brown bears for comparison 20 References Edit Walker Mike Johnsen Sigfus Rasmussen Sune Olander Popp Trevor Steffensen Jorgen Peder Gibrard Phil Hoek Wim Lowe John Andrews John Bjo Rck Svante Cwynar Les C Hughen Konrad Kersahw Peter Kromer Bernd Litt Thomas Lowe David J Nakagawa Takeshi Newnham Rewi Schwander Jakob 2009 Formal definition and dating of the GSSP Global Stratotype Section and Point for the base of the Holocene using the Greenland NGRIP ice core and selected auxiliary records PDF Journal of Quaternary Science 24 1 3 17 Bibcode 2009JQS 24 3W doi 10 1002 jqs 1227 Archived PDF from the original on 2013 11 04 Retrieved 2022 04 24 Bronner G 2015 Cryptochloris wintoni IUCN Red List of Threatened Species 2015 e T5748A21287143 doi 10 2305 IUCN UK 2015 2 RLTS T5748A21287143 en Retrieved 12 November 2021 a b Happold D C D amp Kalina J Ed 2013 Mammals of Africa Introductory chapters and Afrotheria Bloomsbury 352 pages a b c d e f g h Riemer H amp Pollath N 2007 Desert animals in the eastern Sahara status economic significance and cultural reflection in antiquity Proceedings of an interdisciplinary ACACIA workshop held at the University of Cologne December 14 15 2007 Gowers William 1948 African Elephants and Ancient Authors African Affairs 47 188 173 180 doi 10 1093 oxfordjournals afraf a093647 JSTOR 718306 a b BolloK A amp Koncz I 2020 Sixth and Seventh Century Elephant Ivory Finds from the Carpathian Basin The Sources Circulation and Value of Ivory in Late Antiquity and the Early Middle Ages Archaeologiai Ertesito Vol 1 39 68 Braddock A C 2023 Implication An ecocritical dictionary of art history Yale University Press 256 pages Scientific Illustration Scientific Illustration Retrieved 2019 04 18 Kerbis Peterhans J amp Lavrenchenko L 2008 Nilopegamys plumbeus IUCN Red List of Threatened Species 2008 e T40766A10363474 doi 10 2305 IUCN UK 2008 RLTS T40766A10363474 en a b Richardson M 2023 Threatened and recently extinct vertebrates of the world A biogeographic approach Cambridge University Press 750 pages a b Turvey Sam 2009 Holocene extinctions Oxford University Press ISBN 978 0 19 953509 5 Retrieved 29 February 2012 Woodman Neal Ikram Salima Rowland Joanne 2021 04 07 A new addition to the embalmed fauna of ancient Egypt Guldenstaedt s White toothed Shrew Crocidura gueldenstaedtii Pallas 1811 Mammalia Eulipotyphla Soricidae PLOS ONE 16 4 e0249377 Bibcode 2021PLoSO 1649377W doi 10 1371 journal pone 0249377 ISSN 1932 6203 PMC 8026016 PMID 33826664 Bartosiewicz L 2009 A lion s share of attention archaeozoology and the historical record Acta Archaeologica 60 1 275 289 a b c d e Rossi L Scuzzarella C M amp Angelici F M 2020 Extinct or Perhaps Surviving Relict Populations of Big Cats Their Controversial Stories and Implications for Conservation In Problematic Wildlife II pp 393 417 Springer Cham a b Kitchener A C Breitenmoser Wursten C Eizirik E Gentry A Werdelin L Wilting A amp Tobe S 2017 A revised taxonomy of the Felidae The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group Cat News a b Miththapala S Seidensticker J amp O Brien S J 1996 Phylogeographic subspecies recognition in leopards Panthera pardus molecular genetic variation Conservation Biology 10 4 1115 1132 Gopalakrishnan Shyam Sinding Mikkel Holger S Ramos Madrigal Jazmin Niemann Jonas Samaniego Castruita Jose A Vieira Filipe G Caroe Christian de Montero Marc Manuel Kuderna Lukas Serres Aitor Gonzalez Basallote Victor Manuel Liu Yan Hu Wang Guo Dong Marques Bonet Tomas Mirarab Siavash Fernandes Carlos Gaubert Philippe Koepfli Klaus Peter Budd Jane Rueness Eli Knispel Heide Jorgensen Mads Peter Petersen Bent Sicheritz Ponten Thomas Bachmann Lutz Wiig Oystein Hansen Anders J Gilbert M Thomas P 2018 Interspecific gene flow shaped the evolution of the genus Canis Current Biology 28 21 3441 3449 e5 doi 10 1016 j cub 2018 08 041 PMC 6224481 PMID 30344120 a b c d Hamdine W et al 1998 Histoire recente de l ours brun au Maghreb C R Acad Sci Paris Sciences de la Vie Life Sciences Vol 321 pp 565 570 a b Ouchaou B amp Bougariane B 2015 Les extinctions totales et regionales des grands mammiferes durant le Quaternaire terminal au Maroc Travaux de l Institut Scientifique 8 5 20 Calvignac S Hughes S Tougard C Michaux J Thevenot M Philippe M amp Hanni C 2008 Ancient DNA evidence for the loss of a highly divergent brown bear clade during historical times Molecular Ecology 17 8 1962 1970 a b c Moehlman P D 2002 Equids zebras asses and horses status survey and conservation action plan IUCN a b c d Kimura B Marshall F B Chen S Rosenbom S Moehlman P D Tuross N amp Mulligan C J 2011 Ancient DNA from Nubian and Somali wild ass provides insights into donkey ancestry and domestication Proceedings of the Royal Society B Biological Sciences 278 1702 50 57 Guagnin M Shipton C el Dossary S al Rashid M Moussa F Stewart M amp Petraglia M D 2018 Rock art provides new evidence on the biogeography of kudu Tragelaphus imberbis wild dromedary aurochs Bos primigenius and African wild ass Equus africanus in the early and middle Holocene of north western Arabia Journal of Biogeography 45 4 727 740 Groves C 2013 Subgenus Asinus African Wild Ass in The Mammals of Africa Vol V eds Kingdon J Happold DCD Butynski TM Hoffmann M Happold M amp Kalina J 414 417 Kingdon Jonathan 1997 The Kingdon field guide to African mammals Helm London Youcef S A M 2020 African origins of modern asses as seen from paleontology and DNA what about the Atlas wild ass Geobios 58 73 84 a b c d e f g h i j k Faith J T 2014 Late Pleistocene and Holocene mammal extinctions on continental Africa Earth Science Reviews 128 105 121 Ludovic Orlando Jessica L Metcalf Maria T Alberdi Miguel Telles Antunes Dominique Bonjean Marcel Otte Fabiana Martin et al 2009 Revising the recent evolutionary history of equids using ancient DNA Proceedings of the National Academy of Sciences USA 106 51 21754 21759 Bibcode 2009PNAS 10621754O doi 10 1073 pnas 0903672106 PMC 2799835 PMID 20007379 Bryden H 1889 Kloof and Karoo London Longmans Green and Co pp 393 403 ASIN B00CNE0EZC Rookmaaker K 2013 Genus Ceratotherium White Rhinoceros in The Mammals of Africa Vol V eds Kingdon J Happold DCD Butynski TM Hoffmann M Happold M amp Kalina J pp 445 446 Roset J P amp Harbi Riahi M 2007 El Akarit Un site archeologique du Paleolithique moyen dans le sud de la Tunisie Paris Editions Recherche sur les Civilisations2007 Rookmaaker L C amp Groves C P 1977 The extinct Cape rhinoceros Diceros bicornis bicornis Linnaeus 1758 In Szugetierkundliche Mitteilwnge pg 117 126 Emslie R 2020 Diceros bicornis ssp bicornis IUCN Red List of Threatened Species 2020 e T39318A45814419 doi 10 2305 IUCN UK 2020 1 RLTS T39318A45814419 en Retrieved 11 November 2021 Emslie R 2020 Diceros bicornis ssp longipes IUCN Red List of Threatened Species 2020 e T39319A45814470 doi 10 2305 IUCN UK 2020 1 RLTS T39319A45814470 en Retrieved 18 November 2021 Groves C amp Grubb P 2011 Ungulate Taxonomy Johns Hopkins University Press 336 pages a b Emslie R 2020 Ceratotherium simum ssp cottoni IUCN Red List of Threatened Species 2020 e T4183A45813838 doi 10 2305 IUCN UK 2020 1 RLTS T4183A45813838 en Retrieved 12 November 2021 Khayale C Omondi P Kariuki L Muruthi P Gichohi N Stejskal J amp Amin R 2021 Kenya s first White Rhino Conservation and Management Action Plan Pachyderm 62 112 118 Grubb P amp d Huart J P 2010 Rediscovery of the Cape warthog Phacochoerus aethiopicus a review Journal of East African Natural History 99 2 77 102 Fernandez P et al 2015 The last occurrence of Megaceroides algericus Lyddekker 1890 Mammalia Cervidae during the middle Holocene in the cave of Bizmoune Morocco Essaouira region Quaternary International 374 154 167 Kitagawa C 2008 The status of fallow deer in Ancient Egypt autochthonous or introduced MOM Editions 49 1 541 552 Houlihan P F 1986 Some remarks on Deer Cervidae in Ancient Egypt The Journal of Egyptian Archaeology Vol 73 1 Tsahar E Izhaki I Lev Yadun S Bar Oz G 2009 Distribution and Extinction of Ungulates during the Holocene of the Southern Levant PLoS ONE 4 4 e5316 doi 10 1371 journal pone 0005316 a b Extinct and vanishing mammals of the Old World 1945 by Harper Francis from the Internet Archive Meghen C MacHugh D E Bradley D G Genetic characterization and West African cattle fao org Archived from the original on 26 February 2019 Retrieved 20 September 2021 Linseele V 2004 Size and size change of the African aurochs during the Pleistocene and Holocene Journal of African Archaeology 2 2 165 185 doi 10 3213 1612 1651 10026 IUCN SSC Antelope Specialist Group 2017 Kobus leche ssp robertsi IUCN Red List of Threatened Species 2017 e T11038A50190034 doi 10 2305 IUCN UK 2017 2 RLTS T11038A50190034 en Retrieved 12 November 2021 a b Mhorr Gazelle declared extinct in Tunisia Moreno E Sane A Benzal J Ibanez B Sanz Zuasti J amp Espeso G 2012 Changes in habitat structure may explain Decrease in reintroduced mohor gazelle population in the Guembeul Fauna Reserve Senegal Animals 2 3 347 360 Iyengar A Gilbert T Woodfine T Knowles J M Diniz F M Brenneman R A amp Maclean N 2007 Remnants of ancient genetic diversity preserved within captive groups of scimitar horned oryx Oryx dammah Molecular Ecology 16 12 2436 2449 Woodfine T amp Gilbert T 2016 The fall and rise of the scimitar horned oryx a case study of ex situ conservation and reintroduction in practice In Antelope Conservation From Diagnosis to Action 280 296 a b Ana S L Rodrigues Anne Charpentier Dario Bernal Casasola Armelle Gardeisen Carlos Nores Jose Antonio Pis Millan Krista McGrath Camilla F Speller July 11 2018 Forgotten Mediterranean calving grounds of grey and North Atlantic right whales evidence from Roman archaeological records Proceedings of the Royal Society B 285 1882 doi 10 1098 rspb 2018 0961 PMC 6053924 PMID 30051821 Notarbartolo di Sciara G Politi E Bayed A Beaubrun P C amp Knowlton A 1998 A winter cetacean survey off southern Morocco with a special emphasis on right whales Reports of the International Whaling Commission 48 547 550 Silva M A Steiner L Cascao I Cruz M J Prieto R Cole T amp Baumgartner M 2012 Winter sighting of a known western North Atlantic right whale in the Azores J Cetacean Res Manage 12 1 65 69 Cientificos piden ayuda para encontrar en aguas canarias a una cria de ballena franca extinta en el Atlantico Cooke J G 2018 Eschrichtius robustus IUCN Red List of Threatened Species 2018 e T8097A50353881 doi 10 2305 IUCN UK 2018 2 RLTS T8097A50353881 en Retrieved 19 November 2021 Hamilton Alex October 8 2015 The Gray Whale Sneaks Back into the Atlantic Two Centuries Later WNYC Retrieved 2019 10 28 a b c d Hume J P 2017 Extinct Birds Bloomsbury Publishing BirdLife International 2021 Haematopus meadewaldoi IUCN Red List of Threatened Species 2021 e T22693621A205917399 doi 10 2305 IUCN UK 2021 3 RLTS T22693621A205917399 en Retrieved 16 January 2023 BirdLife International 2018 Numenius tenuirostris IUCN Red List of Threatened Species 2018 e T22693185A131111201 doi 10 2305 IUCN UK 2018 2 RLTS T22693185A131111201 en Retrieved 4 April 2023 Campmas E Laroulandie V Michel P Amani F Nespoulet R amp Mohammed A E H 2010 22 A great auk Pinguinus impennis in North Africa discovery of In Birds in Archaeology Proceedings of the 6th Meeting of the ICAZ Bird Working Group in Groningen 23 8 27 8 2008 Vol 12 p 233 Barkhuis Pieper H 1985 The fossil land birds of Madeira and Porto Santo Bocagiana Museu de Historia Natural do Funchal Nº88 BirdLife International 2021 Pinguinus impennis IUCN Red List of Threatened Species 2021 e T22694856A205919631 doi 10 2305 IUCN UK 2021 3 RLTS T22694856A205919631 en Retrieved 6 April 2023 Fernandez Haeger J amp Jordano Barbudo D 2004 Las mariposas del monte mediterraneo Pgs 61 70 a b https elpais com elpais 2015 12 23 ciencia 1450863894 236217 html Ministerio para la Transicion Ecologica y el Reto Demografico Bates M F Jacobsen N 2018 Tetradactylus eastwoodae IUCN Red List of Threatened Species 2018 e T21663A115653635 doi 10 2305 IUCN UK 2018 2 RLTS T21663A115653635 en Retrieved 12 November 2021 IUCN IUCN IUCN FishBase team RMCA Geelhand D 2016 Barbus microbarbis IUCN Red List of Threatened Species 2016 e T61247A47242030 doi 10 2305 IUCN UK 2016 3 RLTS T61247A47242030 en Retrieved 12 November 2021 Freyhof J amp Ford M 2022 Labeobarbus reinii IUCN Red List of Threatened Species 2022 e T60765A137272327 doi 10 2305 IUCN UK 2022 1 RLTS T60765A137272327 en Retrieved 23 July 2022 Freyhof J amp Ford M 2022 Luciobarbus antinorii IUCN Red List of Threatened Species 2022 e T60731A137272558 doi 10 2305 IUCN UK 2022 1 RLTS T60731A137272558 en Retrieved 30 April 2023 Freyhof J amp Ford M 2022 Luciobarbus nasus IUCN Red List of Threatened Species 2022 e T217675561A137276808 doi 10 2305 IUCN UK 2022 1 RLTS T217675561A137276808 en Retrieved 30 April 2023 Marini A amp Talbi M 2008 Desertification and Risk Analysis Using High and Medium Resolution Satellite Data Training Workshop on Mapping Desertification Springer Science amp Business Media 274 pages Hanssens M 2004 Aplocheilichthys sp nov Naivasha IUCN Red List of Threatened Species 2004 e T61225A12451418 doi 10 2305 IUCN UK 2004 RLTS T61225A12451418 en Retrieved 12 November 2021 Williams M C 2020 Deloneura immaculata IUCN Red List of Threatened Species 2020 e T6331A168301470 doi 10 2305 IUCN UK 2020 2 RLTS T6331A168301470 en Retrieved 15 November 2021 Cockburn K N A 2020 Lepidochrysops hypopolia IUCN Red List of Threatened Species 2020 e T11537A168301541 doi 10 2305 IUCN UK 2020 2 RLTS T11537A168301541 en Retrieved 16 November 2021 Inland Water Crustacean Specialist Group 1996 Namibcypris costata IUCN Red List of Threatened Species 1996 e T14316A4431325 doi 10 2305 IUCN UK 1996 RLTS T14316A4431325 en Retrieved 12 November 2021 External links EditThe Extinction Website IUCN Red List of Threatened Species Retrieved from https en wikipedia org w index php title List of African animals extinct in the Holocene amp oldid 1176744318, wikipedia, wiki, book, books, library,

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