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2022 in paleoichthyology

This list of fossil fish research presented in 2022 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes that were described during the year, as well as other significant discoveries and events related to paleoichthyology that occurred in 2022.

List of years in paleoichthyology
In paleontology
2019
2020
2021
2022
2023
2024
2025
In paleobotany
2019
2020
2021
2022
2023
2024
2025
In arthropod paleontology
2019
2020
2021
2022
2023
2024
2025
In paleoentomology
2019
2020
2021
2022
2023
2024
2025
In paleomalacology
2019
2020
2021
2022
2023
2024
2025
In reptile paleontology
2019
2020
2021
2022
2023
2024
2025
In archosaur paleontology
2019
2020
2021
2022
2023
2024
2025
In mammal paleontology
2019
2020
2021
2022
2023
2024
2025

Jawless vertebrates edit

Name Novelty Status Authors Age Type locality Location Notes Images

Aenigmaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

Silurian (Wenlock)

Cape Phillips Formation (Cornwallis Island

  Canada

A member of Pteraspidomorphi. Genus includes new species A. falcata.

Anjiaspis ericius[2]

Sp. nov

In press

Shan et al.

Silurian (Telychian)

Qingshui Formation

  China

A member of Eugaleaspidiformes.

Anomalaspis[1]

Gen. et sp. nov

Thorsteinsson & Elliott

  Canada

A member of Traquairaspidiformes belonging to the family Traquairaspididae. Genus includes new species A. lacruma. The generic name is shared with Anomalaspis Brennan (1952).[3]

Archegonaspis cornwallisensis[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Arctictenaspis borealis[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Ctenaspidae.

Ariaspis cristata[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Ariaspidae.

Ariaspis majuscula[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Ariaspidae.

Ariaspis multijubata[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Ariaspidae.

Ariaspis nassichuki[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Ariaspidae.

Ariaspis perryi[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Ariaspidae.

Canadapteraspis formosa[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Protopteraspididae.

Canadapteraspis uniformis[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Protopteraspididae.

Corvaspis ellesmerensis[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Corvaspidiformes and the family Corvaspididae.

Corvaspis porphyretica[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Corvaspidiformes and the family Corvaspididae.

Corvaspis woodwardi[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Corvaspidiformes and the family Corvaspididae.

Denisonaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Protopteraspididae. Genus includes new species D. borea.

Dinaspidella elegans[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Dinaspidella tenuicostata[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Eumorphaspis[1]

Gen. et comb. et 3 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae. Genus includes E. borealis (Denison, 1963), as well as new species E. goodsiri, E. lata and E. solitaria.

Geissonaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Cyathaspidiformes. Genus includes new species G. mutabilis.

Genetaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Cyathaspidiformes and the family Jarvikaspididae. Genus includes new species G. incohata.

Idanaspis[1]

Gen. et 2 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci. Genus includes new species I. dimidiata and I. reinsoni.

Jarvikaspis[1]

Gen. et 2 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Cyathaspidiformes; the type genus of the new family Jarvikaspididae. Genus includes new species J. arctica and J. mauryensis.

Jiangxialepis jiujiangensis[4]

Sp. nov

In press

Shan, Zhao & Gai

Silurian (Telychian)

Qingshui Formation

  China

A member of Eugaleaspidiformes.

Kyphaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae. Genus includes new species K. boothiaensis.

Nahanniaspis mclintocki[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Orthogoniaspis[1]

Gen. et 2 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Weigeltaspidiformes and the family Weigeltaspididae. Genus includes new species O. magnijubata and O. loefflerae.

Paralaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae. Genus includes new species P. franklini.

Pionaspis ebenina[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Pionaspis rossi[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Prionotaspis[1]

Gen. et 2 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae. Genus includes new species P. miranda and possibly P? abbottensis.

Prosobranchiaspis[1]

Gen. et 2 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae. Genus includes new species P. prolatata and P. smithbayensis.

Pseudoanglaspis[1]

Gen. et 2 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae. Genus includes new species P. aquilonaris and P. minima.

Qingshuiaspis[2]

Gen. et sp. nov

In press

Shan et al.

Silurian (Telychian)

Qingshui Formation

  China

A member of Eugaleaspidiformes belonging to the family Shuyuidae. The type species is Q. junqingi.

Rimasventeraspis? halsteadi[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Natlaspidiformes and the family Rimasventeraspididae.

Rimasventeraspis? septentrionalis[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Natlaspidiformes and the family Rimasventeraspididae.

Soehnaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Cyathaspidiformes. Genus includes new species S. polaris.

Teleaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae. Genus includes new species T. tersa.

Thuleaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Cyathaspidiformes and the family Jarvikaspididae. Genus includes new species T. canadensis.

Toraspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Weigeltaspidiformes; the type genus of the new family Toraspididae. Genus includes new species T. somersetensis.

Traquairaspis longicarinata[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Traquairaspidiformes belonging to the family Traquairaspididae.

Traquairaspis pristina[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Traquairaspidiformes belonging to the family Traquairaspididae.

Trygonaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Protopteraspididae. Genus includes new species T. sicula.

Tujiaaspis[5]

Gen. et sp. nov

Valid

Gai et al.

Silurian (Telychian)

Huixingshao Formation

  China

A member of Eugaleaspidiformes; the anatomy of its articulated remains indicates that galeaspids possessed three unpaired dorsal fins, an approximately symmetrical hypochordal tail and a pair of continuous ventrolateral fins. The type species is T. vividus.

Vernonaspis magna[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Vernonaspis parryi[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Vernonaspis suffusca[1]

Sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of the family Cyathaspididae.

Westollaspis[1]

Gen. et 3 sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Heterostraci belonging to the group Corvaspidiformes; the type genus of the new family Westollaspididae. Genus includes new species W. hyperborea, W. cordata and W. gigas.

Whiteaspis[1]

Gen. et sp. nov

Valid

Thorsteinsson & Elliott

  Canada

A member of Traquairaspidiformes belonging to the family Traquairaspididae. Genus includes new species W. spinifera.

Xitunaspis[6]

Gen. et sp. nov

Valid

Sun et al.

Devonian (Lochkovian)

Xitun Formation

  China

A member of Eugaleaspidiformes belonging to the family Eugaleaspidae. The type species is X. magnus.

Yongdongaspis[7]

Gen. et sp. nov

Valid

Chen et al.

Silurian (Llandovery)

Huixingshao Formation

  China

A member of Eugaleaspidiformes. Genus includes new species Y. littoralis.

Jawless vertebrate research edit

  • A study on the phylogenetic relationships and evolutionary history of lampreys is published by Brownstein & Near (2022), who find Mesomyzon mengae to be a member of the lamprey crown group, and argue that living lamprey biodiversity results from diversifications extending from the Cretaceous to present, rather than gradually accumulating since the Paleozoic.[8]
  • Chevrinais et al. (2022) describe the ontogeny of Euphanerops longaevus.[9]
  • Meng et al. (2022) describe new fossil material of Pterogonaspis yuhaii from the Devonian Xujiachong Formation (Yunnan, China), providing new information on the cranial anatomy of this galeaspid, including the first fossil evidence for the position of the esophagus in galeaspids.[10]
  • The first detailed description of the complex of external endolymphatic structures in headshields of members of the genus Tremataspis from the Silurian of Estonia, with tiny platelets located within the openings of the endolymphatic duct and possibly functioning as a sieve that allowed or prevented material from entering the inner ear, is published by Märss, Wilson & Viljus (2022).[11]

Placoderms edit

Name Novelty Status Authors Age Type locality Location Notes Images

Amazichthys[12]

Gen. et sp. nov

Valid

Jobbins et al.

Devonian (Famennian)

  Morocco

A member of the family Selenosteidae. The type species is A. trinajsticae.

 

Xiushanosteus[13]

Gen. et sp. nov

Valid

Zhu, Li, Ahlberg & Zhu in Zhu et al.

Silurian (Telychian)

Huixingshao Formation

  China

The type species is X. mirabilis.

Placoderm research edit

  • A study on the morphology and function of the antiarch jaw apparatus is published by Lebedev et al. (2022).[14]
  • Wang & Zhu (2022) describe the squamation and scale morphology of Parayunnanolepis xitunensis, recognize at least thirteen morphotypes of scales in P. xitunensis, and interpret their findings as indicative of the high regionalization of squamation at the root of jawed vertebrates.[15]
  • Zhu et al. (2022) redescribe the pelvic region of the holotype of Parayunnanolepis xitunensis, and report that, instead of having two large plates previously designated as dermal pelvic girdles, P. xitunensis had three pairs of lateral pelvic plates and one large oval median pelvic plate.[16]
  • Trinajstic et al. (2022) report preservation of a three-dimensionally mineralized heart, thick-walled stomach and bilobed liver from members of Arthrodira from the Devonian Gogo Formation (Australia), and interpret this finding as indicative of the presence of a flat S-shaped heart separated from the liver and other abdominal organs, and of the absence of lungs in members of Arthrodira;[17] subsequently Jensen et al. (2023) question evidence of the presence of a chambered heart in the studied fossil material presented by Trinajstic et al. (2022).[18][19]

Acanthodians edit

Name Novelty Status Authors Age Type locality Location Notes Images

Fanjingshania[20]

Gen. et sp. nov

Valid

Andreev, Sansom, Li, Zhao & Zhu in Andreev et al.

Silurian (probably Aeronian)

Rongxi Formation

  China

A probable relative of climatiid "acanthodians". The type species is F. renovata.

 

Acanthodian research edit

  • A study aiming to quantify the completeness of the acanthodian fossil record is published by Schnetz et al. (2022).[21]
  • A study on the biomechanical properties and likely function of bony spines in front of the fins of members of the genus Machaeracanthus is published by Ferrón et al. (2022).[22]

Cartilaginous fishes edit

Name Novelty Status Authors Age Type locality Location Notes Images

Carcharhinus dicelmai[23]

Sp. nov

Valid

Collareta et al.

Miocene (Burdigalian)

Chilcatay Formation

  Peru

A species of Carcharhinus.

Cretascymnus beauryi[24]

Sp. nov

Feichtinger et al.

Late Cretaceous (Maastrichtian)

Gerhartsreit Formation

  Austria

A member of the family Somniosidae.

Dalatias orientalis[25]

Sp. nov

Malyshkina et al.

Miocene

Duho Formation

  South Korea

A species of Dalatias.

Dallasiella brachyodon[26]

Sp. nov

Valid

Siversson, Cederström & Ryan

Late Cretaceous (Campanian)

Kristianstad Basin

  Sweden

A member of Lamniformes.

Desinia[27]

Gen. et sp. nov

Valid

Ivanov in Ivanov et al.

Permian

  Russia
(  Kirov Oblast
  Komi Republic
  Mari El
  Tatarstan)

A member of the family Sphenacanthidae. The type species is D. radiata. Published online in 2023, but the issue date is listed as December 2022.[27]

Dracipinna[28]

Gen. et sp. nov

Valid

Pollerspöck & Straube

Late Oligocene and early Miocene

Ebelsberg Formation

  Austria
  Germany

A member of the family Dalatiidae. Genus includes new species D. bracheri.

Eoetmopterus davidi[24]

Sp. nov

Feichtinger et al.

Late Cretaceous (Maastrichtian)

Gerhartsreit Formation

  Austria

A member of the family Etmopteridae.

Fredipristis[24]

Gen et sp. nov

Feichtinger et al.

Late Cretaceous (Maastrichtian)

Gerhartsreit Formation

  Austria

A member of Squaliformes of uncertain affinities. The type species is F. eximia.

Gzhelodus[29]

Gen. et sp. nov

Valid

Ivanov

Carboniferous (KasimovianGzhelian)

  Russia
(  Moscow Oblast
  Samara Oblast
  Volgograd Oblast)

A member of Euselachii belonging to the family Protacrodontidae. The type species is G. serratus.

Heslerodoides[29]

Gen. et sp. nov

Valid

Ivanov

Carboniferous (Bashkirian–Gzhelian)

Zilim Formation

  Brazil
  Russia
(  Bashkortostan
  Moscow Oblast)

A member of Ctenacanthiformes belonging to the family Heslerodidae. The type species is H. triangularis.

Parvodus huizodus[30]

Sp. nov

Wen et al.

Early Triassic

Dongchuan Formation

  China

A member of Hybodontiformes.

Proetmopterus lukasi[24]

Sp. nov

Feichtinger et al.

Late Cretaceous (Maastrichtian)

Gerhartsreit Formation

  Austria

A member of the family Etmopteridae.

Protoxynotus mayrmelnhofi[31]

Sp. nov

Feichtinger et al.

Late Cretaceous (Maastrichtian)

Gerhartsreit Formation

  Austria

A member of the family Somniosidae.

Samarodus[29]

Gen. et sp. nov

Valid

Ivanov

Carboniferous (Moscovian–Gzhelian)

  Russia
(  Bashkortostan
  Moscow Oblast
  Samara Oblast
Pechora Sea)

A shark of uncertain affinities. The type species is S. flexus.

Scyliorhinus weemsi[32]

Sp. nov

Valid

Cicimurri, Knight & Ebersole

Oligocene (Rupelian)

Ashley Formation

  United States
(  South Carolina)

A species of Scyliorhinus.

Strophodus rebecae[33]

Sp. nov

Valid

Carrillo-Briceño & Cadena

Early Cretaceous (Valanginian-Hauterivian)

Rosablanca Formation

  Colombia

A member of Hybodontiformes belonging to the family Acrodontidae.

Taeniurops tosii[34]

Sp. nov

Adnet & Charpentier

Miocene (Aquitanian)

  Oman

A stingray, a species of Taeniurops.

Cartilaginous fish research edit

  • Duffin, Lauer & Lauer (2022) describe chimaeroid egg cases from the Upper Jurassic (Tithonian) Altmühltal Formation (Germany), probably produced by Ischyodus quenstedti, and name a new ichnotaxon Chimaerotheca schernfeldensis.[35]
  • Revision of the fossil material originally attributed to Bibractopiscis niger and Orthacanthus commailli, and a study on the implications of these fossils for the knowledge of the evolution of neurocranium in "ctenacanthiforms" and xenacanthiforms, is published by Luccisano et al. (2022).[36]
  • A study on the evolutionary history of members of the genus Orthacanthus from France and on their relationships with the other European species is published by Luccisano et al. (2022).[37]
  • Greif, Ferrón & Klug (2022) describe the first known fossil cartilage remains from the Devonian Hangenberg black shale from the Moroccan Anti-Atlas, and interpret its morphology as suggestive of ctenacanth affiliation.[38]
  • Taxonomic reassessment of a hybodontiform dental assemblage from the lower Kimmeridgian of Czarnogłowy (Poland), and a study on the implications of this assemblage for the knowledge of ecology and biogeography of cartilaginous fishes prior to the Jurassic/Cretaceous transition, is published by Stumpf, Meng & Kriwet (2022)[39]
  • Fossil teeth of sharks belonging to the groups Hexanchiformes, Echinorhiniformes, Squaliformes and Lamniformes, including the first record of Protosqualus in northwestern Pacific reported to date, are described from the Upper Cretaceous Nishichirashinai and Omagari formations (Yezo Group, Japan) by Kanno et al. (2022).[40]
  • New fossil material of Xampylodon dentatus, including more complete teeth or specimens representing teeth of different positions than most previous records, and the oldest fossil material of Rolfodon tatere reported to date is described from the Upper Cretaceous (Campanian) of James Ross Island (Antarctica) by dos Santos et al. (2022).[41]
  • Feichtinger et al. (2022) describe isolated teeth of Protoxynotus misburgensis from the Santonian of Lebanon, representing the first known record of this species from the southern Tethyan Realm, and interpret this finding as indicating that Protoxynotus and Cretascymnus occupied overlapping or similar habitats during the Late Cretaceous.[42]
  • Herraiz et al. (2022) describe teeth of a member of the genus Trigonognathus from the El Ferriol outcrop (Miocene of Spain), representing the first known record of this genus from the Mediterranean realm.[43]
  • Revision of the fossil record of the genus Echinorhinus in South America is published by Bogan & Agnolín (2022), who consider Echinorhinus pozzi and Echinorhinus maremagnum to be valid species, and consider E. maremagnum to be distinct from Echinorhinus lapaoi.[44]
  • A study on the anatomy, growth and ecology of Cretodus crassidens, based on data from a specimen from the Turonian "Lastame" lithofacies of the Scaglia Rossa Veneta (Lessini Mountains, Veneto, northeastern Italy), is published by Amalfitano et al. (2022).[45]
  • A tooth of Cetorhinus huddlestoni, as well as gill rakers differing from previously described cetorhinids and referred to the same species as the tooth, are described from the Miocene Duho Formation (South Korea) by Malyshkina, Nam & Kwon (2022).[46]
  • A study aiming to determine whether the observed body forms of lamniform sharks are influenced by thermophysiology, and reevaluating the body form of Otodus megalodon proposed by Cooper et al. (2020),[47] is published by Sternes, Wood & Shimada (2022).[48]
  • A study on the putative nursery areas and body size patterns across different populations of Otodus megalodon is published by Shimada et al. (2022), who report that specimens of O. megalodon are on average larger in cooler water than those in warmer water, and argue that the previously identified nursery areas may reflect temperature-dependent trends rather than the inferred reproductive strategy.[49]
  • McCormack et al. (2022) demonstrate the use of zinc isotopes to assess the trophic level in extant and extinct sharks, and interpret their findings as indicative of dietary shifts throughout the Neogene in sharks belonging to the genera Otodus and Carcharodon, and indicating that Early Pliocene sympatric great white sharks and Otodus megalodon likely occupied a similar mean trophic level.[50]
  • Evidence from nitrogen isotope ratios in fossil teeth of members of the genus Otodus, indicating that O. megalodon occupied a higher trophic level than is known for any marine species, extinct or extant, is presented by Kast et al. (2022).[51]
  • Cooper et al. (2022) create the first three-dimensional model of the body of Otodus megalodon and use it to infer its movement and feeding ecology, interpreting it as likely able to swim great distances and to feed on prey as large as modern apex predators.[52]
  • A study on tooth marks on physeteroid bones from the Miocene Pisco Formation (Peru) is published by Benites-Palomino et al. (2022), who interpret their findings as indicating that Miocene sharks were actively targeting the foreheads of physeteroids to feed on their lipid-rich nasal complexes, with the shape and distribution of the bite marks suggesting a series of consecutive scavenging events by members of different shark species.[53]
  • A study on the evolutionary history of carcharhiniform sharks is published by Brée, Condamine & Guinot (2022), who interpret their findings as indicative of an early low diversity period followed by a radiation exacerbated since 30 million years ago, as well as indicating that variations in diversification through time were likely linked to reef expansion and temperature change.[54]
  • Greenfield, Delsate & Candoni (2022) coin a new name Toarcibatidae for the family of Toarcian batomorphs previously referred to as Archaeobatidae.[55]
  • A study on the microstructure of rostral denticles of Ischyrhiza mira is published by Cook et al. (2022)[56]
  • New record of large dermal tubercles and bucklers, including tubercles similar in morphology to "Ceratoptera unios" and dermal bucklers similar in morphology to those of the extant roughtail stingray, is reported from the Lower Pleistocene Waccamaw Formation (South Carolina, United States) by Boessenecker & Gibson (2022), who interpret this findings as likely fossils of large stingrays in excess of 3 m disc width.[57]
  • A study on the phylogenetic relationships of extant and fossil rays and skates is published by Villalobos-Segura et al. (2022).[58]
  • A study on the completeness of the chondrichthyan fossil record from Florida, aiming to determine patterns in taxonomic and ecomorphological diversity of Eocene to Pleistocene chondrichthyans from the Florida Platform, is published by Perez (2022).[59]
  • Szabó et al. (2022) describe an assemblage of cartilaginous fishes from the Miocene Tekeres Schlieren Member of the Baden Formation (Hungary), including the first known records of deepwater cartilaginous fishes from the Badenian of the Central Paratethys.[60]
  • Fossil material of a diverse shark and ray fauna is reported from the early Pleistocene of Taiwan by Lin, Lin & Shimada (2022).[61]

Ray-finned fishes edit

Name Novelty Status Authors Age Type locality Location Notes Images

Acentrogobius matsya[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Acentrogobius.

Acipenser amnisinferos[63]

Sp. nov

Valid

Hilton & Grande

Late Cretaceous (Maastrichtian)

Hell Creek Formation

  United States
(  North Dakota)

A species of Acipenser.

Acipenser praeparatorum[63]

Sp. nov

Valid

Hilton & Grande

Late Cretaceous (Maastrichtian)

Hell Creek Formation

  United States
(  North Dakota)

A species of Acipenser.

Allocyclostoma[64]

Gen. et sp. nov

In press

Schwarzhans, Stringer & Welton

Early Cretaceous (Albian)

Pawpaw Formation

  United States
(  Texas)

Possibly a member of Polymixiiformes. The type species is A. alienus.

Amblyeleotris kireedam[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Amblyeleotris.

Ancistrogobius indicus[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Ancistrogobius.

Anisotremus rambo[65]

Sp. nov

Valid

Lin & Nolf

Eocene

  United States
(  Alabama
  Louisiana
  Texas)

A species of Anisotremus.

Archaeotolithus doppelsteini[66]

Sp. nov

Valid

Schwarzhans & Keupp

Early Jurassic (Pliensbachian)

Amaltheenton Formation

  Germany

A member of Actinopterygii of uncertain affinities.

Arconiapogon[67]

Gen. et sp. nov

Valid

Marramà, Giusberti & Carnevale

Oligocene (Rupelian)

  Italy

A member of the family Apogonidae belonging to the subfamily Apogoninae. The type species is A. deangelii.

Argentina? texana[64]

Sp. nov

In press

Schwarzhans, Stringer & Welton

Early Cretaceous (Albian)

Pawpaw Formation

  United States
(  Texas)

Possibly a species of Argentina.

Armigatus felixi[68]

Sp. nov

In press

Than-Marchese & Alvarado-Ortega

Early Cretaceous (Albian)

Tlayúa Formation

  Mexico

Astroscopus compactus[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Gosport Sand

  United States
(  Alabama
  Mississippi)

A species of Astroscopus.

Atractosteus grandei[69]

Sp. nov

Brownstein & Lyson

Paleogene (Danian)

Fort Union Formation

  United States
(  North Dakota)

A gar, a species of Atractosteus.

Bellottia verecunda[70]

Sp. nov

Valid

Carnevale & Schwarzhans

Miocene (Messinian)

  Italy

A species of Bellottia.

Benthosema taurinense[70]

Sp. nov

Valid

Carnevale & Schwarzhans

Miocene (Tortonian and Messinian)

  Italy

A species of Benthosema.

Blennius vernyhorovae[71]

Sp. nov

Schwarzhans, Klots & Kovalchuk in Schwarzhans et al.

Miocene

  Ukraine

A species of Blennius.

Bolinichthys higashibesshoensis[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Higashibessho Formation

  Japan

A species of Bolinichthys.

Bostrychus marsilii[70]

Sp. nov

Valid

Carnevale & Schwarzhans

Miocene (Messinian)

  Italy

A species of Bostrychus.

Bradyurus alessandroi[73]

Sp. nov

Valid

Bannikov & Zorzin

Eocene

Monte Bolca

  Italy

A member of Percoidei of uncertain phylogenetic placement.

Buenia pulvinus[74]

Sp. nov

Valid

Van Hinsbergh & Hoedemakers

Pliocene

  Morocco[75]
  Spain

A species of Buenia.

Caboellimma[76]

Gen. et comb. nov

De Figueiredo & Gallo

Early Cretaceous

Cabo Formation

  Brazil

A member of Clupeomorpha belonging to the group Ellimmichthyiformes; a new genus for "Ellimma" cruzae Santos (1990).

Callionymus vyali[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Callionymus.

Calypsoichthys[77]

Gen. et sp. nov

Valid

Argyriou et al.

Late Cretaceous (Maastrichtian)

  Greece

A member of the family Enchodontidae. The type species is C. pavlakiensis.

Cantarius ohei[78]

Sp. nov

Schwarzhans et al.

Miocene

Pebas Formation

  Peru

A member of the family Ariidae.

Capassopiscis[79]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Cenomanian)

  Lebanon

A member of the family Pantodontidae. The type species is C. pankowskii.

Centroberyx vaalsensis[80]

Sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Campanian)

Vaals Formation

  Netherlands

A species of Centroberyx.

Ceratoscopelus brevis[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Takakura Formation

  Japan

A species of Ceratoscopelus.

Chaetodon wattsi[67]

Sp. nov

Valid

Marramà, Giusberti & Carnevale

Oligocene (Rupelian)

  Italy

A species of Chaetodon.

Cichlasoma bluntschlii[78]

Sp. nov

Schwarzhans et al.

Miocene

Pebas Formation

  Peru

A species of Cichlasoma.

Cirripectes biconvexus[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Cirripectes.

"Citharus" varians[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Cook Mountain Formation

  United States
(  Alabama
  Texas)

A member of the family Citharidae.

Concentrilepis[81]

Gen. et sp. nov

Valid

Stack & Gottfried

Permian (Kungurian)

Minnekahta Limestone

  United States
(  South Dakota)

An early ray-finned fish with anatomical features of the paraphyletic "paleoniscoids". The type species is C. minnekahtaensis.

Coreoperca chosun[82]

Sp. nov

In press

Nam, Nazarkin & Bannikov

Early Miocene

Geumgwangdong Formation

  South Korea

A species of Coreoperca.

Coris medoboryensis[71]

Sp. nov

Schwarzhans, Klots & Kovalchuk in Schwarzhans et al.

Miocene

  Ukraine

A species of Coris.

Cuneatus maximus[83]

Sp. nov

Valid

Brownstein

Late Paleocene to early Eocene

Willwood Formation

  United States
(  Wyoming)

A gar.

Diaphus epipedus[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Takakura Formation

  Japan

A species of Diaphus.

Diaphus postcavallonis[74]

Sp. nov

Valid

Van Hinsbergh & Hoedemakers

Pliocene

  Spain

A species of Diaphus.

Diaphus watatsumi[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Takakura Formation

  Japan

A species of Diaphus.

Dicentrarchus oligocenicus[84]

Sp. nov

In press

Grădianu, Bordeianu & Codrea

Oligocene

Bituminous Marls Formation

  Romania

A species of Dicentrarchus.

Drombus thackerae[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Drombus.

Elopothrissus bernardlemorti[65]

Sp. nov

Valid

Lin & Nolf

Eocene

  United States
(  Alabama
  Texas)

A member of Albuliformes.

Elopothrissus pawpawensis[64]

Sp. nov

In press

Schwarzhans, Stringer & Welton

Early Cretaceous (Albian)

Pawpaw Formation

  United States
(  Texas)

Eosciaena[85]

Gen. et sp. nov

Valid

Stringer, Parmley & Quinn

Eocene (Bartonian)

Clinchfield Formation

  United States
(  Georgia (U.S. state)
  Louisiana[65])

A member of the family Sciaenidae. The type species is E. ebersolei.

Fibramia keralensis[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Fibramia.

Francischanos[86]

Gen. et comb. nov

In press

Ribeiro, Bockmann & Poyato-Ariza

Early Cretaceous (Aptian)

Quiricó Formation

  Brazil

A member of the family Chanidae; a new genus for "Dastilbe" moraesi Silva-Santos (1955).

Fusigobius? venadicus[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

Possibly a species of Fusigobius.

Genartina princeps[64]

Sp. nov

In press

Schwarzhans, Stringer & Welton

Early Cretaceous (Albian)

Pawpaw Formation

  United States
(  Texas)

A teleost of uncertain affinities, possibly near Albuliformes.

Germanostomus[87]

Gen. et sp. nov

Valid

Cooper et al.

Early Jurassic (Toarcian)

Posidonia Shale

  Germany

A member of the family Pachycormidae belonging to the subfamily Asthenocorminae. The type species is G. pectopteri.

Gerres mlynskyi[88]

Sp. nov

Valid

Brzobohatý, Zahradníková & Hudáčková

Miocene

Vienna Basin

  Austria
  France
  Poland
  Slovakia

A species of Gerres.

"aff. Glyptophidium" stringeri[65]

Sp. nov

Valid

Lin & Nolf

Eocene

  United States
(  Alabama
  Mississippi
  Texas)

A cusk-eel.

Gobiodon burdigalicus[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Gobiodon.

Gobius bratishkoi[71]

Sp. nov

Schwarzhans, Klots & Kovalchuk in Schwarzhans et al.

Miocene

  Ukraine

A species of Gobius.

Gobius ukrainicus[71]

Sp. nov

Schwarzhans, Klots & Kovalchuk in Schwarzhans et al.

Miocene

  Ukraine

A species of Gobius.

Harenaichthys[89]

Gen. et sp. nov

In press

Kim et al.

Late Cretaceous (Maastrichtian)

Nemegt Formation

  Mongolia

A member of Osteoglossomorpha. The type species is H. lui.

Ichthyotringa? cuneata[64]

Sp. nov

In press

Schwarzhans, Stringer & Welton

Early Cretaceous (Albian)

Pawpaw Formation

  United States
(  Texas)

A member of Aulopiformes belonging to the family Ichthyotringidae.

Ichthyotringa pindica[77]

Sp. nov

Valid

Argyriou et al.

Late Cretaceous (Maastrichtian)

  Greece

Italopterus[90]

Gen. et comb. nov

Valid

Shen & Arratia

Triassic

  Italy

A member of the family Thoracopteridae. Genus includes "Thoracopterus" magnificus Tintori & Sassi (1987) and "Thoracopterus" martinisi Tintori & Sassi (1992).

Jaydia? quilonica[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

Possibly a species of Jaydia.

Jenynsia herbsti[91]

Sp. nov

Valid

Sferco et al.

Late Miocene

  Argentina

A species of Jenynsia.

Kaykay[92]

Gen. et sp. nov

Valid

Gouiric-Cavalli & Arratia

Jurassic (Oxfordian stage)

  Argentina

A member of Pachycormiformes. Genus includes new species K. lafken.

Kutaichthys[93]

Gen et 2 sp. nov

Valid

Bakaev in Esin & Bakaev

Permian

  Russia
(  Komi Republic
  Perm Krai
  Samara Oblast)

An early ray-finned fish belonging to the group Palaeonisciformes and the family Palaeoniscidae. The type species is K. gubini Esin & Bakaev; genus also includes K. dozmerensis Esin & Bakaev. Published online in 2023, but the issue date is listed as December 2022.[93]

Lampadena exima[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Takakura Formation

  Japan

A species of Lampadena.

Lampanyctus beczynensis[94]

Sp. nov

Valid

Schwarzhans & Radwańska

Miocene (Langhian)

  Poland

A species of Lampanyctus.

Lampanyctus lenticularis[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Takakura Formation

  Japan

A species of Lampanyctus.

Lampanyctus tsuyamaensis[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Takakura Formation

  Japan

A species of Lampanyctus.

Leptolepis buttenheimensis[66]

Sp. nov

Valid

Schwarzhans & Keupp

Early Jurassic (Pliensbachian)

Amaltheenton Formation

  Germany

Extinct species of ray finned fish.

Leptolepis steberae[66]

Sp. nov

Valid

Schwarzhans & Keupp

Early Jurassic (Pliensbachian)

Amaltheenton Formation

  Germany

Libyachromis[95]

Gen. et sp. nov

Valid

Přikryl, Kaur & Murray

Oligocene

  Libya

A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is L. fugacior.

Lophionotus parnaibensis[96]

Sp. nov

In press

Gallo et al.

Late Jurassic

Pastos Bons Formation

  Brazil

A member of the family Semionotidae.

Makaira colonense[97]

Sp. nov

Valid

De Gracia et al.

Late Miocene

Chagres Formation

  Panama

A species of Makaira.

Makaira fierstini[97]

Sp. nov

Valid

De Gracia et al.

Late Miocene

Gatún Formation

  Panama

A species of Makaira.

Marcopoloichthys furreri[98]

Sp. nov

Valid

Arratia

Middle Triassic (Ladinian)

Upper Prosanto Formation

   Switzerland

A member of Teleosteomorpha belonging to the family Marcopoloichthyidae.

Medoborichthys[71]

Gen. et 2 sp. nov

Schwarzhans, Klots & Kovalchuk in Schwarzhans et al.

Miocene

  Ukraine

A member of the family Gobiidae belonging to the Priolepis lineage within the subfamily Gobiinae. The type species is M. renesulcis; genus also includes M. podolicus.

Mene garviei[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Cook Mountain Formation

  United States
(  Texas)

A species of Mene.

Minicholepis[99]

Gen et sp. nov

Valid

Bulanov, Minikh & Golubev

Permian

  Russia
(  Kirov Oblast)

A member of Eurynotoidiformes. The type species is M. primus. Published online in 2023, but the issue date is listed as December 2022.[99]

Moldavigobius[100]

Gen. et sp. et comb. nov

Valid

Reichenbacher & Bannikov

Miocene

  Moldova
  Turkey

A member of the family Gobiidae. The type species is M. helenae; genus also includes "Knipowitschia" suavis Schwarzhans (2014).

Morgula[97]

Gen. et sp. nov

Valid

De Gracia et al.

Late Miocene

Chagres Formation

  Panama

A marlin. The type species is M. donosochagrense.

"Muraenesox" barrytownensis[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Lisbon Formation

  United States
(  Alabama
  Texas)

A member of the family Muraenesocidae.

Neoopisthopterus weltoni[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Cook Mountain Formation

  United States
(  Texas)

A species of Neoopisthopterus.

Nezumia prikryli[101]

Sp. nov

Valid

Schwarzhans

Miocene (Langhian)

  Czech Republic

A species of Nezumia.

Notogoneus maarvelis[102]

Sp. nov

Valid

Grande & Wilson in Grande et al.

Late Cretaceous (Campanian)

  Canada
(  Northwest Territories)

A member of the family Gonorynchidae.

Nusaviichthys[103]

Gen. et sp. nov

In press

Alvarado-Ortega & Alves

Early Cretaceous (Albian)

  Mexico

A member of Crossognathiformes belonging to the family Notelopidae. The type species is N. nerivelai.

Oechsleria[104]

Gen. et sp. nov

In press

Micklich & Bannikov

Oligocene (Rupelian)

Bodenheim Formation

  Germany

A member of the family Veliferidae. The type species is O. unterfeldensis.

Oligophus bartonensis[105]

Sp. nov

Valid

Schwarzhans & Carnevale

Eocene (Bartonian)

Marne di Monte Piano Formation

  Italy

A lanternfish.

Oligopseudamia[67]

Gen. et sp. nov

Valid

Marramà, Giusberti & Carnevale

Oligocene (Rupelian)

  Italy

A member of the family Apogonidae belonging to the subfamily Pseudaminae. The type species is O. iancurtisi.

Oniketia[67]

Gen. et sp. nov

Valid

Marramà, Giusberti & Carnevale

Oligocene (Rupelian)

  Italy

A member of the family Gobiidae. The type species is O. akihitoi.

Pagellus pamunkeyensis[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Piney Point Formation

  United States
(  Virginia)

A species of Pagellus.

Palaeoneiros[106]

Gen. et sp. nov

Valid

Giles et al.

Devonian (Famennian)

Chadakoin Formation

  United States
(  Pennsylvania)

An early ray-finned fish with affinities to post-Devonian species Wendyichthys dicksoni and Cyranorhis bergeraci. Genus includes new species P. clackorum.

Paleoschizothorax diluculum[107]

Sp. nov

Valid

Yang et al.

Oligocene

Shangganchaigou Formation

  China

A member of the family Cyprinidae related to Schizothorax.

Paraconger wechesensis[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Weches Formation

  United States
(  Texas)

A species of Paraconger.

Parascombrops yanceyi[65]

Sp. nov

Valid

Lin & Nolf

Eocene

  United States
(  Texas)

A species of Parascombrops.

Paraulopus wichitae[64]

Sp. nov

In press

Schwarzhans, Stringer & Welton

Early Cretaceous (Albian)

Pawpaw Formation

  United States
(  Texas)

A species of Paraulopus.

Parenypnias[71]

Gen. et 2 sp. nov

Schwarzhans, Klots & Kovalchuk in Schwarzhans et al.

Miocene

  Czech Republic
  Ukraine

A member of the family Gobiidae belonging to the subfamily Gobiinae and the tribe Gobiosomatini. The type species is P. inauditus; genus also includes P. kiselevi.

Pavarottia astescalpone[108]

Sp. nov

Valid

Bannikov & Zorzin

Eocene

Monte Bolca

  Italy

A member of Percoidei belonging to the family Pavarottiidae.

Pebasciaena[78]

Gen. et sp. nov

Schwarzhans et al.

Miocene

Pebas Formation

  Peru

A member of the family Sciaenidae. The type species is P. amazonensis.

Piratata[109]

Gen. et sp. nov

In press

Richter et al.

Permian (Cisuralian)

Pedra de Fogo Formation

  Brazil

A deep-scaled ray-finned fish. Genus includes new species P. rogersmithii.

Plagioscion peyeri[78]

Sp. nov

Schwarzhans et al.

Miocene

Pebas Formation

  Peru

A species of Plagioscion.

Pleuropholis germinalis[110]

Sp. nov

Valid

Olive, Taverne & Brito

Early Cretaceous (Barremian to Aptian)

Sainte-Barbe Clays Formation

  Belgium

Pogonias tetragonus[78]

Sp. nov

Schwarzhans et al.

Miocene

Pebas Formation

  Peru

A species of Pogonias.

Polyspinatus[111]

Gen. et sp. nov

Schrøder et al.

Eocene

Fur Formation

  Denmark

A beardfish. Genus includes new species P. fluere.

Protoholocentrus[80]

Gen. et sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Campanian)

Vaals Formation

  Netherlands

A member of Holocentriformes of uncertain affinities. The type species is P. janjanssensi.

Prototetrapturus[97]

Gen. et comb. nov

Valid

De Gracia et al.

Miocene (Messinian)

  Algeria

A marlin. The type species is "Xiphiorhynchus" courcelli Arambourg (1927).

Pseudophichthys texanus[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Weches Formation

  United States
(  Alabama
  Mississippi
  Texas
  Virginia)

A species of Pseudophichthys.

Saurenchelys silex[74]

Sp. nov

Valid

Van Hinsbergh & Hoedemakers

Pliocene

  Morocco[75]
  Spain

A species of Saurenchelys.

Saurichthys sui[112]

Sp. nov

In press

Fang & Wu

Late Triassic

Junggar Basin

  China

Saurichthys taotie[113]

Sp. nov

Valid

Fang et al.

Late Triassic (Carnian)

Xiaowa Formation

  China

Announced in 2022; the final article version was published in 2023.

Scomber collettei[114]

Sp. nov

Valid

Bannikov & Erebakan

Miocene

  Russia
(  Krasnodar Krai)

A species of Scomber.

Seinstedtia[115]

Gen. et sp. nov

Valid

Schultze et al.

Late Triassic (Norian)

  Germany

A member of Teleosteomorpha of uncertain affinities. The type species is S. parva.

Sillaginocentrus crispus[80]

Sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Campanian)

Vaals Formation

  Netherlands

A member of Holocentriformes of uncertain affinities.

Siphamia minor[62]

Sp. nov

In press

Carolin et al.

Miocene (Burdigalian)

Quilon Formation

  India

A species of Siphamia.

Smithconger[116]

Gen. et comb. nov

Valid

Carnevale et al.

Eocene

Lillebælt Clay Formation

  Denmark
  New Zealand

A member of the family Congridae. The type species is "Pseudoxenomystax" treldeensis Schwarzhans (2007); genus also includes "Bathycongrus" waihaoensis Schwarzhans (2019).

Somalichromis[117]

Gen. et sp. nov

Murray

Oligocene

Daban Series

Somaliland

A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is S. hadrocephalus.

Spathochoira[97]

Gen. et comb. nov

Valid

De Gracia et al.

Late Miocene

Eastover Formation

  United States
(  Virginia)

A marlin. The type species is "Istiophorus" calvertensis Berry (1917).

Spinascutichthys[118]

Gen. et sp. nov

Valid

Murray, Chida & Holmes

Late Cretaceous (Cenomanian)

  Lebanon

A member of Aulopiformes belonging to the group Enchodontoidei. Genus includes new species S. pankowskiae.

Stenobrachius ohashii[72]

Sp. nov

Valid

Schwarzhans et al.

Miocene

Kurosedani Formation

  Japan

A species of Stenobrachius.

Stenobrachius sangsunii[119]

Sp. nov

Nam & Nazarkin

Miocene

Duho Formation

  South Korea

A species of Stenobrachius.

Sturisomatichthys podgornyi[120]

Sp. nov

Valid

Bogan & Agnolin

Late Miocene

  Argentina

A species of Sturisomatichthys.

Symmetrosulcus dockeryi[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Weches Formation

  United States
(  Alabama
  Mississippi
  Texas)

A cusk-eel.

Teffichthys elegans[121]

Sp. nov

Valid

Yuan et al.

Early Triassic (Induan)

Daye Formation

  China

A member of the family Perleididae.

 

Texoma[64]

Gen. et sp. nov

In press

Schwarzhans, Stringer & Welton

Early Cretaceous (Albian)

Pawpaw Formation

  United States
(  Texas)

Possibly a member of Polymixiiformes. The type species is T. cyclogaster.

Thorogobius antirostratus[88]

Sp. nov

Valid

Brzobohatý, Zahradníková & Hudáčková

Miocene

Vienna Basin

  Slovakia

A species of Thorogobius.

Trachinus meridianus[122]

Sp. nov

Valid

Schwarzhans & Kovalchuk

Miocene

  Poland
  Ukraine

A species of Trachinus.

Umbrina pachaula[78]

Sp. nov

Schwarzhans et al.

Miocene

Pebas Formation

  Peru

A species of Umbrina.

Vologdinia[99]

Gen et comb. nov

Valid

Bulanov, Minikh & Golubev

Permian

Poldarsa/Poldarskaya Formation

  Russia
(  Orenburg Oblast
  Vologda Oblast)

A member of Eurynotoidiformes. The type species is "Isadia" opokiensis Minikh & Andrushkevich (2017). Published online in 2023, but the issue date is listed as December 2022.[99]

Waitakia beelzebub[65]

Sp. nov

Valid

Lin & Nolf

Eocene

Piney Point Formation

  United States
(  Alabama
  Virginia)

A member of the subfamily Hemerocoetinae.

Xinjiangodus[123]

Gen. et sp. nov

Junior homonym

Zhou et al.

Late Cretaceous

Donggou Formation

  China

A member of the family Pycnodontidae. Genus includes new species X. gyrodoides. The generic name is preoccupied by Xinjiangodus Yue & Gao (1992).

Ray-finned fish research edit

  • A new database of the occurrences of Paleozoic ray-finned fishes is presented by Henderson et al. (2022), who evaluate the impact of fossil record biases, as well as taxonomic and phylogenetic issues, on the knowledge of the early evolution of ray-finned fishes;[124] subsequently Henderson, Dunne & Giles (2022) use this database to study patterns of diversity of ray-finned fishes through the Paleozoic, taking the extent and impact of sampling biases into account.[125]
  • A novel mode of fang accommodation, with teeth of the lower jaw inserting into fenestrae of the upper jaw, is reported in Brazilichthys macrognathus by Figueroa & Andrews (2022).[126]
  • Redescription and a study on the affinities of Toyemia is published by Bakaev & Kogan (2022).[127]
  • Redescription of the anatomy and a study on the affinities of Brachydegma caelatum is published by Argyriou, Giles & Friedman (2022).[128]
  • Osteoderms providing evidence of presence of large sturgeons (within the upper size bracket for Acipenseridae) in early-middle Paleocene freshwater ecosystems of western North America are described from the Fort Union Formation (Montana, United States) by Brownstein (2022).[129]
  • Fossil material of a member or a relative of the genus Eomesodon, representing the oldest record of pycnodonts from East Gondwana reported to date, is described from the Middle Jurassic (Bathonian) Jaisalmer Formation (Rajasthan, India) by Kumar et al. (2022).[130]
  • A study on the tooth replacement pattern and implantation in Serrasalmimus secans is published by Matsui & Kimura (2022), who interpret their findings as indicating that serrasalmimid pycnodont fish independently acquired a vertical replacement in true thecodont implantation, i.e. a characteristic tooth replacement pattern of mammals.[131]
  • A study on the phylogenetic relationships and evolutionary history of extant and extinct gars is published by Brownstein et al. (2022).[132]
  • Redescription and a study on the affinities of Saurostomus esocinus is published by Cooper & Maxwell (2022), who interpret this taxon as the basalmost transitional member of the suspension-feeding clade of pachycormids.[133]
  • A study on bone repair in response to damage in Leedsichthys problematicus is published by Johanson et al. (2022).[134]
  • Redescription and a study on the affinities of Thaumaturus intermedius is published by Micklich & Arratia (2022).[135]
  • Redescription of "Diplomystus" solignaci is published by Marramà, Khalloufi & Carnevale (2022), who interpret this fish as a paraclupeid ellimmichthyiform, and transfer it to the genus Paraclupea.[136]
  • A study on cranial morphological features that diagnose known families of catfishes, and on their implications for the knowledge of the affinities of catfishes from the Paleogene of Africa, is published by Murray & Holmes (2022), who reassess the familial placement of the Paleogene African catfishes and assign Eomacrones wilsoni to the family Bagridae sensu stricto.[137]
  • Description of new fossil material of Enchodus from the Cenomanian of Ukraine, and a revision of earlier records of Enchodus from Ukraine, is published by Kovalchuk, Barkaszi & Anfimova (2022).[138]
  • Redescription and a study on the phylogenetic affinities of Protosyngnathus sumatrensis is published by Murray (2022).[139]
  • A study on the phylogenetic affinities of fossil gobioids is published by Gierl et al. (2022).[140]
  • New specimen of Mene rhombea with extensive soft tissue preservation and striking colour patterning is described from the Eocene (Ypresian) Monte Bolca Lagerstätte (Italy) by Rossi et al. (2022).[141]
  • A study on patterns of body size evolution of tetraodontiforms in relation to paleoclimate events is published by Troyer et al. (2022).[142]
  • Přikryl et al. (2022) describe a new specimen of Archaeotetraodon winterbottomi from the Oligocene Rybnytsia Member of the Menilite Series (Ukraine), providing new information on the anatomy of this tetraodontid.[143]

Lobe-finned fishes edit

Name Novelty Status Authors Age Type locality Location Notes Images

Ceratodus guanganensis[144]

Sp. nov

Valid

Wang et al.

Late Jurassic

Shaximiao Formation

  China

Announced in 2021; the final version of the article naming it was published in 2022.

Ceratodus shishkini[145]

Sp. nov

Valid

Minikh

Triassic

  Russia
(  Orenburg Oblast)

Published online in 2023, but the issue date is listed as December 2022.[145]

Dianodipterus[146]

Gen. et sp. nov

Valid

Luo et al.

Devonian (Eifelian)

Qujing Formation

  China

A lungfish. The type species is D. huizeensis.

Diplurus enigmaticus[147]

Sp. nov

Brownstein & Bissell

Triassic

  United States
(  New Jersey)

A coelacanth.

Langlieria smalingi[148]

Sp. nov

Valid

Downs & Daeschler

Devonian (Frasnian)

Catskill Formation

  United States
(  Pennsylvania)

A member of the family Tristichopteridae.

Libys callolepis[149]

Sp. nov

Valid

Ferrante, Menkveld-Gfeller & Cavin

Early Jurassic (Toarcian)

Stadelgraben Formation

   Switzerland

A coelacanth belonging to the family Latimeriidae.

Qikiqtania[150]

Gen. et sp. nov

Valid

Stewart et al.

Devonian (Frasnian)

Fram Formation

  Canada
(  Nunavut)

An elpistostegalian. The type species is Q. wakei.

 

Rhizodus serpukhovensis[151]

Sp. nov

Valid

Smirnova

Carboniferous (Serpukhovian)

  Russia
(  Moscow Oblast)

Rinconodus[152]

Gen. et sp. nov

Valid

Panzeri et al.

Late Cretaceous (Santonian)

Bajo de la Carpa Formation

  Argentina

A lungfish belonging to the family Ceratodontidae. The type species is R. salvadori.

Lobe-finned fish research edit

  • Review of the phylogenetic analyses of onychodont relationships, aiming to determine the sources of discrepancies in the different phylogenetic hypotheses, is published by Ciudad Real et al. (2022).[153]
  • A study on the histology of the median fin bones and life history of Miguashaia bureaui is published by Mondéjar Fernandez et al. (2022).[154]
  • Toriño et al. (2022) describe a large mawsoniid coelacanth from the Upper Jurassic Kimmeridge Clay (United Kingdom), interpret its morphology as unexpectedly similar to the morphology of Mawsonia, and consider the studied coelacanth to be either an unknown Mawsonia-like form or a member of the lineage of Trachymetopon with some morphological characters previously assumed as diagnostic for Mawsonia.[155]
  • Description of cranial endocasts of six Paleozoic lungfish (Iowadipterus halli, Gogodipterus paddyensis, Pillararhynchus longi, Griphognathus whitei, Orlovichthys limnatis, Rhinodipterus ulrichi), and a study on the evolution of crania, brains and sensory abilities of lungfish, is published by Clement et al. (2022).[156]
  • Description of two well-preserved specimens of Youngolepis praecursor from the Devonian Xitun Formation (China), and a study on the implications of these specimens for the knowledge of the evolution of the specialized lungfish feeding mechanism, is published by Cui et al. (2022).[157]
  • A study on the anatomy of the neurocrania of Scaumenacia curta and Pentlandia macroptera is published by Boirot, Challands & Cloutier (2022), who report that the neurocranium of P. macroptera was at least partially ossified, while S. curta had a cartilaginous neurocranium, and evaluate the implications of their findings for the knowledge of paedomorphosis in lungfish evolution.[158]
  • A study on the histology of the tooth plates of Metaceratodus baibianorum from the Upper Cretaceous La Colonia Formation (Argentina) is published by Panzeri, Pereyra & Cione (2022).[159]
  • A study on the anatomy and affinities of Palaeospondylus gunni is published by Hirasawa et al. (2022), who interpret this taxon as a sarcopterygian, and likely a stem-tetrapod;[160] their conclusions are subsequently contested by Brownstein (2023).[161][162]

Other edit

Name Novelty Status Authors Age Type locality Location Notes Images

Permodontodus[163]

Gen. et sp. nov

Valid

McMenamin

Permian (Kungurian)

Wellington Formation

  United States
(  Oklahoma)

A bony fish of uncertain affinities, possibly a lobe-finned fish. Genus includes new species P. waurikensis.

Qianodus[164]

Gen. et sp. nov

Valid

Andreev, Sansom, Li, Zhao & Zhu in Andreev et al.

Silurian (probably Aeronian)

Rongxi Formation

  China

An early jawed vertebrate, likely a stem-chondrichthyan. The type species is Q. duplicis.

 

Shenacanthus[13]

Gen. et sp. nov

Valid

Zhu, Li, Ahlberg & Zhu in Zhu et al.

Silurian (Telychian)

Huixingshao Formation

  China

A stem-chondrichthyan. The type species is S. vermiformis.

 

General research edit

  • A study on the evolution of swimming speed in early vertebrates, inferred from caudal fin morphology of Paleozoic cyclostomes (Myxinidae and Petromyzontidae), jawless stem gnathostomes (Conodonta, Anaspida, Pteraspidomorphi, Thelodonti and Osteostraci) and placoderms, is published by Ferrón & Donoghue (2022), who interpret their findings as indicating that microsquamous taxa (thelodonts and anaspids) had higher swimming capabilities than vertebrates with rigid bony carapaces (including placoderms), that demonstrating that the rise of active nektonic vertebrates long-predated the Devonian.[165]
  • A study on the morphological similarities of Silurian and Devonian jawless and jawed vertebrates, aiming to determine which groups were most and least likely to have competed (and whether competition with jawed vertebrates was likely to cause the extinction of the majority of jawless vertebrates), is published by Scott & Anderson (2022), who don't find support for overall competitive exclusion of jawless vertebrates by jawed vertebrates.[166]
  • A study on the evolution of the vertebrate spiracular region from jawless vertebrates to tetrapods is published by Gai et al. (2022).[167]
  • A study on the mandibular morphology of Silurian and Devonian jawed vertebrates, and on the functional capabilities of their jaws, is published by Deakin et al. (2022).[168]
  • Description of the ichthyolith assemblage from the Upper Triassic Luning Formation (Nevada, United States), increasing known diversity of marine vertebrates in the western United States in the Late Triassic from four to at least 14 genera, is published by Tackett, Zierer & Clement (2022), who report evidence of the presence of taxa that were previously known only from Europe during the Late Triassic.[169]
  • Revision of the marine fish fauna from the Upper Cretaceous (Campanian) Rybushka Formation (Saratov Oblast, Russia) is published by Ebersole et al. (2022).[170]
  • A study aiming to reconstruct the fish community and oceanographic conditions off the coast of Peru during the last interglacial, based on data from sediments from the northern Humboldt Current system, is published by Salvatteci et al. (2022).[171]
  • Review of the fossil material providing information on the reproduction of extinct fishes is published by Capasso (2022), who names new ootaxa Theutonicootheca primigenia (possible egg capsule of jawless vertebrates or placoderms from the Emsian Kaub Formation, Germany), Beargulchootheca carbonifera (possible egg capsule of cartilaginous fishes from the Mississippian Bear Gulch Limestone of the Heath Formation, Montana, United States), Palaeochimaerootheca browni (egg capsule of chimaerids from the Pennsylvanian Cherokee Shale, Missouri, and Mazon Creek fossil beds, Illinois, United States), and Parascylliootheca libanica (egg capsule of parascylliids from the Cretaceous Sannine Formation, Lebanon).[172]

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  41. ^ dos Santos, R. O.; Riff, D.; Amenábar, C. R.; Ramos, R. R. C.; Rodrigues, I. F.; Scheffler, S. M.; Carvalho, M. A. (2022). "New records of hexanchiform sharks (Elasmobranchii: Neoselachii) from the Late Cretaceous of Antarctica with comments on previous reports and described taxa". New Zealand Journal of Geology and Geophysics: 1–16. doi:10.1080/00288306.2022.2143382. S2CID 253570784.
  42. ^ Feichtinger, I.; Kindlimann, R.; Guinot, G.; Harzhauser, M.; Pollerspöck, J. (2022). "First record of the somniosid shark Protoxynotus misburgensis from the Santonian (Late Cretaceous) of the Southern Tethyan Realm". Historical Biology: An International Journal of Paleobiology: 1–6. doi:10.1080/08912963.2022.2162401. S2CID 255331620.
  43. ^ Herraiz, J. L.; Carrillo-Briceño, J. D.; Ferrón, H. G.; Adnet, S.; Botella, H.; Martínez-Pérez, C. (2022). "First fossil record (Middle Miocene) of the viper shark Trigonognathus Mochizuki and Ohe, 1990, in the Mediterranean realm". Journal of Vertebrate Paleontology. 42 (1). e2114360. Bibcode:2022JVPal..42E4360H. doi:10.1080/02724634.2022.2114360. hdl:10550/84833. S2CID 253065446.
  44. ^ Bogan, S.; Agnolín, F. L. (2022). "The fossil record of the Bramble-shark Echinorhinus (Echinorhiniformes, Echinorhinidae) in South America". Journal of South American Earth Sciences. 120. 104083. Bibcode:2022JSAES.12004083B. doi:10.1016/j.jsames.2022.104083. S2CID 253173677.
  45. ^ Amalfitano, J.; Dalla Vecchia, F. M.; Carnevale, G.; Fornaciari, E.; Roghi, G.; Giusberti, L. (2022). "Morphology and paleobiology of the Late Cretaceous large-sized shark Cretodus crassidens (Dixon, 1850) (Neoselachii; Lamniformes)". Journal of Paleontology. 96 (5): 1166–1188. Bibcode:2022JPal...96.1166A. doi:10.1017/jpa.2022.23. S2CID 248702856.
  46. ^ Malyshkina, T. P.; Nam, G.-S.; Kwon, S. H. (2022). "Basking shark remains (Lamniformes, Cetorhinidae) from the Miocene of South Korea". Journal of Vertebrate Paleontology. 41 (5): e2037625. doi:10.1080/02724634.2021.2037625. S2CID 247466156.
  47. ^ Cooper, J. A.; Pimiento, C.; Ferrón, H. G.; Benton, M. J. (2020). "Body dimensions of the extinct giant shark Otodus megalodon: a 2D reconstruction". Scientific Reports. 10 (1): Article number 14596. Bibcode:2020NatSR..1014596C. doi:10.1038/s41598-020-71387-y. PMC 7471939. PMID 32883981.
  48. ^ Sternes, P. C.; Wood, J. J.; Shimada, K. (2023). "Body forms of extant lamniform sharks (Elasmobranchii: Lamniformes), and comments on the morphology of the extinct megatooth shark, Otodus megalodon, and the evolution of lamniform thermophysiology". Historical Biology: An International Journal of Paleobiology. 35: 139–151. doi:10.1080/08912963.2021.2025228. S2CID 246655344.
  49. ^ Shimada, K.; Maisch, H. M.; Perez, V. J.; Becker, M. A.; Griffiths, M. L. (2023). "Revisiting body size trends and nursery areas of the Neogene megatooth shark, Otodus megalodon (Lamniformes: Otodontidae), reveals Bergmann's rule possibly enhanced its gigantism in cooler waters". Historical Biology: An International Journal of Paleobiology. 35 (2): 208–217. doi:10.1080/08912963.2022.2032024. S2CID 247311831.
  50. ^ McCormack, J.; Griffiths, M. L.; Kim, S. L.; Shimada, K.; Karnes, M.; Maisch, H.; Pederzani, S.; Bourgon, N.; Jaouen, K.; Becker, M. A.; Jöns, N.; Sisma-Ventura, G.; Straube, N.; Pollerspöck, J.; Hublin, J.-J.; Eagle, R. A.; Tütken, T. (2022). "Trophic position of Otodus megalodon and great white sharks through time revealed by zinc isotopes". Nature Communications. 13 (1): Article number 2980. Bibcode:2022NatCo..13.2980M. doi:
2022, paleoichthyology, this, list, fossil, fish, research, presented, 2022, list, taxa, jawless, vertebrates, placoderms, acanthodians, fossil, cartilaginous, fishes, bony, fishes, other, fishes, that, were, described, during, year, well, other, significant, . This list of fossil fish research presented in 2022 is a list of new taxa of jawless vertebrates placoderms acanthodians fossil cartilaginous fishes bony fishes and other fishes that were described during the year as well as other significant discoveries and events related to paleoichthyology that occurred in 2022 List of years in paleoichthyology 2019 2020 2021 2022 2023 2024 2025 In paleontology 2019 2020 2021 2022 2023 2024 2025In paleobotany 2019 2020 2021 2022 2023 2024 2025In arthropod paleontology 2019 2020 2021 2022 2023 2024 2025In paleoentomology 2019 2020 2021 2022 2023 2024 2025In paleomalacology 2019 2020 2021 2022 2023 2024 2025In reptile paleontology 2019 2020 2021 2022 2023 2024 2025In archosaur paleontology 2019 2020 2021 2022 2023 2024 2025In mammal paleontology 2019 2020 2021 2022 2023 2024 2025Art Archaeology Architecture Literature Music Philosophy Science Contents 1 Jawless vertebrates 1 1 Jawless vertebrate research 2 Placoderms 2 1 Placoderm research 3 Acanthodians 3 1 Acanthodian research 4 Cartilaginous fishes 4 1 Cartilaginous fish research 5 Ray finned fishes 5 1 Ray finned fish research 6 Lobe finned fishes 6 1 Lobe finned fish research 7 Other 8 General research 9 ReferencesJawless vertebrates editName Novelty Status Authors Age Type locality Location Notes ImagesAenigmaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott Silurian Wenlock Cape Phillips Formation Cornwallis Island nbsp Canada A member of Pteraspidomorphi Genus includes new species A falcata Anjiaspis ericius 2 Sp nov In press Shan et al Silurian Telychian Qingshui Formation nbsp China A member of Eugaleaspidiformes Anomalaspis 1 Gen et sp nov Thorsteinsson amp Elliott nbsp Canada A member of Traquairaspidiformes belonging to the family Traquairaspididae Genus includes new species A lacruma The generic name is shared with Anomalaspis Brennan 1952 3 Archegonaspis cornwallisensis 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Arctictenaspis borealis 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Ctenaspidae Ariaspis cristata 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Ariaspidae Ariaspis majuscula 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Ariaspidae Ariaspis multijubata 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Ariaspidae Ariaspis nassichuki 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Ariaspidae Ariaspis perryi 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Ariaspidae Canadapteraspis formosa 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Protopteraspididae Canadapteraspis uniformis 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Protopteraspididae Corvaspis ellesmerensis 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Corvaspidiformes and the family Corvaspididae Corvaspis porphyretica 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Corvaspidiformes and the family Corvaspididae Corvaspis woodwardi 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Corvaspidiformes and the family Corvaspididae Denisonaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Protopteraspididae Genus includes new species D borea Dinaspidella elegans 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Dinaspidella tenuicostata 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Eumorphaspis 1 Gen et comb et 3 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Genus includes E borealis Denison 1963 as well as new species E goodsiri E lata and E solitaria Geissonaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Cyathaspidiformes Genus includes new species G mutabilis Genetaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Cyathaspidiformes and the family Jarvikaspididae Genus includes new species G incohata Idanaspis 1 Gen et 2 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci Genus includes new species I dimidiata and I reinsoni Jarvikaspis 1 Gen et 2 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Cyathaspidiformes the type genus of the new family Jarvikaspididae Genus includes new species J arctica and J mauryensis Jiangxialepis jiujiangensis 4 Sp nov In press Shan Zhao amp Gai Silurian Telychian Qingshui Formation nbsp China A member of Eugaleaspidiformes Kyphaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Genus includes new species K boothiaensis Nahanniaspis mclintocki 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Orthogoniaspis 1 Gen et 2 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Weigeltaspidiformes and the family Weigeltaspididae Genus includes new species O magnijubata and O loefflerae Paralaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Genus includes new species P franklini Pionaspis ebenina 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Pionaspis rossi 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Prionotaspis 1 Gen et 2 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Genus includes new species P miranda and possibly P abbottensis Prosobranchiaspis 1 Gen et 2 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Genus includes new species P prolatata and P smithbayensis Pseudoanglaspis 1 Gen et 2 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Genus includes new species P aquilonaris and P minima Qingshuiaspis 2 Gen et sp nov In press Shan et al Silurian Telychian Qingshui Formation nbsp China A member of Eugaleaspidiformes belonging to the family Shuyuidae The type species is Q junqingi Rimasventeraspis halsteadi 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Natlaspidiformes and the family Rimasventeraspididae Rimasventeraspis septentrionalis 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Natlaspidiformes and the family Rimasventeraspididae Soehnaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Cyathaspidiformes Genus includes new species S polaris Teleaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Genus includes new species T tersa Thuleaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Cyathaspidiformes and the family Jarvikaspididae Genus includes new species T canadensis Toraspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Weigeltaspidiformes the type genus of the new family Toraspididae Genus includes new species T somersetensis Traquairaspis longicarinata 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Traquairaspidiformes belonging to the family Traquairaspididae Traquairaspis pristina 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Traquairaspidiformes belonging to the family Traquairaspididae Trygonaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Protopteraspididae Genus includes new species T sicula Tujiaaspis 5 Gen et sp nov Valid Gai et al Silurian Telychian Huixingshao Formation nbsp China A member of Eugaleaspidiformes the anatomy of its articulated remains indicates that galeaspids possessed three unpaired dorsal fins an approximately symmetrical hypochordal tail and a pair of continuous ventrolateral fins The type species is T vividus Vernonaspis magna 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Vernonaspis parryi 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Vernonaspis suffusca 1 Sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of the family Cyathaspididae Westollaspis 1 Gen et 3 sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Heterostraci belonging to the group Corvaspidiformes the type genus of the new family Westollaspididae Genus includes new species W hyperborea W cordata and W gigas Whiteaspis 1 Gen et sp nov Valid Thorsteinsson amp Elliott nbsp Canada A member of Traquairaspidiformes belonging to the family Traquairaspididae Genus includes new species W spinifera Xitunaspis 6 Gen et sp nov Valid Sun et al Devonian Lochkovian Xitun Formation nbsp China A member of Eugaleaspidiformes belonging to the family Eugaleaspidae The type species is X magnus Yongdongaspis 7 Gen et sp nov Valid Chen et al Silurian Llandovery Huixingshao Formation nbsp China A member of Eugaleaspidiformes Genus includes new species Y littoralis Jawless vertebrate research edit A study on the phylogenetic relationships and evolutionary history of lampreys is published by Brownstein amp Near 2022 who find Mesomyzon mengae to be a member of the lamprey crown group and argue that living lamprey biodiversity results from diversifications extending from the Cretaceous to present rather than gradually accumulating since the Paleozoic 8 Chevrinais et al 2022 describe the ontogeny of Euphanerops longaevus 9 Meng et al 2022 describe new fossil material of Pterogonaspis yuhaii from the Devonian Xujiachong Formation Yunnan China providing new information on the cranial anatomy of this galeaspid including the first fossil evidence for the position of the esophagus in galeaspids 10 The first detailed description of the complex of external endolymphatic structures in headshields of members of the genus Tremataspis from the Silurian of Estonia with tiny platelets located within the openings of the endolymphatic duct and possibly functioning as a sieve that allowed or prevented material from entering the inner ear is published by Marss Wilson amp Viljus 2022 11 Placoderms editName Novelty Status Authors Age Type locality Location Notes ImagesAmazichthys 12 Gen et sp nov Valid Jobbins et al Devonian Famennian nbsp Morocco A member of the family Selenosteidae The type species is A trinajsticae nbsp Xiushanosteus 13 Gen et sp nov Valid Zhu Li Ahlberg amp Zhu in Zhu et al Silurian Telychian Huixingshao Formation nbsp China The type species is X mirabilis Placoderm research edit A study on the morphology and function of the antiarch jaw apparatus is published by Lebedev et al 2022 14 Wang amp Zhu 2022 describe the squamation and scale morphology of Parayunnanolepis xitunensis recognize at least thirteen morphotypes of scales in P xitunensis and interpret their findings as indicative of the high regionalization of squamation at the root of jawed vertebrates 15 Zhu et al 2022 redescribe the pelvic region of the holotype of Parayunnanolepis xitunensis and report that instead of having two large plates previously designated as dermal pelvic girdles P xitunensis had three pairs of lateral pelvic plates and one large oval median pelvic plate 16 Trinajstic et al 2022 report preservation of a three dimensionally mineralized heart thick walled stomach and bilobed liver from members of Arthrodira from the Devonian Gogo Formation Australia and interpret this finding as indicative of the presence of a flat S shaped heart separated from the liver and other abdominal organs and of the absence of lungs in members of Arthrodira 17 subsequently Jensen et al 2023 question evidence of the presence of a chambered heart in the studied fossil material presented by Trinajstic et al 2022 18 19 Acanthodians editName Novelty Status Authors Age Type locality Location Notes ImagesFanjingshania 20 Gen et sp nov Valid Andreev Sansom Li Zhao amp Zhu in Andreev et al Silurian probably Aeronian Rongxi Formation nbsp China A probable relative of climatiid acanthodians The type species is F renovata nbsp Acanthodian research edit A study aiming to quantify the completeness of the acanthodian fossil record is published by Schnetz et al 2022 21 A study on the biomechanical properties and likely function of bony spines in front of the fins of members of the genus Machaeracanthus is published by Ferron et al 2022 22 Cartilaginous fishes editName Novelty Status Authors Age Type locality Location Notes ImagesCarcharhinus dicelmai 23 Sp nov Valid Collareta et al Miocene Burdigalian Chilcatay Formation nbsp Peru A species of Carcharhinus Cretascymnus beauryi 24 Sp nov Feichtinger et al Late Cretaceous Maastrichtian Gerhartsreit Formation nbsp Austria A member of the family Somniosidae Dalatias orientalis 25 Sp nov Malyshkina et al Miocene Duho Formation nbsp South Korea A species of Dalatias Dallasiella brachyodon 26 Sp nov Valid Siversson Cederstrom amp Ryan Late Cretaceous Campanian Kristianstad Basin nbsp Sweden A member of Lamniformes Desinia 27 Gen et sp nov Valid Ivanov in Ivanov et al Permian nbsp Russia nbsp Kirov Oblast nbsp Komi Republic nbsp Mari El nbsp Tatarstan A member of the family Sphenacanthidae The type species is D radiata Published online in 2023 but the issue date is listed as December 2022 27 Dracipinna 28 Gen et sp nov Valid Pollerspock amp Straube Late Oligocene and early Miocene Ebelsberg Formation nbsp Austria nbsp Germany A member of the family Dalatiidae Genus includes new species D bracheri Eoetmopterus davidi 24 Sp nov Feichtinger et al Late Cretaceous Maastrichtian Gerhartsreit Formation nbsp Austria A member of the family Etmopteridae Fredipristis 24 Gen et sp nov Feichtinger et al Late Cretaceous Maastrichtian Gerhartsreit Formation nbsp Austria A member of Squaliformes of uncertain affinities The type species is F eximia Gzhelodus 29 Gen et sp nov Valid Ivanov Carboniferous Kasimovian Gzhelian nbsp Russia nbsp Moscow Oblast nbsp Samara Oblast nbsp Volgograd Oblast A member of Euselachii belonging to the family Protacrodontidae The type species is G serratus Heslerodoides 29 Gen et sp nov Valid Ivanov Carboniferous Bashkirian Gzhelian Zilim Formation nbsp Brazil nbsp Russia nbsp Bashkortostan nbsp Moscow Oblast A member of Ctenacanthiformes belonging to the family Heslerodidae The type species is H triangularis Parvodus huizodus 30 Sp nov Wen et al Early Triassic Dongchuan Formation nbsp China A member of Hybodontiformes Proetmopterus lukasi 24 Sp nov Feichtinger et al Late Cretaceous Maastrichtian Gerhartsreit Formation nbsp Austria A member of the family Etmopteridae Protoxynotus mayrmelnhofi 31 Sp nov Feichtinger et al Late Cretaceous Maastrichtian Gerhartsreit Formation nbsp Austria A member of the family Somniosidae Samarodus 29 Gen et sp nov Valid Ivanov Carboniferous Moscovian Gzhelian nbsp Russia nbsp Bashkortostan nbsp Moscow Oblast nbsp Samara OblastPechora Sea A shark of uncertain affinities The type species is S flexus Scyliorhinus weemsi 32 Sp nov Valid Cicimurri Knight amp Ebersole Oligocene Rupelian Ashley Formation nbsp United States nbsp South Carolina A species of Scyliorhinus Strophodus rebecae 33 Sp nov Valid Carrillo Briceno amp Cadena Early Cretaceous Valanginian Hauterivian Rosablanca Formation nbsp Colombia A member of Hybodontiformes belonging to the family Acrodontidae Taeniurops tosii 34 Sp nov Adnet amp Charpentier Miocene Aquitanian nbsp Oman A stingray a species of Taeniurops Cartilaginous fish research edit Duffin Lauer amp Lauer 2022 describe chimaeroid egg cases from the Upper Jurassic Tithonian Altmuhltal Formation Germany probably produced by Ischyodus quenstedti and name a new ichnotaxon Chimaerotheca schernfeldensis 35 Revision of the fossil material originally attributed to Bibractopiscis niger and Orthacanthus commailli and a study on the implications of these fossils for the knowledge of the evolution of neurocranium in ctenacanthiforms and xenacanthiforms is published by Luccisano et al 2022 36 A study on the evolutionary history of members of the genus Orthacanthus from France and on their relationships with the other European species is published by Luccisano et al 2022 37 Greif Ferron amp Klug 2022 describe the first known fossil cartilage remains from the Devonian Hangenberg black shale from the Moroccan Anti Atlas and interpret its morphology as suggestive of ctenacanth affiliation 38 Taxonomic reassessment of a hybodontiform dental assemblage from the lower Kimmeridgian of Czarnoglowy Poland and a study on the implications of this assemblage for the knowledge of ecology and biogeography of cartilaginous fishes prior to the Jurassic Cretaceous transition is published by Stumpf Meng amp Kriwet 2022 39 Fossil teeth of sharks belonging to the groups Hexanchiformes Echinorhiniformes Squaliformes and Lamniformes including the first record of Protosqualus in northwestern Pacific reported to date are described from the Upper Cretaceous Nishichirashinai and Omagari formations Yezo Group Japan by Kanno et al 2022 40 New fossil material of Xampylodon dentatus including more complete teeth or specimens representing teeth of different positions than most previous records and the oldest fossil material of Rolfodon tatere reported to date is described from the Upper Cretaceous Campanian of James Ross Island Antarctica by dos Santos et al 2022 41 Feichtinger et al 2022 describe isolated teeth of Protoxynotus misburgensis from the Santonian of Lebanon representing the first known record of this species from the southern Tethyan Realm and interpret this finding as indicating that Protoxynotus and Cretascymnus occupied overlapping or similar habitats during the Late Cretaceous 42 Herraiz et al 2022 describe teeth of a member of the genus Trigonognathus from the El Ferriol outcrop Miocene of Spain representing the first known record of this genus from the Mediterranean realm 43 Revision of the fossil record of the genus Echinorhinus in South America is published by Bogan amp Agnolin 2022 who consider Echinorhinus pozzi and Echinorhinus maremagnum to be valid species and consider E maremagnum to be distinct from Echinorhinus lapaoi 44 A study on the anatomy growth and ecology of Cretodus crassidens based on data from a specimen from the Turonian Lastame lithofacies of the Scaglia Rossa Veneta Lessini Mountains Veneto northeastern Italy is published by Amalfitano et al 2022 45 A tooth of Cetorhinus huddlestoni as well as gill rakers differing from previously described cetorhinids and referred to the same species as the tooth are described from the Miocene Duho Formation South Korea by Malyshkina Nam amp Kwon 2022 46 A study aiming to determine whether the observed body forms of lamniform sharks are influenced by thermophysiology and reevaluating the body form of Otodus megalodon proposed by Cooper et al 2020 47 is published by Sternes Wood amp Shimada 2022 48 A study on the putative nursery areas and body size patterns across different populations of Otodus megalodon is published by Shimada et al 2022 who report that specimens of O megalodon are on average larger in cooler water than those in warmer water and argue that the previously identified nursery areas may reflect temperature dependent trends rather than the inferred reproductive strategy 49 McCormack et al 2022 demonstrate the use of zinc isotopes to assess the trophic level in extant and extinct sharks and interpret their findings as indicative of dietary shifts throughout the Neogene in sharks belonging to the genera Otodus and Carcharodon and indicating that Early Pliocene sympatric great white sharks and Otodus megalodon likely occupied a similar mean trophic level 50 Evidence from nitrogen isotope ratios in fossil teeth of members of the genus Otodus indicating that O megalodon occupied a higher trophic level than is known for any marine species extinct or extant is presented by Kast et al 2022 51 Cooper et al 2022 create the first three dimensional model of the body of Otodus megalodon and use it to infer its movement and feeding ecology interpreting it as likely able to swim great distances and to feed on prey as large as modern apex predators 52 A study on tooth marks on physeteroid bones from the Miocene Pisco Formation Peru is published by Benites Palomino et al 2022 who interpret their findings as indicating that Miocene sharks were actively targeting the foreheads of physeteroids to feed on their lipid rich nasal complexes with the shape and distribution of the bite marks suggesting a series of consecutive scavenging events by members of different shark species 53 A study on the evolutionary history of carcharhiniform sharks is published by Bree Condamine amp Guinot 2022 who interpret their findings as indicative of an early low diversity period followed by a radiation exacerbated since 30 million years ago as well as indicating that variations in diversification through time were likely linked to reef expansion and temperature change 54 Greenfield Delsate amp Candoni 2022 coin a new name Toarcibatidae for the family of Toarcian batomorphs previously referred to as Archaeobatidae 55 A study on the microstructure of rostral denticles of Ischyrhiza mira is published by Cook et al 2022 56 New record of large dermal tubercles and bucklers including tubercles similar in morphology to Ceratoptera unios and dermal bucklers similar in morphology to those of the extant roughtail stingray is reported from the Lower Pleistocene Waccamaw Formation South Carolina United States by Boessenecker amp Gibson 2022 who interpret this findings as likely fossils of large stingrays in excess of 3 m disc width 57 A study on the phylogenetic relationships of extant and fossil rays and skates is published by Villalobos Segura et al 2022 58 A study on the completeness of the chondrichthyan fossil record from Florida aiming to determine patterns in taxonomic and ecomorphological diversity of Eocene to Pleistocene chondrichthyans from the Florida Platform is published by Perez 2022 59 Szabo et al 2022 describe an assemblage of cartilaginous fishes from the Miocene Tekeres Schlieren Member of the Baden Formation Hungary including the first known records of deepwater cartilaginous fishes from the Badenian of the Central Paratethys 60 Fossil material of a diverse shark and ray fauna is reported from the early Pleistocene of Taiwan by Lin Lin amp Shimada 2022 61 Ray finned fishes editName Novelty Status Authors Age Type locality Location Notes ImagesAcentrogobius matsya 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Acentrogobius Acipenser amnisinferos 63 Sp nov Valid Hilton amp Grande Late Cretaceous Maastrichtian Hell Creek Formation nbsp United States nbsp North Dakota A species of Acipenser Acipenser praeparatorum 63 Sp nov Valid Hilton amp Grande Late Cretaceous Maastrichtian Hell Creek Formation nbsp United States nbsp North Dakota A species of Acipenser Allocyclostoma 64 Gen et sp nov In press Schwarzhans Stringer amp Welton Early Cretaceous Albian Pawpaw Formation nbsp United States nbsp Texas Possibly a member of Polymixiiformes The type species is A alienus Amblyeleotris kireedam 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Amblyeleotris Ancistrogobius indicus 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Ancistrogobius Anisotremus rambo 65 Sp nov Valid Lin amp Nolf Eocene nbsp United States nbsp Alabama nbsp Louisiana nbsp Texas A species of Anisotremus Archaeotolithus doppelsteini 66 Sp nov Valid Schwarzhans amp Keupp Early Jurassic Pliensbachian Amaltheenton Formation nbsp Germany A member of Actinopterygii of uncertain affinities Arconiapogon 67 Gen et sp nov Valid Marrama Giusberti amp Carnevale Oligocene Rupelian nbsp Italy A member of the family Apogonidae belonging to the subfamily Apogoninae The type species is A deangelii Argentina texana 64 Sp nov In press Schwarzhans Stringer amp Welton Early Cretaceous Albian Pawpaw Formation nbsp United States nbsp Texas Possibly a species of Argentina Armigatus felixi 68 Sp nov In press Than Marchese amp Alvarado Ortega Early Cretaceous Albian Tlayua Formation nbsp MexicoAstroscopus compactus 65 Sp nov Valid Lin amp Nolf Eocene Gosport Sand nbsp United States nbsp Alabama nbsp Mississippi A species of Astroscopus Atractosteus grandei 69 Sp nov Brownstein amp Lyson Paleogene Danian Fort Union Formation nbsp United States nbsp North Dakota A gar a species of Atractosteus Bellottia verecunda 70 Sp nov Valid Carnevale amp Schwarzhans Miocene Messinian nbsp Italy A species of Bellottia Benthosema taurinense 70 Sp nov Valid Carnevale amp Schwarzhans Miocene Tortonian and Messinian nbsp Italy A species of Benthosema Blennius vernyhorovae 71 Sp nov Schwarzhans Klots amp Kovalchuk in Schwarzhans et al Miocene nbsp Ukraine A species of Blennius Bolinichthys higashibesshoensis 72 Sp nov Valid Schwarzhans et al Miocene Higashibessho Formation nbsp Japan A species of Bolinichthys Bostrychus marsilii 70 Sp nov Valid Carnevale amp Schwarzhans Miocene Messinian nbsp Italy A species of Bostrychus Bradyurus alessandroi 73 Sp nov Valid Bannikov amp Zorzin Eocene Monte Bolca nbsp Italy A member of Percoidei of uncertain phylogenetic placement Buenia pulvinus 74 Sp nov Valid Van Hinsbergh amp Hoedemakers Pliocene nbsp Morocco 75 nbsp Spain A species of Buenia Caboellimma 76 Gen et comb nov De Figueiredo amp Gallo Early Cretaceous Cabo Formation nbsp Brazil A member of Clupeomorpha belonging to the group Ellimmichthyiformes a new genus for Ellimma cruzae Santos 1990 Callionymus vyali 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Callionymus Calypsoichthys 77 Gen et sp nov Valid Argyriou et al Late Cretaceous Maastrichtian nbsp Greece A member of the family Enchodontidae The type species is C pavlakiensis Cantarius ohei 78 Sp nov Schwarzhans et al Miocene Pebas Formation nbsp Peru A member of the family Ariidae Capassopiscis 79 Gen et sp nov Valid Taverne Late Cretaceous Cenomanian nbsp Lebanon A member of the family Pantodontidae The type species is C pankowskii Centroberyx vaalsensis 80 Sp nov In press Schwarzhans amp Jagt Late Cretaceous Campanian Vaals Formation nbsp Netherlands A species of Centroberyx Ceratoscopelus brevis 72 Sp nov Valid Schwarzhans et al Miocene Takakura Formation nbsp Japan A species of Ceratoscopelus Chaetodon wattsi 67 Sp nov Valid Marrama Giusberti amp Carnevale Oligocene Rupelian nbsp Italy A species of Chaetodon Cichlasoma bluntschlii 78 Sp nov Schwarzhans et al Miocene Pebas Formation nbsp Peru A species of Cichlasoma Cirripectes biconvexus 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Cirripectes Citharus varians 65 Sp nov Valid Lin amp Nolf Eocene Cook Mountain Formation nbsp United States nbsp Alabama nbsp Texas A member of the family Citharidae Concentrilepis 81 Gen et sp nov Valid Stack amp Gottfried Permian Kungurian Minnekahta Limestone nbsp United States nbsp South Dakota An early ray finned fish with anatomical features of the paraphyletic paleoniscoids The type species is C minnekahtaensis Coreoperca chosun 82 Sp nov In press Nam Nazarkin amp Bannikov Early Miocene Geumgwangdong Formation nbsp South Korea A species of Coreoperca Coris medoboryensis 71 Sp nov Schwarzhans Klots amp Kovalchuk in Schwarzhans et al Miocene nbsp Ukraine A species of Coris Cuneatus maximus 83 Sp nov Valid Brownstein Late Paleocene to early Eocene Willwood Formation nbsp United States nbsp Wyoming A gar Diaphus epipedus 72 Sp nov Valid Schwarzhans et al Miocene Takakura Formation nbsp Japan A species of Diaphus Diaphus postcavallonis 74 Sp nov Valid Van Hinsbergh amp Hoedemakers Pliocene nbsp Spain A species of Diaphus Diaphus watatsumi 72 Sp nov Valid Schwarzhans et al Miocene Takakura Formation nbsp Japan A species of Diaphus Dicentrarchus oligocenicus 84 Sp nov In press Grădianu Bordeianu amp Codrea Oligocene Bituminous Marls Formation nbsp Romania A species of Dicentrarchus Drombus thackerae 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Drombus Elopothrissus bernardlemorti 65 Sp nov Valid Lin amp Nolf Eocene nbsp United States nbsp Alabama nbsp Texas A member of Albuliformes Elopothrissus pawpawensis 64 Sp nov In press Schwarzhans Stringer amp Welton Early Cretaceous Albian Pawpaw Formation nbsp United States nbsp Texas Eosciaena 85 Gen et sp nov Valid Stringer Parmley amp Quinn Eocene Bartonian Clinchfield Formation nbsp United States nbsp Georgia U S state nbsp Louisiana 65 A member of the family Sciaenidae The type species is E ebersolei Fibramia keralensis 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Fibramia Francischanos 86 Gen et comb nov In press Ribeiro Bockmann amp Poyato Ariza Early Cretaceous Aptian Quirico Formation nbsp Brazil A member of the family Chanidae a new genus for Dastilbe moraesi Silva Santos 1955 Fusigobius venadicus 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India Possibly a species of Fusigobius Genartina princeps 64 Sp nov In press Schwarzhans Stringer amp Welton Early Cretaceous Albian Pawpaw Formation nbsp United States nbsp Texas A teleost of uncertain affinities possibly near Albuliformes Germanostomus 87 Gen et sp nov Valid Cooper et al Early Jurassic Toarcian Posidonia Shale nbsp Germany A member of the family Pachycormidae belonging to the subfamily Asthenocorminae The type species is G pectopteri Gerres mlynskyi 88 Sp nov Valid Brzobohaty Zahradnikova amp Hudackova Miocene Vienna Basin nbsp Austria nbsp France nbsp Poland nbsp Slovakia A species of Gerres aff Glyptophidium stringeri 65 Sp nov Valid Lin amp Nolf Eocene nbsp United States nbsp Alabama nbsp Mississippi nbsp Texas A cusk eel Gobiodon burdigalicus 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Gobiodon Gobius bratishkoi 71 Sp nov Schwarzhans Klots amp Kovalchuk in Schwarzhans et al Miocene nbsp Ukraine A species of Gobius Gobius ukrainicus 71 Sp nov Schwarzhans Klots amp Kovalchuk in Schwarzhans et al Miocene nbsp Ukraine A species of Gobius Harenaichthys 89 Gen et sp nov In press Kim et al Late Cretaceous Maastrichtian Nemegt Formation nbsp Mongolia A member of Osteoglossomorpha The type species is H lui Ichthyotringa cuneata 64 Sp nov In press Schwarzhans Stringer amp Welton Early Cretaceous Albian Pawpaw Formation nbsp United States nbsp Texas A member of Aulopiformes belonging to the family Ichthyotringidae Ichthyotringa pindica 77 Sp nov Valid Argyriou et al Late Cretaceous Maastrichtian nbsp GreeceItalopterus 90 Gen et comb nov Valid Shen amp Arratia Triassic nbsp Italy A member of the family Thoracopteridae Genus includes Thoracopterus magnificus Tintori amp Sassi 1987 and Thoracopterus martinisi Tintori amp Sassi 1992 Jaydia quilonica 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India Possibly a species of Jaydia Jenynsia herbsti 91 Sp nov Valid Sferco et al Late Miocene nbsp Argentina A species of Jenynsia Kaykay 92 Gen et sp nov Valid Gouiric Cavalli amp Arratia Jurassic Oxfordian stage nbsp Argentina A member of Pachycormiformes Genus includes new species K lafken Kutaichthys 93 Gen et 2 sp nov Valid Bakaev in Esin amp Bakaev Permian nbsp Russia nbsp Komi Republic nbsp Perm Krai nbsp Samara Oblast An early ray finned fish belonging to the group Palaeonisciformes and the family Palaeoniscidae The type species is K gubini Esin amp Bakaev genus also includes K dozmerensis Esin amp Bakaev Published online in 2023 but the issue date is listed as December 2022 93 Lampadena exima 72 Sp nov Valid Schwarzhans et al Miocene Takakura Formation nbsp Japan A species of Lampadena Lampanyctus beczynensis 94 Sp nov Valid Schwarzhans amp Radwanska Miocene Langhian nbsp Poland A species of Lampanyctus Lampanyctus lenticularis 72 Sp nov Valid Schwarzhans et al Miocene Takakura Formation nbsp Japan A species of Lampanyctus Lampanyctus tsuyamaensis 72 Sp nov Valid Schwarzhans et al Miocene Takakura Formation nbsp Japan A species of Lampanyctus Leptolepis buttenheimensis 66 Sp nov Valid Schwarzhans amp Keupp Early Jurassic Pliensbachian Amaltheenton Formation nbsp Germany Extinct species of ray finned fish Leptolepis steberae 66 Sp nov Valid Schwarzhans amp Keupp Early Jurassic Pliensbachian Amaltheenton Formation nbsp GermanyLibyachromis 95 Gen et sp nov Valid Prikryl Kaur amp Murray Oligocene nbsp Libya A cichlid belonging to the subfamily Pseudocrenilabrinae The type species is L fugacior Lophionotus parnaibensis 96 Sp nov In press Gallo et al Late Jurassic Pastos Bons Formation nbsp Brazil A member of the family Semionotidae Makaira colonense 97 Sp nov Valid De Gracia et al Late Miocene Chagres Formation nbsp Panama A species of Makaira Makaira fierstini 97 Sp nov Valid De Gracia et al Late Miocene Gatun Formation nbsp Panama A species of Makaira Marcopoloichthys furreri 98 Sp nov Valid Arratia Middle Triassic Ladinian Upper Prosanto Formation nbsp Switzerland A member of Teleosteomorpha belonging to the family Marcopoloichthyidae Medoborichthys 71 Gen et 2 sp nov Schwarzhans Klots amp Kovalchuk in Schwarzhans et al Miocene nbsp Ukraine A member of the family Gobiidae belonging to the Priolepis lineage within the subfamily Gobiinae The type species is M renesulcis genus also includes M podolicus Mene garviei 65 Sp nov Valid Lin amp Nolf Eocene Cook Mountain Formation nbsp United States nbsp Texas A species of Mene Minicholepis 99 Gen et sp nov Valid Bulanov Minikh amp Golubev Permian nbsp Russia nbsp Kirov Oblast A member of Eurynotoidiformes The type species is M primus Published online in 2023 but the issue date is listed as December 2022 99 Moldavigobius 100 Gen et sp et comb nov Valid Reichenbacher amp Bannikov Miocene nbsp Moldova nbsp Turkey A member of the family Gobiidae The type species is M helenae genus also includes Knipowitschia suavis Schwarzhans 2014 Morgula 97 Gen et sp nov Valid De Gracia et al Late Miocene Chagres Formation nbsp Panama A marlin The type species is M donosochagrense Muraenesox barrytownensis 65 Sp nov Valid Lin amp Nolf Eocene Lisbon Formation nbsp United States nbsp Alabama nbsp Texas A member of the family Muraenesocidae Neoopisthopterus weltoni 65 Sp nov Valid Lin amp Nolf Eocene Cook Mountain Formation nbsp United States nbsp Texas A species of Neoopisthopterus Nezumia prikryli 101 Sp nov Valid Schwarzhans Miocene Langhian nbsp Czech Republic A species of Nezumia Notogoneus maarvelis 102 Sp nov Valid Grande amp Wilson in Grande et al Late Cretaceous Campanian nbsp Canada nbsp Northwest Territories A member of the family Gonorynchidae Nusaviichthys 103 Gen et sp nov In press Alvarado Ortega amp Alves Early Cretaceous Albian nbsp Mexico A member of Crossognathiformes belonging to the family Notelopidae The type species is N nerivelai Oechsleria 104 Gen et sp nov In press Micklich amp Bannikov Oligocene Rupelian Bodenheim Formation nbsp Germany A member of the family Veliferidae The type species is O unterfeldensis Oligophus bartonensis 105 Sp nov Valid Schwarzhans amp Carnevale Eocene Bartonian Marne di Monte Piano Formation nbsp Italy A lanternfish Oligopseudamia 67 Gen et sp nov Valid Marrama Giusberti amp Carnevale Oligocene Rupelian nbsp Italy A member of the family Apogonidae belonging to the subfamily Pseudaminae The type species is O iancurtisi Oniketia 67 Gen et sp nov Valid Marrama Giusberti amp Carnevale Oligocene Rupelian nbsp Italy A member of the family Gobiidae The type species is O akihitoi Pagellus pamunkeyensis 65 Sp nov Valid Lin amp Nolf Eocene Piney Point Formation nbsp United States nbsp Virginia A species of Pagellus Palaeoneiros 106 Gen et sp nov Valid Giles et al Devonian Famennian Chadakoin Formation nbsp United States nbsp Pennsylvania An early ray finned fish with affinities to post Devonian species Wendyichthys dicksoni and Cyranorhis bergeraci Genus includes new species P clackorum Paleoschizothorax diluculum 107 Sp nov Valid Yang et al Oligocene Shangganchaigou Formation nbsp China A member of the family Cyprinidae related to Schizothorax Paraconger wechesensis 65 Sp nov Valid Lin amp Nolf Eocene Weches Formation nbsp United States nbsp Texas A species of Paraconger Parascombrops yanceyi 65 Sp nov Valid Lin amp Nolf Eocene nbsp United States nbsp Texas A species of Parascombrops Paraulopus wichitae 64 Sp nov In press Schwarzhans Stringer amp Welton Early Cretaceous Albian Pawpaw Formation nbsp United States nbsp Texas A species of Paraulopus Parenypnias 71 Gen et 2 sp nov Schwarzhans Klots amp Kovalchuk in Schwarzhans et al Miocene nbsp Czech Republic nbsp Ukraine A member of the family Gobiidae belonging to the subfamily Gobiinae and the tribe Gobiosomatini The type species is P inauditus genus also includes P kiselevi Pavarottia astescalpone 108 Sp nov Valid Bannikov amp Zorzin Eocene Monte Bolca nbsp Italy A member of Percoidei belonging to the family Pavarottiidae Pebasciaena 78 Gen et sp nov Schwarzhans et al Miocene Pebas Formation nbsp Peru A member of the family Sciaenidae The type species is P amazonensis Piratata 109 Gen et sp nov In press Richter et al Permian Cisuralian Pedra de Fogo Formation nbsp Brazil A deep scaled ray finned fish Genus includes new species P rogersmithii Plagioscion peyeri 78 Sp nov Schwarzhans et al Miocene Pebas Formation nbsp Peru A species of Plagioscion Pleuropholis germinalis 110 Sp nov Valid Olive Taverne amp Brito Early Cretaceous Barremian to Aptian Sainte Barbe Clays Formation nbsp BelgiumPogonias tetragonus 78 Sp nov Schwarzhans et al Miocene Pebas Formation nbsp Peru A species of Pogonias Polyspinatus 111 Gen et sp nov Schroder et al Eocene Fur Formation nbsp Denmark A beardfish Genus includes new species P fluere Protoholocentrus 80 Gen et sp nov In press Schwarzhans amp Jagt Late Cretaceous Campanian Vaals Formation nbsp Netherlands A member of Holocentriformes of uncertain affinities The type species is P janjanssensi Prototetrapturus 97 Gen et comb nov Valid De Gracia et al Miocene Messinian nbsp Algeria A marlin The type species is Xiphiorhynchus courcelli Arambourg 1927 Pseudophichthys texanus 65 Sp nov Valid Lin amp Nolf Eocene Weches Formation nbsp United States nbsp Alabama nbsp Mississippi nbsp Texas nbsp Virginia A species of Pseudophichthys Saurenchelys silex 74 Sp nov Valid Van Hinsbergh amp Hoedemakers Pliocene nbsp Morocco 75 nbsp Spain A species of Saurenchelys Saurichthys sui 112 Sp nov In press Fang amp Wu Late Triassic Junggar Basin nbsp ChinaSaurichthys taotie 113 Sp nov Valid Fang et al Late Triassic Carnian Xiaowa Formation nbsp China Announced in 2022 the final article version was published in 2023 Scomber collettei 114 Sp nov Valid Bannikov amp Erebakan Miocene nbsp Russia nbsp Krasnodar Krai A species of Scomber Seinstedtia 115 Gen et sp nov Valid Schultze et al Late Triassic Norian nbsp Germany A member of Teleosteomorpha of uncertain affinities The type species is S parva Sillaginocentrus crispus 80 Sp nov In press Schwarzhans amp Jagt Late Cretaceous Campanian Vaals Formation nbsp Netherlands A member of Holocentriformes of uncertain affinities Siphamia minor 62 Sp nov In press Carolin et al Miocene Burdigalian Quilon Formation nbsp India A species of Siphamia Smithconger 116 Gen et comb nov Valid Carnevale et al Eocene Lillebaelt Clay Formation nbsp Denmark nbsp New Zealand A member of the family Congridae The type species is Pseudoxenomystax treldeensis Schwarzhans 2007 genus also includes Bathycongrus waihaoensis Schwarzhans 2019 Somalichromis 117 Gen et sp nov Murray Oligocene Daban Series Somaliland A cichlid belonging to the subfamily Pseudocrenilabrinae The type species is S hadrocephalus Spathochoira 97 Gen et comb nov Valid De Gracia et al Late Miocene Eastover Formation nbsp United States nbsp Virginia A marlin The type species is Istiophorus calvertensis Berry 1917 Spinascutichthys 118 Gen et sp nov Valid Murray Chida amp Holmes Late Cretaceous Cenomanian nbsp Lebanon A member of Aulopiformes belonging to the group Enchodontoidei Genus includes new species S pankowskiae Stenobrachius ohashii 72 Sp nov Valid Schwarzhans et al Miocene Kurosedani Formation nbsp Japan A species of Stenobrachius Stenobrachius sangsunii 119 Sp nov Nam amp Nazarkin Miocene Duho Formation nbsp South Korea A species of Stenobrachius Sturisomatichthys podgornyi 120 Sp nov Valid Bogan amp Agnolin Late Miocene nbsp Argentina A species of Sturisomatichthys Symmetrosulcus dockeryi 65 Sp nov Valid Lin amp Nolf Eocene Weches Formation nbsp United States nbsp Alabama nbsp Mississippi nbsp Texas A cusk eel Teffichthys elegans 121 Sp nov Valid Yuan et al Early Triassic Induan Daye Formation nbsp China A member of the family Perleididae nbsp Texoma 64 Gen et sp nov In press Schwarzhans Stringer amp Welton Early Cretaceous Albian Pawpaw Formation nbsp United States nbsp Texas Possibly a member of Polymixiiformes The type species is T cyclogaster Thorogobius antirostratus 88 Sp nov Valid Brzobohaty Zahradnikova amp Hudackova Miocene Vienna Basin nbsp Slovakia A species of Thorogobius Trachinus meridianus 122 Sp nov Valid Schwarzhans amp Kovalchuk Miocene nbsp Poland nbsp Ukraine A species of Trachinus Umbrina pachaula 78 Sp nov Schwarzhans et al Miocene Pebas Formation nbsp Peru A species of Umbrina Vologdinia 99 Gen et comb nov Valid Bulanov Minikh amp Golubev Permian Poldarsa Poldarskaya Formation nbsp Russia nbsp Orenburg Oblast nbsp Vologda Oblast A member of Eurynotoidiformes The type species is Isadia opokiensis Minikh amp Andrushkevich 2017 Published online in 2023 but the issue date is listed as December 2022 99 Waitakia beelzebub 65 Sp nov Valid Lin amp Nolf Eocene Piney Point Formation nbsp United States nbsp Alabama nbsp Virginia A member of the subfamily Hemerocoetinae Xinjiangodus 123 Gen et sp nov Junior homonym Zhou et al Late Cretaceous Donggou Formation nbsp China A member of the family Pycnodontidae Genus includes new species X gyrodoides The generic name is preoccupied by Xinjiangodus Yue amp Gao 1992 Ray finned fish research edit A new database of the occurrences of Paleozoic ray finned fishes is presented by Henderson et al 2022 who evaluate the impact of fossil record biases as well as taxonomic and phylogenetic issues on the knowledge of the early evolution of ray finned fishes 124 subsequently Henderson Dunne amp Giles 2022 use this database to study patterns of diversity of ray finned fishes through the Paleozoic taking the extent and impact of sampling biases into account 125 A novel mode of fang accommodation with teeth of the lower jaw inserting into fenestrae of the upper jaw is reported in Brazilichthys macrognathus by Figueroa amp Andrews 2022 126 Redescription and a study on the affinities of Toyemia is published by Bakaev amp Kogan 2022 127 Redescription of the anatomy and a study on the affinities of Brachydegma caelatum is published by Argyriou Giles amp Friedman 2022 128 Osteoderms providing evidence of presence of large sturgeons within the upper size bracket for Acipenseridae in early middle Paleocene freshwater ecosystems of western North America are described from the Fort Union Formation Montana United States by Brownstein 2022 129 Fossil material of a member or a relative of the genus Eomesodon representing the oldest record of pycnodonts from East Gondwana reported to date is described from the Middle Jurassic Bathonian Jaisalmer Formation Rajasthan India by Kumar et al 2022 130 A study on the tooth replacement pattern and implantation in Serrasalmimus secans is published by Matsui amp Kimura 2022 who interpret their findings as indicating that serrasalmimid pycnodont fish independently acquired a vertical replacement in true thecodont implantation i e a characteristic tooth replacement pattern of mammals 131 A study on the phylogenetic relationships and evolutionary history of extant and extinct gars is published by Brownstein et al 2022 132 Redescription and a study on the affinities of Saurostomus esocinus is published by Cooper amp Maxwell 2022 who interpret this taxon as the basalmost transitional member of the suspension feeding clade of pachycormids 133 A study on bone repair in response to damage in Leedsichthys problematicus is published by Johanson et al 2022 134 Redescription and a study on the affinities of Thaumaturus intermedius is published by Micklich amp Arratia 2022 135 Redescription of Diplomystus solignaci is published by Marrama Khalloufi amp Carnevale 2022 who interpret this fish as a paraclupeid ellimmichthyiform and transfer it to the genus Paraclupea 136 A study on cranial morphological features that diagnose known families of catfishes and on their implications for the knowledge of the affinities of catfishes from the Paleogene of Africa is published by Murray amp Holmes 2022 who reassess the familial placement of the Paleogene African catfishes and assign Eomacrones wilsoni to the family Bagridae sensu stricto 137 Description of new fossil material of Enchodus from the Cenomanian of Ukraine and a revision of earlier records of Enchodus from Ukraine is published by Kovalchuk Barkaszi amp Anfimova 2022 138 Redescription and a study on the phylogenetic affinities of Protosyngnathus sumatrensis is published by Murray 2022 139 A study on the phylogenetic affinities of fossil gobioids is published by Gierl et al 2022 140 New specimen of Mene rhombea with extensive soft tissue preservation and striking colour patterning is described from the Eocene Ypresian Monte Bolca Lagerstatte Italy by Rossi et al 2022 141 A study on patterns of body size evolution of tetraodontiforms in relation to paleoclimate events is published by Troyer et al 2022 142 Prikryl et al 2022 describe a new specimen of Archaeotetraodon winterbottomi from the Oligocene Rybnytsia Member of the Menilite Series Ukraine providing new information on the anatomy of this tetraodontid 143 Lobe finned fishes editName Novelty Status Authors Age Type locality Location Notes ImagesCeratodus guanganensis 144 Sp nov Valid Wang et al Late Jurassic Shaximiao Formation nbsp China Announced in 2021 the final version of the article naming it was published in 2022 Ceratodus shishkini 145 Sp nov Valid Minikh Triassic nbsp Russia nbsp Orenburg Oblast Published online in 2023 but the issue date is listed as December 2022 145 Dianodipterus 146 Gen et sp nov Valid Luo et al Devonian Eifelian Qujing Formation nbsp China A lungfish The type species is D huizeensis Diplurus enigmaticus 147 Sp nov Brownstein amp Bissell Triassic nbsp United States nbsp New Jersey A coelacanth Langlieria smalingi 148 Sp nov Valid Downs amp Daeschler Devonian Frasnian Catskill Formation nbsp United States nbsp Pennsylvania A member of the family Tristichopteridae Libys callolepis 149 Sp nov Valid Ferrante Menkveld Gfeller amp Cavin Early Jurassic Toarcian Stadelgraben Formation nbsp Switzerland A coelacanth belonging to the family Latimeriidae Qikiqtania 150 Gen et sp nov Valid Stewart et al Devonian Frasnian Fram Formation nbsp Canada nbsp Nunavut An elpistostegalian The type species is Q wakei nbsp Rhizodus serpukhovensis 151 Sp nov Valid Smirnova Carboniferous Serpukhovian nbsp Russia nbsp Moscow Oblast Rinconodus 152 Gen et sp nov Valid Panzeri et al Late Cretaceous Santonian Bajo de la Carpa Formation nbsp Argentina A lungfish belonging to the family Ceratodontidae The type species is R salvadori Lobe finned fish research edit Review of the phylogenetic analyses of onychodont relationships aiming to determine the sources of discrepancies in the different phylogenetic hypotheses is published by Ciudad Real et al 2022 153 A study on the histology of the median fin bones and life history of Miguashaia bureaui is published by Mondejar Fernandez et al 2022 154 Torino et al 2022 describe a large mawsoniid coelacanth from the Upper Jurassic Kimmeridge Clay United Kingdom interpret its morphology as unexpectedly similar to the morphology of Mawsonia and consider the studied coelacanth to be either an unknown Mawsonia like form or a member of the lineage of Trachymetopon with some morphological characters previously assumed as diagnostic for Mawsonia 155 Description of cranial endocasts of six Paleozoic lungfish Iowadipterus halli Gogodipterus paddyensis Pillararhynchus longi Griphognathus whitei Orlovichthys limnatis Rhinodipterus ulrichi and a study on the evolution of crania brains and sensory abilities of lungfish is published by Clement et al 2022 156 Description of two well preserved specimens of Youngolepis praecursor from the Devonian Xitun Formation China and a study on the implications of these specimens for the knowledge of the evolution of the specialized lungfish feeding mechanism is published by Cui et al 2022 157 A study on the anatomy of the neurocrania of Scaumenacia curta and Pentlandia macroptera is published by Boirot Challands amp Cloutier 2022 who report that the neurocranium of P macroptera was at least partially ossified while S curta had a cartilaginous neurocranium and evaluate the implications of their findings for the knowledge of paedomorphosis in lungfish evolution 158 A study on the histology of the tooth plates of Metaceratodus baibianorum from the Upper Cretaceous La Colonia Formation Argentina is published by Panzeri Pereyra amp Cione 2022 159 A study on the anatomy and affinities of Palaeospondylus gunni is published by Hirasawa et al 2022 who interpret this taxon as a sarcopterygian and likely a stem tetrapod 160 their conclusions are subsequently contested by Brownstein 2023 161 162 Other editName Novelty Status Authors Age Type locality Location Notes ImagesPermodontodus 163 Gen et sp nov Valid McMenamin Permian Kungurian Wellington Formation nbsp United States nbsp Oklahoma A bony fish of uncertain affinities possibly a lobe finned fish Genus includes new species P waurikensis Qianodus 164 Gen et sp nov Valid Andreev Sansom Li Zhao amp Zhu in Andreev et al Silurian probably Aeronian Rongxi Formation nbsp China An early jawed vertebrate likely a stem chondrichthyan The type species is Q duplicis nbsp Shenacanthus 13 Gen et sp nov Valid Zhu Li Ahlberg amp Zhu in Zhu et al Silurian Telychian Huixingshao Formation nbsp China A stem chondrichthyan The type species is S vermiformis nbsp General research editA study on the evolution of swimming speed in early vertebrates inferred from caudal fin morphology of Paleozoic cyclostomes Myxinidae and Petromyzontidae jawless stem gnathostomes Conodonta Anaspida Pteraspidomorphi Thelodonti and Osteostraci and placoderms is published by Ferron amp Donoghue 2022 who interpret their findings as indicating that microsquamous taxa thelodonts and anaspids had higher swimming capabilities than vertebrates with rigid bony carapaces including placoderms that demonstrating that the rise of active nektonic vertebrates long predated the Devonian 165 A study on the morphological similarities of Silurian and Devonian jawless and jawed vertebrates aiming to determine which groups were most and least likely to have competed and whether competition with jawed vertebrates was likely to cause the extinction of the majority of jawless vertebrates is published by Scott amp Anderson 2022 who don t find support for overall competitive exclusion of jawless vertebrates by jawed vertebrates 166 A study on the evolution of the vertebrate spiracular region from jawless vertebrates to tetrapods is published by Gai et al 2022 167 A study on the mandibular morphology of Silurian and Devonian jawed vertebrates and on the functional capabilities of their jaws is published by Deakin et al 2022 168 Description of the ichthyolith assemblage from the Upper Triassic Luning Formation Nevada United States increasing known diversity of marine vertebrates in the western United States in the Late Triassic from four to at least 14 genera is published by Tackett Zierer amp Clement 2022 who report evidence of the presence of taxa that were previously known only from Europe during the Late Triassic 169 Revision of the marine fish fauna from the Upper Cretaceous Campanian Rybushka Formation Saratov Oblast Russia is published by Ebersole et al 2022 170 A study aiming to reconstruct the fish community and oceanographic conditions off the coast of Peru during the last interglacial based on data from sediments from the northern Humboldt Current system is published by Salvatteci et al 2022 171 Review of the fossil material providing information on the reproduction of extinct fishes is published by Capasso 2022 who names new ootaxa Theutonicootheca primigenia possible egg capsule of jawless vertebrates or placoderms from the Emsian Kaub Formation Germany Beargulchootheca carbonifera possible egg capsule of cartilaginous fishes from the Mississippian Bear Gulch Limestone of the Heath Formation Montana United States Palaeochimaerootheca browni egg capsule of chimaerids from the Pennsylvanian Cherokee Shale Missouri and Mazon Creek fossil beds Illinois United States and Parascylliootheca libanica egg capsule of parascylliids from the Cretaceous Sannine Formation Lebanon 172 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