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Yamnaya culture

The Yamnaya culture[a] or the Yamna culture,[b] also known as the Pit Grave culture or Ochre Grave culture, was a late Copper Age to early Bronze Age archaeological culture of the region between the Southern Bug, Dniester, and Ural rivers (the Pontic–Caspian steppe), dating to 3300–2600 BCE.[2] It was discovered by Vasily Gorodtsov following his archaeological excavations near the Donets River in 1901–1903. Its name derives from its characteristic burial tradition: Я́мная (romanization: yamnaya) is a Russian adjective that means 'related to pits (yama)', as these people used to bury their dead in tumuli (kurgans) containing simple pit chambers.

Yamnaya culture
Alternative names
  • Pit Grave culture
  • Yamna culture
  • Ochre Grave culture
  • Yamnaya Horizon
Geographical rangePontic–Caspian steppe in Europe
PeriodCopper Age, Bronze Age
Datesc. 3300 – 2600 BCE
Preceded bySamara culture, Khvalynsk culture, Dnieper–Donets culture, Sredny Stog culture, Repin culture, Maykop culture, Cucuteni–Trypillia culture, Cernavodă culture, Usatove culture
Followed by

West: Catacomb culture

East: Poltavka culture
Defined byVasily Gorodtsov

The Yamnaya economy was based upon animal husbandry, fishing, and foraging, and the manufacture of ceramics, tools, and weapons.[3] The people of the Yamnaya culture lived primarily as nomads, with a chiefdom system and wheeled carts and wagons that allowed them to manage large herds.[4] They are also closely connected to Final Neolithic cultures, which later spread throughout Europe and Central Asia, especially the Corded Ware people and the Bell Beaker culture,[4] as well as the peoples of the Sintashta, Andronovo, and Srubnaya cultures. Back migration from Corded Ware also contributed to Sintashta and Andronovo.[5] In these groups, several aspects of the Yamnaya culture are present.[c] Yamnaya material culture was very similar to the Afanasevo culture of South Siberia, and the populations of both cultures are genetically indistinguishable.[1] This suggests that the Afanasevo culture may have originated from the migration of Yamnaya groups to the Altai region or, alternatively, that both cultures developed from an earlier shared cultural source.[6]

Genetic studies have suggested that the people of the Yamnaya culture can be modelled as a genetic admixture between a population related to Eastern European Hunter-Gatherers (EHG)[d] and people related to hunter-gatherers from the Caucasus (CHG) in roughly equal proportions,[7] an ancestral component which is often named "Steppe ancestry", with additional admixture from Anatolian, Levantine, or Early European farmers.[8][9] Genetic studies also indicate that populations associated with the Corded Ware, Bell Beaker, Sintashta, and Andronovo cultures derived large parts of their ancestry from the Yamnaya or a closely related population.[1][10][11][12]

Origins edit

 
Largest expansion of the Yamna(ya) culture. Modified from [13] c. 3500 origins of Usatovo culture; 3300 origins of Yamna; c. 3300–3200 expansion of Yamna across the Pontic-Caspian steppe; c. 2700 end of Trypillia culture,[14] and transformation of Yamna into Corded Ware in the contact zone east of the Carpathian mountains; 3100–2600 Yamnaya expansion into the Danube Valley.[15][16][17]

The Yamnaya culture was defined by Vasily Gorodtsov in order to differentiate it from the Catacomb and Srubnaya cultures that existed in the area, but were considered to be of a later period. Due to the time interval to the Yamnaya culture, and the reliance on archaeological findings, debate as to its origin is ongoing.[18] In 1996, Pavel Dolukhanov suggested that the emergence of the Pit-Grave culture represents a social development of various different local Bronze Age cultures, thus representing "an expression of social stratification and the emergence of chiefdom-type nomadic social structures" which in turn intensified inter-group contacts between essentially heterogeneous social groups.[19]

The origin of the Yamnaya culture continues to be debated, with proposals for its origins pointing to both the Khvalynsk and Sredny Stog cultures.[18] The Khvalynsk culture (4700–3800 BCE)[20] (middle Volga) and the Don-based Repin culture (c. 3950–3300 BCE)[21] in the eastern Pontic-Caspian steppe, and the closely related Sredny Stog culture (c. 4500–3500 BCE) in the western Pontic-Caspian steppe, preceded the Yamnaya culture (3300–2500 BCE).[22][23]

Further efforts to pinpoint the location came from Anthony (2007), who suggested that the Yamnaya culture (3300–2600 BCE) originated in the DonVolga area at c. 3400 BC,[24][2] preceded by the middle Volga-based Khvalynsk culture and the Don-based Repin culture (c. 3950–3300 BCE),[21][2] arguing that late pottery from these two cultures can barely be distinguished from early Yamnaya pottery.[25] Earlier continuity from eneolithic but largely hunter-gatherer Samara culture and influences from the more agricultural Dnieper–Donets II are apparent.

He argues that the early Yamnaya horizon spread quickly across the Pontic–Caspian steppes between c. 3400 and 3200 BCE:[24]

The spread of the Yamnaya horizon was the material expression of the spread of late Proto-Indo-European across the Pontic–Caspian steppes.[26]
[...] The Yamnaya horizon is the visible archaeological expression of a social adjustment to high mobility – the invention of the political infrastructure to manage larger herds from mobile homes based in the steppes.[27]

Alternatively, Parpola (2015) relates both the Corded ware culture and the Yamnaya culture to the late Trypillia (Tripolye) culture.[28] He hypothesizes that "the Tripolye culture was taken over by PIE speakers by c. 4000 BCE,"[29] and that in its final phase the Trypillian culture expanded to the steppes, morphing into various regional cultures which fused with the late Sredny Stog (Serednii Stih) pastoralist cultures, which, he suggests, gave rise to the Yamnaya culture.[30] Dmytro Telegin viewed Sredny Stog and Yamna as one cultural continuum and considered Sredny Stog to be the genetic foundation of the Yamna.[31]

The Yamnaya culture was succeeded in its western range by the Catacomb culture (2800–2200 BCE); in the east, by the Poltavka culture (2700–2100 BCE) at the middle Volga. These two cultures were followed by the Srubnaya culture (18th–12th century BCE).

Characteristics edit

 
Yamnaya culture grave, Volgograd Oblast

The Yamnaya culture was nomadic[4] or semi-nomadic, with some agriculture practiced near rivers, and a few fortified sites, the largest of which is Mikhaylivka.[32]

Characteristic for the culture are the burials in pit graves under kurgans (tumuli), often accompanied by animal offerings. Some graves contain large anthropomorphic stelae, with carved human heads, arms, hands, belts, and weapons.[33] The dead bodies were placed in a supine position with bent knees and covered in ochre. Some kurgans contained "stratified sequences of graves".[34] Kurgan burials may have been rare, and were perhaps reserved for special adults, who were predominantly, but not necessarily, male.[35] Status and gender are marked by grave goods and position, and in some areas, elite individuals are buried with complete wooden wagons.[36] Grave goods are more common in eastern Yamnaya burials, which are also characterized by a higher proportion of male burials and more male-centred rituals than western areas.[37]

The Yamnaya culture had and used two-wheeled carts and four-wheeled wagons, which are thought to have been oxen-drawn at this time, and there is evidence that they rode horses.[38][39] For instance, several Yamnaya skeletons exhibit specific characteristics in their bone morphology that may have been caused by long-term horseriding.[38]

Metallurgists and other craftsmen are given a special status in Yamnaya society, and metal objects are sometimes found in large quantities in elite graves. New metalworking technologies and weapon designs are used.[36]

Stable isotope ratios of Yamna individuals from the Dnipro Valley suggest the Yamnaya diet was terrestrial protein based with insignificant contribution from freshwater or aquatic resources.[40] Anthony speculates that the Yamnaya ate meat, milk, yogurt, cheese, and soups made from seeds and wild vegetables, and probably consumed mead.[41]

Mallory and Adams suggest that Yamnaya society may have had a tripartite structure of three differentiated social classes, although the evidence available does not demonstrate the existence of specific classes such as priests, warriors, and farmers.[42]

Gallery edit

Archaeogenetics edit

 
Main genetic ancestry of Western Steppe Herders (Yamnaya pastoralists): a confluence of Eastern Hunter-Gatherers (EHG) and Caucasus Hunter-Gatherers (CHG).[44]
 
Admixture proportions of Yamnaya populations. They combined Eastern Hunter Gatherer ( EHG), Caucasian Hunter-Gatherer ( CHG), Anatolian Neolithic () and Western Hunter Gatherer ( WHG) ancestry.[45]

According to Jones et al. (2015) and Haak et al. (2015), autosomal tests indicate that the Yamnaya people were the result of a genetic admixture between two different hunter-gatherer populations: distinctive "Eastern Hunter-Gatherers" (EHG), from Eastern Europe, with high affinity to the Mal'ta–Buret' culture or other, closely related people from Siberia[10] and a population of "Caucasus hunter-gatherers" (CHG) who probably arrived from the Caucasus[46][7] or Iran.[47] Each of those two populations contributed about half the Yamnaya DNA.[11][7] This admixture is referred to in archaeogenetics as Western Steppe Herder (WSH) ancestry.

Admixture between EHGs and CHGs is believed to have occurred on the eastern Pontic-Caspian steppe starting around 5,000 BCE, while admixture with Early European Farmers (EEF) happened in the southern parts of the Pontic-Caspian steppe sometime later. More recent genetic studies have found that the Yamnaya were a mixture of EHGs, CHGs, and to a lesser degree Anatolian farmers and Levantine farmers, but not EEFs from Europe due to lack of WHG DNA in the Yamnaya. This occurred in two distinct admixture events from West Asia into the Pontic-Caspian steppe.[48][49]

Haplogroup R1b, specifically the Z2103 suBCElade of R1b-L23, is the most common Y-DNA haplogroup found among the Yamnaya specimens. This haplogroup is rare in Western Europe and mainly exists in Southeastern Europe today.[50] Additionally, a minority are found to belong to haplogroup I2.[11] They are found to belong to a wider variety of West Eurasian mtDNA haplogroups, including U, T, and haplogroups associated with Caucasus Hunter-Gatherers and Early European Farmers.[51][52] A small but significant number of Yamnaya kurgan specimens from Northern Ukraine carried the East Asian mtDNA haplogroup C4.[53][54]

People of the Yamnaya culture are believed to have had mostly brown eye colour, light to intermediate skin, and brown hair colour, with some variation.[55][56]

Some Yamnaya individuals are believed to have carried a mutation to the KITLG gene associated with blond hair, as several individuals with Steppe ancestry are later found to carry this mutation. The Ancient North Eurasian Afontova Gora group, who contributed significant ancestry to Western Steppe Herders, are believed to be the source of this mutation.[57] A study in 2015 found that Yamnaya had the highest ever calculated genetic selection for height of any of the ancient populations tested.[58][59] It has been hypothesized that an allele associated with lactase persistence (conferring lactose tolerance into adulthood) was brought to Europe from the steppe by Yamnaya-related migrations.[60][61][62][63]

A 2022 study by Lazaridis et al. found that the typical phenotype among the Yamnaya population was brown eyes, brown hair, and intermediate skin colour. None of their Yamnaya samples were predicted to have either blue eyes or blond hair, in contrast with later Steppe groups in Russia and Central Asia, as well as the Bell Beaker culture in Europe, who did carry these phenotypes in high proportions.[64]

The geneticist David Reich has argued that the genetic data supports the likelihood that the people of the Yamnaya culture were a "single, genetically coherent group" who were responsible for spreading many Indo-European languages.[65] Reich's group recently suggested that the source of Anatolian and Indo-European subfamilies of the Proto-Indo-European (PIE) language may have been in west Asia and the Yamna were responsible for the dissemination of the latter.[9] Reich also argues that the genetic evidence shows that Yamnaya society was an oligarchy dominated by a small number of elite males.[66]

The genetic evidence for the extent of the role of the Yamnaya culture in the spread of Indo-European languages has been questioned by Russian archaeologist Leo Klejn[67] and Balanovsky et al.,[68] who note a lack of male haplogroup continuity between the people of the Yamnaya culture and the contemporary populations of Europe. Klejn has also suggested that the autosomal evidence does not support a Yamnaya migration, arguing that Western Steppe Herder ancestry in both contemporary and Bronze Age samples is lowest around the Danube in Hungary, near the western limits of the Yamnaya culture, and highest in Northern Europe, which Klejn argues is the opposite of what would be expected if the geneticists' hypothesis is correct.[69]

Language edit

 
Illustration of the closely related Afanasievo culture

Marija Gimbutas identified the Yamnaya culture with the late Proto-Indo-Europeans (PIE) in her Kurgan hypothesis. In the view of David Anthony, the Pontic-Caspian steppe is the strongest candidate for the Urheimat (original homeland) of the Proto-Indo-European language, citing evidence from linguistics and genetics[10][70] which suggests that the Yamnaya culture may be the homeland of the Indo-European languages, with the possible exception of the Anatolian languages.[71][72] On the other hand, Colin Renfrew has argued for a Near Eastern origin of the earliest Indo-European speakers.[73]

According to David W. Anthony, the genetic evidence suggests that the leading clans of the Yamnaya were of EHG (Eastern European hunter-gatherer) and WHG (Western European hunter-gatherer) paternal origin[74] and implies that the Indo-European languages were the result of "a dominant language spoken by EHGs that absorbed Caucasus-like elements in phonology, morphology, and lexicon."[75] It has also been suggested that the PIE language evolved through trade interactions in the circum-Pontic area in the 4th millennium BCE, mediated by the Yamna predecessors in the North Pontic steppe.[76]

Guus Kroonen et al. 2022 found that the "basal Indo-European stage", also known as Indo-Anatolian or Pre-Proto-Indo-European language, largely but not totally, lacked agricultural-related vocabulary, and only the later "core Indo-European languages" saw an increase in agriculture-associated words. According to them, this fits a homeland of early core Indo-European within the westernmost Yamnaya horizon, around and west of the Dnieper, while its basal stage, Indo-Anatolian, may have originated in the Sredny Stog culture, as opposed to the eastern Yamnaya horizon. The Corded Ware culture may have acted as major source for the spread of later Indo-European languages, including Indo-Iranian, while Tocharian languages may have been mediated via the Catacomb culture. They also argue that this new data contradicts a possible earlier origin of Pre-Proto-Indo-European among agricultural societies South of the Caucasus, rather "this may support a scenario of linguistic continuity of local non-mobile herders in the Lower Dnieper region and their genetic persistence after their integration into the successive and expansive Yamnaya horizon". Furthermore the authors mention that this scenario can explain the difference in paternal haplogroup frequency between the Yamnaya and Corded Ware cultures, while both sharing similar autosomal DNA ancestry.[77]

Yamnaya-related migrations edit

 
Scheme of Indo-European dispersals from a Yamanaya-Western Steppe Herders homeland (), c. 4000 to 1000 BCE, according to the widely held Steppe hypothesis.

Western Europe edit

Genetic studies have found that Yamnaya autosomal characteristics are very close to the Corded Ware culture people, with up to 75% Yamnaya-like ancestry in the DNA of Corded Ware skeletons from Central and Eastern Europe.[78] Yamnaya–related ancestry is found in the DNA of modern Central, and Northern Europeans (c. 38.8–50.4 %), and is also found in lower levels in present-day Southern Europeans (c. 18.5–32.6 %), Sardinians (c. 2.4–7.1 %), and Sicilians (c. 5.9–11.6 %).[79][70][12]

However, according to Heyd, et al. (2023), the specific paternal DNA haplogroup that is most commonly found in male Yamnaya specimens cannot be found in modern Western Europeans, or in males from the nearby Corded Ware culture. This makes it unlikely that the Corded Ware culture can be directly descended from the Yamnaya culture, at least along the paternal line.[80]

Autosomal tests also indicate that the Yamnaya are the vector for "Ancient North Eurasian" admixture into Europe.[10] "Ancient North Eurasian" is the name given in literature to a genetic component that represents descent from the people of the Mal'ta–Buret' culture[10] or a population closely related to them. That genetic component is visible in tests of the Yamnaya people[10] as well as modern-day Europeans. [81]

Eastern Europe and Finland edit

 
According to Allentoft (2015), the Sintashta culture probably derived from the Corded Ware Culture.

In the Baltic, Jones et al. (2017) found that the Neolithic transition – the passage from a hunter-gatherer economy to a farming-based economy – coincided with the arrival en masse of individuals with Yamnaya-like ancestry. This is different from what happened in Western and Southern Europe, where the Neolithic transition was caused by a population that came from Anatolia, with Pontic steppe ancestry being detected from only the late Neolithic onward.[82]

Per Haak et al. (2015), the Yamnaya contribution in the modern populations of Eastern Europe ranges from 46.8% among Russians to 42.8% in Ukrainians. Finland has the highest Yamnaya contributions in all of Europe (50.4%).[83][e]

Central and South Asia edit

 
Map of the approximate maximal extent of the Andronovo culture. The formative Sintashta-Petrovka culture is shown in darker red. The location of the earliest spoke-wheeled chariot finds is indicated in purple. Adjacent and overlapping cultures (Afanasevo, Srubna, Bactria-Margiana Culture are shown in green.
 
Archaeological cultures associated with Indo-Iranian migrations and Indo-Aryan migrations (after EIEC). The Andronovo, BMAC and Yaz cultures have often been associated with Indo-Iranian migrations. The GGC, Cemetery H, Copper Hoard and PGW cultures are candidates for cultures associated with Indo-Aryan migrations.

Studies also point to the strong presence of Yamnaya descent in the current nations of South Asia, especially in groups that are referred to as Indo-Aryans.[84][85] According to Pathak et al. (2018), the "North-Western Indian & Pakistani" populations (PNWI) showed significant Middle-Late Bronze Age Steppe (Steppe_MLBA) ancestry along with Yamnaya Early-Middle Bronze Age (Steppe_EMBA) ancestry, but the Indo-Europeans of Gangetic Plains and Dravidian people only showed significant Yamnaya (Steppe_EMBA) ancestry and no Steppe_MLBA. The study also noted that ancient south Asian samples had significantly higher Steppe_MLBA than Steppe_EMBA (or Yamnaya). The study identified the Rors as the population in South Asia with the highest proportion of Steppe ancestry.[85] Lazaridis et al. (2016) estimated (6.5–50.2 %) steppe-related admixture in South Asians, though the proportion of Steppe ancestry varies widely across ethnic groups.[47][f] According to Narasimhan et al. (2019), the Yamnaya-related ancestry, termed Western_Steppe_EMBA, that reached central and south Asia was not the initial expansion from the steppe to the east, but a secondary expansion that involved a group possessing ~67% Western_Steppe_EMBA ancestry and ~33% ancestry from the European cline. This group included people similar to that of Corded Ware, Srubnaya, Petrovka, and Sintashta. Moving further east in the central steppe, it acquired ~9% ancestry from a group of people that possessed West Siberian Hunter Gatherer ancestry, thus forming the Central Steppe MLBA cluster, which is the primary source of steppe ancestry in South Asia, contributing up to 30% of the ancestry of the modern groups in the region.[84]

According to Unterländer et al. (2017), all Iron Age Scythian Steppe nomads can best be described as a mixture of Yamnaya-related ancestry and an East Asian-related component, which most closely corresponds to the modern North Siberian Nganasan people of the lower Yenisey River, to varying degrees, but generally higher among Eastern Scythians. [86]

See also edit

Notes edit

  1. ^ /ˈjæmnə/ YAM-ny-yə; Russian: Ямная культура, romanizedYámnaya kultúra, pronounced [ˈjamnəjə kʊlʲˈturə], from яма "pit, hole"
  2. ^ /ˈjæmnə/ YAM-nə; Ukrainian: Ямна культура, romanizedJamna kuľtura, Ukrainian pronunciation: [ˈjamnɐ kʊlʲˈturɐ], lit.'culture of pits'
  3. ^ Yamnayan cultural aspects, for example, were horse-riding, burial styles, and to some extent the pastoralist economy.
  4. ^ The Eastern European hunter-gatherers were themselves mostly descended from ancient North Eurasians, related to the palaeolithic Mal'ta–Buret' culture.
  5. ^ Per Haak et al. (2015), adding a north-Siberian people as a fourth reference population improves residuals for northeastern European populations. This accounts for the higher than expected Yamnaya contribution and brings it down to expected levels (67.8–50.4 % in Finns, 64.9–46.8 % in Russians).
  6. ^ Lazaridis et al. (2016) Supplementary Information, Table S9.1: "Kalash – 50.2 %, Tiwari Brahmins – 44.1 %, Gujarati (four samples) – 46.1 % to 27.5 %, Pathan – 44.6 %, Burusho – 42.5 %, Sindhi – 37.7 %, Punjabi – 32.6 %, Balochi – 32.4 %, Brahui – 30.2 %, Lodhi – 29.3 %, Bengali – 24.6 %, Vishwabhramin – 20.4 %, Makrani – 19.2 %, Mala – 18.4 %, Kusunda – 8.9 %, Kharia – 6.5 %."

References edit

  1. ^ a b c Allentoft 2015.
  2. ^ a b c Morgunova & Khokhlova 2013.
  3. ^ Shishlina, Natalia I. (11 May 2023), Kristiansen, Kristian; Kroonen, Guus; Willerslev, Eske (eds.), "Yamnaya Pastoralists in the Eurasian Desert Steppe Zone: New Perspectives on Mobility", The Indo-European Puzzle Revisited (1 ed.), Cambridge University Press, pp. 34–41, doi:10.1017/9781009261753.006, ISBN 978-1-00-926175-3, retrieved 2023-05-23
  4. ^ a b c Anthony, David W. (11 May 2023), Kristiansen, Kristian; Kroonen, Guus; Willerslev, Eske (eds.), "The Yamnaya Culture and the Invention of Nomadic Pastoralism in the Eurasian Steppes", The Indo-European Puzzle Revisited (1 ed.), Cambridge University Press, pp. 13–33, doi:10.1017/9781009261753.005, ISBN 978-1-00-926175-3, retrieved 2023-05-13
  5. ^ Novembre 2015, "evidence to support theories of a back-migration from Corded Ware-related populations that contributed to the origins of the Sintashta culture in the Urals and their descendants, the Andronovo."
  6. ^ Hermes, Taylor R.; Tishkin, Alexey A.; Kosintsev, Pavel A.; Stepanova, Nadezhda F.; Krause-Kyora, Ben; Makarewicz, Cheryl A. (1 December 2020). "Mitochondrial DNA of domesticated sheep confirms pastoralist component of Afanasievo subsistence economy in the Altai Mountains (3300-2900 cal BC)". Archaeological Research in Asia. 24: 100232. doi:10.1016/j.ara.2020.100232. ISSN 2352-2267. S2CID 225136827.
  7. ^ a b c "Europe's fourth ancestral 'tribe' uncovered". BBC. 16 November 2015.
  8. ^ Wang, Chuan-Chao; Reinhold, Sabine; Kalmykov, Alexey; Wissgott, Antje; Brandt, Guido; Jeong, Choongwon; Cheronet, Olivia; Ferry, Matthew; Harney, Eadaoin; Keating, Denise; Mallick, Swapan (4 February 2019). "Ancient human genome-wide data from a 3000-year interval in the Caucasus corresponds with eco-geographic regions". Nature Communications. 10 (1): 590. Bibcode:2019NatCo..10..590W. doi:10.1038/s41467-018-08220-8. ISSN 2041-1723. PMC 6360191. PMID 30713341.
  9. ^ a b Lazaridis, Iosif; Alpaslan-Roodenberg, Songül; Acar, Ayşe; Açıkkol, Ayşen; Agelarakis, Anagnostis; Aghikyan, Levon; Akyüz, Uğur; Andreeva, Desislava; Andrijašević, Gojko; Antonović, Dragana; Armit, Ian; Atmaca, Alper; Avetisyan, Pavel; Aytek, Ahmet İhsan; Bacvarov, Krum (26 August 2022). "The genetic history of the Southern Arc: A bridge between West Asia and Europe". Science. 377 (6609): eabm4247. doi:10.1126/science.abm4247. ISSN 0036-8075. PMC 10064553. PMID 36007055. S2CID 251843620.
  10. ^ a b c d e f Haak et al. 2015.
  11. ^ a b c Mathieson, et al. 2015.
  12. ^ a b Gibbons, Ann (10 June 2015). "Nomadic herders left a strong genetic mark on Europeans and Asians". Science. AAAS.
  13. ^ Nikitin, Alexey G.; Ivanova, Svetlana; Kiosak, Dmytro; Badgerow, Jessica; Pashnick, Jeff (June 2017). "Subdivisions of haplogroups U and C encompass mitochondrial DNA lineages of Eneolithic–Early Bronze Age Kurgan populations of western North Pontic steppe". Journal of Human Genetics. 62 (6): 605–613. doi:10.1038/jhg.2017.12. ISSN 1435-232X. PMID 28148921. S2CID 7459815.
  14. ^ Nikitin, Alexey G.; Potekhina, Inna; Rohland, Nadin; Mallick, Swapan; Reich, David; Lillie, Malcolm (24 February 2017). "Mitochondrial DNA analysis of eneolithic trypillians from Ukraine reveals neolithic farming genetic roots". PLOS ONE. 12 (2): e0172952. Bibcode:2017PLoSO..1272952N. doi:10.1371/journal.pone.0172952. ISSN 1932-6203. PMC 5325568. PMID 28235025.
  15. ^ Anthony 2007, p. 300-370.
  16. ^ Nordgvist & Heyd 2020.
  17. ^ Mallory 1999.
  18. ^ a b Mallory 1999, p. 215.
  19. ^ Dolukhanov 1996, p. 94.
  20. ^ Anthony 2007, p. 182.
  21. ^ a b Anthony 2007, p. 275.
  22. ^ Anthony 2007, p. 300.
  23. ^ Mallory 1999, p. 210-211.
  24. ^ a b Anthony 2007, p. 321.
  25. ^ Anthony 2007, pp. 274–277, 317–320.
  26. ^ Anthony 2007, pp. 301–302.
  27. ^ Anthony 2007, p. 303.
  28. ^ Parpola 2015, p. 49.
  29. ^ Parpola 2015, p. 45.
  30. ^ Parpola 2015, p. 47.
  31. ^ Telegin, D. Y. (1973). Serednʹo-stohivsʹka kulʹtura epokhy midi. Kyiv: Naukova Dumka. p. 147.
  32. ^ Mallory 1997, p. 212.
  33. ^ Anthony 2007, p. 339.
  34. ^ Anthony 2007, p. 319.
  35. ^ Anthony, David. The Horse, the Wheel, and Language. pp. 328–329. OCLC 1102387902.
  36. ^ a b J., Harrison, Richard. The transformation of Europe in the third millennium BC: the example of "Le Petit-Chasseur I + II" (Sion, Valais, Switzerland). p. 196. OCLC 718304072.{{cite book}}: CS1 maint: multiple names: authors list (link)
  37. ^ Anthony 2007, p. 305.
  38. ^ a b Trautmann, Martin (2023). "First bioanthropological evidence for Yamnaya horsemanship". Science Advances. 9 (9): eade2451. Bibcode:2023SciA....9E2451T. doi:10.1126/sciadv.ade2451. hdl:10138/356326. PMID 36867690. S2CID 257334477.
  39. ^ P., Mallory, J. (2003) [1989]. In search of the Indo-Europeans : language, archaeology, and myth. Thames and Hudson. p. 213. ISBN 0-500-27616-1. OCLC 886668216.{{cite book}}: CS1 maint: multiple names: authors list (link)
  40. ^ Gerling, Claudia (1 July 2015). Prehistoric Mobility and Diet in the West Eurasian Steppes 3500 to 300 BC: An Isotopic Approach. De Gruyter. doi:10.1515/9783110311211. ISBN 978-3-11-031121-1.
  41. ^ Anthony 2007, p. 430.
  42. ^ J. P. Mallory; Douglas Q. Adams, eds. (1997). Encyclopedia of Indo-European culture. London: Fitzroy Dearborn. p. 653. ISBN 1-884964-98-2. OCLC 37931209.
  43. ^ Librado, Pablo (2021). "The origins and spread of domestic horses from the Western Eurasian steppes". Nature. 598 (7882): 634–640. Bibcode:2021Natur.598..634L. doi:10.1038/s41586-021-04018-9. PMC 8550961. PMID 34671162.
  44. ^ Hanel, Andrea; Carlberg, Carsten (3 July 2020). "Skin colour and vitamin D: An update". Experimental Dermatology. 29 (9): 864–875. doi:10.1111/exd.14142. PMID 32621306.
  45. ^ Wang, Chuan-Chao; Reinhold, Sabine; Kalmykov, Alexey (4 February 2019). "Ancient human genome-wide data from a 3000-year interval in the Caucasus corresponds with eco-geographic regions". Nature Communications. 10 (1): 590. Bibcode:2019NatCo..10..590W. doi:10.1038/s41467-018-08220-8. ISSN 2041-1723. PMC 6360191. PMID 30713341.
  46. ^ Jones et al. 2015.
  47. ^ a b Lazaridis et al. 2016.
  48. ^ Lazaridis, Iosif; Alpaslan-Roodenberg, Songül; Acar, Ayşe; Açıkkol, Ayşen; Agelarakis, Anagnostis; Aghikyan, Levon; Akyüz, Uğur; Andreeva, Desislava; Andrijašević, Gojko; Antonović, Dragana; Armit, Ian; Atmaca, Alper; Avetisyan, Pavel; Aytek, Ahmet İhsan; Bacvarov, Krum (26 August 2022). "The genetic history of the Southern Arc: A bridge between West Asia and Europe". Science. 377 (6609): eabm4247. doi:10.1126/science.abm4247. ISSN 0036-8075. PMC 10064553. PMID 36007055. S2CID 251843620.
  49. ^ Chintalapati, Manjusha; Patterson, Nick; Moorjani, Priya (30 May 2022). Perry, George H (ed.). "The spatiotemporal patterns of major human admixture events during the European Holocene". eLife. 11: e77625. doi:10.7554/eLife.77625. ISSN 2050-084X. PMC 9293011. PMID 35635751.
  50. ^ Balanovsky 2017, p. 437.
  51. ^ Wang 2019.
  52. ^ Eske, Allentoft; Morten E. Pokutta; Dalia Willerslev (2015). Population genomics of Bronze Age Eurasia. University of Copenhagen, Denmark. OCLC 1234973657.
  53. ^ Pilipenko, Aleksandr S.; Trapezov, Rostislav O.; Cherdantsev, Stepan V.; Babenko, Vladimir N.; Nesterova, Marina S.; Pozdnyakov, Dmitri V.; Molodin, Vyacheslav I.; Polosmak, Natalia V. (20 September 2018). "Maternal genetic features of the Iron Age Tagar population from Southern Siberia (1st millennium BC)". PLOS ONE. Public Library of Science (PLoS). 13 (9): e0204062. Bibcode:2018PLoSO..1304062P. doi:10.1371/journal.pone.0204062. ISSN 1932-6203. PMC 6147448. PMID 30235269.
  54. ^ Nikitin, Alexey G; Ivanova, Svetlana; Kiosak, Dmytro; Badgerow, Jessica; Pashnick, Jeff (2 February 2017). "Subdivisions of haplogroups U and C encompass mitochondrial DNA lineages of Eneolithic–Early Bronze Age Kurgan populations of western North Pontic steppe". Journal of Human Genetics. Springer Science and Business Media LLC. 62 (6): 605–613. doi:10.1038/jhg.2017.12. ISSN 1434-5161. PMID 28148921. S2CID 7459815. "In the 12 successfully haplotyped specimens, 75% of mtDNA lineages consisted of west Eurasian haplogroup U and its U4 and U5 sublineages. Furthermore, we identified a subgroup of east Eurasian haplogroup C in two representatives of the Yamna culture in one of the studied kurgans."
  55. ^ Gibbons, A. (24 July 2015). "Revolution in human evolution". Science. 349 (6246): 362–366. Bibcode:2015Sci...349..362G. doi:10.1126/science.349.6246.362. ISSN 0036-8075. PMID 26206910.
  56. ^ Hanel, Andrea; Carlberg, Carsten (September 2020). "Skin colour and vitamin D: An update". Experimental Dermatology. 29 (9): 864–875. doi:10.1111/exd.14142. ISSN 0906-6705. PMID 32621306.
  57. ^ Mathieson, Iain; Alpaslan-Roodenberg, Songül; Posth, Cosimo; Szécsényi-Nagy, Anna; Rohland, Nadin; Mallick, Swapan; Olalde, Iñigo; Broomandkhoshbacht, Nasreen; Candilio, Francesca; Cheronet, Olivia; Fernandes, Daniel (March 2018). "The genomic history of southeastern Europe". Nature. 555 (7695): 197–203. Bibcode:2018Natur.555..197M. doi:10.1038/nature25778. ISSN 1476-4687. PMC 6091220. PMID 29466330.
  58. ^ Heyd, Volker (April 2017). "Kossinna's smile". Antiquity. 91 (356): 348–359. doi:10.15184/aqy.2017.21. hdl:10138/255652. ISSN 0003-598X. S2CID 164376362.
  59. ^ Mathieson, Iain; Lazaridis, Iosif; Rohland, Nadin; Mallick, Swapan; Patterson, Nick; Roodenberg, Songül Alpaslan; Harney, Eadaoin; Stewardson, Kristin; Fernandes, Daniel; Novak, Mario; Sirak, Kendra (24 December 2015). "Genome-wide patterns of selection in 230 ancient Eurasians". Nature. 528 (7583): 499–503. Bibcode:2015Natur.528..499M. doi:10.1038/nature16152. ISSN 0028-0836. PMC 4918750. PMID 26595274.
  60. ^ Segurel, Laure (2020). "Why and when was lactase persistence selected for? Insights from Central Asian herders and ancient DNA". PLOS Biology. 18 (6): e3000742. doi:10.1371/journal.pbio.3000742. PMC 7302802. PMID 32511234. "Furthermore, ancient DNA studies found that the LP mutation was absent or very rare in Europe until the end of the Bronze Age [26–29] and appeared first in individuals with steppe ancestry [19,20]. Thus, it was proposed that the mutation originated in Yamnaya-associated populations and arrived later in Europe by migration of these steppe herders."
  61. ^ Callaway, Ewen. "DNA data explosion lights up the Bronze Age". Nature. "the 101 sequenced individuals, the Yamnaya were most likely to have the DNA variation responsible for lactose tolerance, hinting that the steppe migrants might have eventually introduced the trait to Europe"
  62. ^ Furholt, Martin (2018). "Massive Migrations? The Impact of Recent DNA Studies on our View of Third Millennium Europe". European Journal of Archaeology. 21 (2): 159–191. doi:10.1017/eaa.2017.43. "For example, one lineage could have a biological evolutionary advantage over the other. Allentoft et al. (2015: 171) have found a remarkably high rate of lactose tolerance among individuals connected to Yamnaya and to Corded Ware, as opposed to the majority of Late Neolithic individuals."
  63. ^ Saag, L (2020). "Human Genetics: Lactase Persistence in a Battlefield". Current Biology. 30 (21): R1311–R1313. doi:10.1016/j.cub.2020.08.087. PMID 33142099. S2CID 226229587.
  64. ^ Lazaridis et al. 2022c.
  65. ^ Reich, David (2018). Who we are and how we got here: ancient DNA and the new science of the human past. Oxford, United Kingdom. p. 121. ISBN 978-0-19-882125-0. OCLC 1006478846.{{cite book}}: CS1 maint: location missing publisher (link)
  66. ^ Reich, David (2018). Who we are and how we got here: ancient DNA and the new science of the human past. Oxford, United Kingdom. p. 239. ISBN 978-0-19-882125-0. OCLC 1006478846.{{cite book}}: CS1 maint: location missing publisher (link)
  67. ^ Klejn, Leo (2017). "The Steppe Hypothesis of Indo-European Origins Remains to be Proven". Acta Archaeologica. 88 (1): 193–204. doi:10.1111/j.1600-0390.2017.12184.x.
  68. ^ Balanovsky, O. (2017). "Genetic differentiation between upland and lowland populations shapes the Y-chromosomal landscape of West Asia". Human Genetics. 136 (4): 437–450. doi:10.1007/s00439-017-1770-2. PMID 28281087. S2CID 3735168. "The ancient Yamnaya samples are located on the "eastern" R-GG400 branch of haplogroup R1b-L23, showing that the paternal descendants of the Yamnaya still live in the Pontic steppe and that the ancient Yamnaya population was not an important source of paternal lineages in present-day West Europeans."
  69. ^ Klejn 2017, p. 201: "In the tables presented in the article by Reichs’ team (Haak et al. 2015) the genetic pool connecting the Yamnaya culture with the Corded Ware people is shown to be more intense in Northern Europe (Norway and Sweden) and decreases gradually from the North to the South (Fig. 6). It is weakest around the Danube, in Hungary, i. e. areas neighbouring the western branch of the Yamnaya culture! This is the reverse image to what the proposed hypothesis by the geneticists would lead us to expect. It is true that this gradient is traced back from the contemporary materials, but it was already present during the Bronze Age [...]"
  70. ^ a b Zimmer, Carl (10 June 2015). "DNA Deciphers Roots of Modern Europeans". New York Times. Retrieved 2020-12-12.
  71. ^ Olsen, Birgit A.; Olander, Thomas; Kristiansen, Kristian (23 August 2019), Tracing the Indo-Europeans, Oxbow Books, pp. 1–6, doi:10.2307/j.ctvmx3k2h.6, ISBN 978-1-78925-273-6, S2CID 202354040
  72. ^ Olsen, Birgit A. (11 May 2023), Kristiansen, Kristian; Kroonen, Guus; Willerslev, Eske (eds.), "Marriage Strategies and Fosterage among the Indo-Europeans: A Linguistic Perspective", The Indo-European Puzzle Revisited (1 ed.), Cambridge University Press, pp. 296–302, doi:10.1017/9781009261753.027, ISBN 978-1-00-926175-3, retrieved 2023-05-13
  73. ^ Schneider, Thomas (2023). Language Contact in Ancient Egypt. LIT Verlag Münster. p. 110-113. ISBN 978-3-643-91507-8.
  74. ^ Anthony 2019b, p. 36.
  75. ^ Anthony 2019a, p. 1-19.
  76. ^ Nikitin, Alexey; Ivanova, Svetlana (28 November 2022). "Long-distance exchanges along the Black Sea coast in the Eneolithic and the steppe genetic ancestry problem". Cambridge Open Engage. doi:10.33774/coe-2022-7m315.
  77. ^ Kroonen, Guus; Jakob, Anthony; Palmér, Axel I.; van Sluis, Paulus; Wigman, Andrew (12 October 2022). "Indo-European cereal terminology suggests a Northwest Pontic homeland for the core Indo-European languages". PLOS ONE. 17 (10): e0275744. Bibcode:2022PLoSO..1775744K. doi:10.1371/journal.pone.0275744. ISSN 1932-6203. PMC 9555676. PMID 36223379.
  78. ^ Stoneking, Mark; Arias, Leonardo; Liu, Dang; Oliveira, Sandra; Pugach, Irina; Rodriguez, Jae Joseph Russell B. (24 January 2023). "Genomic perspectives on human dispersals during the Holocene". Proceedings of the National Academy of Sciences. 120 (4): e2209475119. Bibcode:2023PNAS..12009475S. doi:10.1073/pnas.2209475119. ISSN 0027-8424. PMC 9942792. PMID 36649433.
  79. ^ Haak et al. 2015, pp. 121–124.
  80. ^ Kristiansen, Kristian; Kroonen, Guus; Willerslev, Eske (11 May 2023). The Indo-European Puzzle Revisited: Integrating Archaeology, Genetics, and Linguistics. Cambridge University Press. p. 70-76. ISBN 978-1-009-26174-6.
  81. ^ Lazaridis et al. 2014.
  82. ^ Jones et al. 2017.
  83. ^ Haak et al. 2015, pp. 121–122.
  84. ^ a b Narasimhan et al. 2019.
  85. ^ a b Pathak et al. 2018.
  86. ^ Unterländer et al. 2017Genomic inference reveals that Scythians in the east and the west of the steppe zone can best be described as a mixture of Yamnaya-related ancestry and an East Asian component. Demographic modelling suggests independent origins for eastern and western groups with ongoing gene-flow between them, plausibly explaining the striking uniformity of their material culture. We also find evidence that significant gene-flow from east to west Eurasia must have occurred early during the Iron Age." and "The blend of EHG [European hunter-gatherer] and Caucasian elements in carriers of the Yamnaya culture was formed on the European steppe and exported into Central Asia and Siberia". We therefore considered an alternative model in which we treat them as a mix of Yamnaya and the Han (Supplementary Table 25). This model fits all of the Iron Age Scythian groups, consistent with these groups having ancestry related to East Asians not found in the other populations. Alternatively, the Iron Age Scythian groups can also be modelled as a mix of Yamnaya and the north Siberian Nganasan (Supplementary Note 2, Supplementary Table 26).

Sources edit

  • Allentoft, Morten E.; et al. (2015). "Population genomics of Bronze Age Eurasia". Nature. 522 (7555): 167–172. Bibcode:2015Natur.522..167A. doi:10.1038/nature14507. PMID 26062507. S2CID 4399103.
  • Anthony, David W. (2007). The Horse, the Wheel, and Language: How Bronze-Age Riders from the Eurasian Steppes Shaped the Modern World. Princeton University Press. ISBN 978-0-691-05887-0.
  • Anthony, David (2017). "Archaeology and Language: Why Archaeologists Care About the Indo-European Problem". In Crabtree, P.J.; Bogucki, P. (eds.). European Archaeology as Anthropology: Essays in Memory of Bernard Wailes.
  • Anthony, David (Spring–Summer 2019a). "Archaeology, Genetics, and Language in the Steppes: A Comment on Bomhard". Journal of Indo-European Studies. 47 (1–2). Retrieved 2020-01-09.
  • Anthony, David W. (2019b). "Ancient DNA, Mating Networks, and the Anatolian Split". In Serangeli, Matilde; Olander, Thomas (eds.). Dispersals and Diversification: Linguistic and Archaeological Perspectives on the Early Stages of Indo-European. BRILL. pp. 21–54. ISBN 978-9004416192.
  • Cassidy LM, Martiniano R, Murphy EM, Teasdale MD, Mallory J, Hartwell B, Bradley DG (2016). "Neolithic and Bronze Age migration to Ireland and establishment of the insular Atlantic genome". PNAS. 113 (2): 368–373. Bibcode:2016PNAS..113..368C. doi:10.1073/pnas.1518445113. PMC 4720318. PMID 26712024.
  • Dolukhanov, Pavel M. (1996), The Early Slavs: Eastern Europe from the Initial Settlement to the Kievan Rus, New York: Longman, ISBN 0-582-23627-4
  • Fortson, Benjamin W. (2004), Indo-European Language and Culture: An Introduction, Blackwell Publishing
  • Gallego-Llorente M, Connell S, Jones ER, Merrett DC, Jeon Y, Eriksson A, et al. (2016). "The genetics of an early Neolithic pastoralist from the Zagros, Iran". Scientific Reports. 6: 31326. Bibcode:2016NatSR...631326G. doi:10.1038/srep31326. PMC 4977546. PMID 27502179.
  • Haak W, Lazaridis I, Patterson N, Rohland N, Mallick S, Llamas B, et al. (2015). "Massive migration from the steppe was a source for Indo-European languages in Europe". Nature. 522 (7555): 207–211. arXiv:1502.02783. Bibcode:2015Natur.522..207H. bioRxiv 10.1101/013433. doi:10.1038/nature14317. PMC 5048219. PMID 25731166.
  • Jeong, Choongwon; Balanovsky, Oleg; Lukianova, Elena; Kahbatkyzy, Nurzhibek; et al. (29 April 2019). "The genetic history of admixture across inner Eurasia languages in Europe". Nature Ecology and Evolution. Nature Research. 3 (6): 966–976. doi:10.1038/s41559-019-0878-2. hdl:10871/36562. PMC 6542712. PMID 31036896.
  • Jones, Eppie R.; Gonzalez-Fortes, Gloria; Connell, Sarah; Siska, Veronika; et al. (2015). "Upper Palaeolithic genomes reveal deep roots of modern Eurasians". Nature Communications. 6: 8912. Bibcode:2015NatCo...6.8912J. doi:10.1038/ncomms9912. PMC 4660371. PMID 26567969.
  • Jones ER, Zarina G, Moiseyev V, Lightfoot E, Nigst PR, Manica A, Pinhasi R, Bradley DG (2017). "The Neolithic transition in the Baltic was not driven by admixture with early European farmers". Current Biology. 27 (4): 576–582. doi:10.1016/j.cub.2016.12.060. PMC 5321670. PMID 28162894.
  • Kuzmina, Elena E. (2007). Mallory, J. P. (ed.). The Origin of the Indo-Iranians. BRILL. ISBN 978-9004160545.
  • Lazaridis, Iosif; Patterson, Nick; Mittnik, Alissa; Renaud, Gabriel; et al. (2014). "Ancient human genomes suggest three ancestral populations for present-day Europeans". Nature. 513 (7518): 409–413. arXiv:1312.6639. Bibcode:2014Natur.513..409L. doi:10.1038/nature13673. PMC 4170574. PMID 25230663.
  • Lazaridis I, Nadel D, Rollefson G, Merrett DC, Rohland N, Mallick S, et al. (16 June 2016). "The genetic structure of the world's first farmers". bioRxiv 10.1101/059311. Supplementary Information
  • Lazaridis I, Nadel D, Rollefson G, Merrett DC, Rohland N, Mallick S, et al. (25 July 2016). "Genomic insights into the origin of farming in the ancient Near East". Nature (published August 2016). 536 (7617): 419–424. Bibcode:2016Natur.536..419L. doi:10.1038/nature19310. PMC 5003663. PMID 27459054.
  • Lazaridis, Iosif; Alpaslan-Roodenberg, Songül; Acar, Ayşe; Açıkkol, Ayşen; Agelarakis, Anagnostis; Aghikyan, Levon; Akyüz, Uğur; Andreeva, Desislava; Andrijašević, Gojko; Antonović, Dragana; Armit, Ian; Atmaca, Alper; Avetisyan, Pavel; Aytek, Ahmet İhsan; Bacvarov, Krum (26 August 2022). "A genetic probe into the ancient and medieval history of Southern Europe and West Asia". Science. 377 (6609): 940–951. Bibcode:2022Sci...377..940L. doi:10.1126/science.abq0755. ISSN 0036-8075. PMC 10019558. PMID 36007020. S2CID 251844202.
  • Mallory, J. P. (1997), "Yamna Culture", Encyclopedia of Indo-European Culture, Fitzroy Dearborn
  • Mallory, J.P. (1999), In Search of the Indo-Europeans: Language, Archaeology, and Myth (reprint ed.), London: Thames & Hudson, ISBN 978-0-500-27616-7
  • Martiniano, R, et al. (2017). "The population genomics of archaeological transition in west Iberia: Investigation of ancient substructure using imputation and haplotype-based methods". PLOS Genet. 13 (7): e1006852. doi:10.1371/journal.pgen.1006852. PMC 5531429. PMID 28749934.
  • Mathieson, Iain; et al. (10 October 2015). "Eight thousand years of natural selection in Europe". bioRxiv 10.1101/016477.
    Mathieson, Iain; et al. (23 November 2015). "Genome-wide patterns of selection in 230 ancient Eurasians". Nature (published 24 December 2015). 528 (7583): 499–503. Bibcode:2015Natur.528..499M. doi:10.1038/nature16152. PMC 4918750. PMID 26595274.
  • Mathieson, Iain; et al. (21 February 2018). "The Genomic History of Southeastern Europe". Nature. Nature Research. 555 (7695): 197–203. Bibcode:2018Natur.555..197M. doi:10.1038/nature25778. PMC 6091220. PMID 29466330.
  • Morgunova, Nina; Khokhlova, Olga (2013). "Chronology and Periodization of the Pit-Grave Culture in the Area Between the Volga and Ural Rivers Based on 14C Dating and Paleopedological Research". Radiocarbon. 55 (2–3): 1286–1296. doi:10.2458/azu_js_rc.55.16087. ISSN 0033-8222.
  • Narasimhan VM, Patterson N, Moorjani P, Rohland N, et al. (2019). "The formation of human populations in South and Central Asia". Science. 365 (6457): eaat7487. doi:10.1126/science.aat7487. PMC 6822619. PMID 31488661.
  • Nordgvist, Kerkko; Heyd, Volker (2020). "The Forgotten Child of the Wider Corded Ware Family: Russian Fatyanovo Culture in Context". Proceedings of the Prehistoric Society. 86: 65–93. doi:10.1017/ppr.2020.9. S2CID 228923806.
  • Novembre, John (11 June 2015). "Ancient DNA steps into the language debate". Nature. 522 (7555): 164–165. doi:10.1038/522164a. PMID 26062506. S2CID 205085294.
  • Parpola, Asko (2015), The Roots of Hinduism, Oxford University Press
  • Pashnick, Jeff (August 2014). Genetic Analysis of Ancient Human Remains from the Early Bronze Age Cultures of the North PonticSteppe Region (Masters Theses thesis). Vol. 737. Grand Valley State University. Retrieved 2020-01-12.
  • Pathak AK, Kadian A, Kushniarevich A, Montinaro F, Mondal M, Ongaro L, et al. (6 December 2018). "The Genetic Ancestry of Modern Indus Valley Populations from Northwest India". The American Journal of Human Genetics. 103 (6): 918–929. doi:10.1016/j.ajhg.2018.10.022. PMC 6288199. PMID 30526867.
  • Unterländer, Martina; Palstra, Friso; Lazaridis, Iosif; Pilipenko, Aleksandr; Hofmanová, Zuzana; Groß, Melanie; et al. (2017). "Ancestry and demography and descendants of Iron Age nomads of the Eurasian Steppe". Nature Communications. 8: 14615. Bibcode:2017NatCo...814615U. doi:10.1038/ncomms14615. ISSN 2041-1723. PMC 5337992. PMID 28256537.
  • Wang, Chuan-Chao (4 February 2019). "Ancient human genome-wide data from a 3000-year interval in the Caucasus corresponds with eco-geographic regions Eurasia". Nature Communications. Nature Research. 10 (1): 590. Bibcode:2019NatCo..10..590W. bioRxiv 10.1101/322347. doi:10.1038/s41467-018-08220-8. PMC 6360191. PMID 30713341.
  • Wilde S, Timpson A, Kirsanow K, Kaiser E, Kayser M, Unterländer M, et al. (2014). "Direct evidence for positive selection of skin, hair, and eye pigmentation in Europeans during the last 5,000 y". Proceedings of the National Academy of Sciences of the United States of America. 111 (13): 4832–4837. Bibcode:2014PNAS..111.4832W. doi:10.1073/pnas.1316513111. PMC 3977302. PMID 24616518.

External links edit

  • "Genetic study revives debate on origin and expansion of Indo-European Languages". Science Daily. March 2015.
  • "First Horse Warriors". PBS.org NOVA series. 15 May 2019.

yamnaya, culture, yamna, culture, redirects, here, confused, with, yamna, language, papua, indonesia, yamna, culture, also, known, grave, culture, ochre, grave, culture, late, copper, early, bronze, archaeological, culture, region, between, southern, dniester,. Yamna culture redirects here Not to be confused with the Yamna language of Papua Indonesia The Yamnaya culture a or the Yamna culture b also known as the Pit Grave culture or Ochre Grave culture was a late Copper Age to early Bronze Age archaeological culture of the region between the Southern Bug Dniester and Ural rivers the Pontic Caspian steppe dating to 3300 2600 BCE 2 It was discovered by Vasily Gorodtsov following his archaeological excavations near the Donets River in 1901 1903 Its name derives from its characteristic burial tradition Ya mnaya romanization yamnaya is a Russian adjective that means related to pits yama as these people used to bury their dead in tumuli kurgans containing simple pit chambers Yamnaya cultureAlternative namesPit Grave culture Yamna culture Ochre Grave culture Yamnaya HorizonGeographical rangePontic Caspian steppe in EuropePeriodCopper Age Bronze AgeDatesc 3300 2600 BCEPreceded bySamara culture Khvalynsk culture Dnieper Donets culture Sredny Stog culture Repin culture Maykop culture Cucuteni Trypillia culture Cernavodă culture Usatove cultureFollowed byNorth Corded Ware culture 1 West Catacomb culture East Poltavka cultureDefined byVasily GorodtsovThe Yamnaya economy was based upon animal husbandry fishing and foraging and the manufacture of ceramics tools and weapons 3 The people of the Yamnaya culture lived primarily as nomads with a chiefdom system and wheeled carts and wagons that allowed them to manage large herds 4 They are also closely connected to Final Neolithic cultures which later spread throughout Europe and Central Asia especially the Corded Ware people and the Bell Beaker culture 4 as well as the peoples of the Sintashta Andronovo and Srubnaya cultures Back migration from Corded Ware also contributed to Sintashta and Andronovo 5 In these groups several aspects of the Yamnaya culture are present c Yamnaya material culture was very similar to the Afanasevo culture of South Siberia and the populations of both cultures are genetically indistinguishable 1 This suggests that the Afanasevo culture may have originated from the migration of Yamnaya groups to the Altai region or alternatively that both cultures developed from an earlier shared cultural source 6 Genetic studies have suggested that the people of the Yamnaya culture can be modelled as a genetic admixture between a population related to Eastern European Hunter Gatherers EHG d and people related to hunter gatherers from the Caucasus CHG in roughly equal proportions 7 an ancestral component which is often named Steppe ancestry with additional admixture from Anatolian Levantine or Early European farmers 8 9 Genetic studies also indicate that populations associated with the Corded Ware Bell Beaker Sintashta and Andronovo cultures derived large parts of their ancestry from the Yamnaya or a closely related population 1 10 11 12 Contents 1 Origins 2 Characteristics 3 Gallery 4 Archaeogenetics 5 Language 6 Yamnaya related migrations 6 1 Western Europe 6 2 Eastern Europe and Finland 6 3 Central and South Asia 7 See also 8 Notes 9 References 9 1 Sources 10 External linksOrigins editSee also Kurgan hypothesis and Marija Gimbutas nbsp Largest expansion of the Yamna ya culture Modified from 13 c 3500 origins of Usatovo culture 3300 origins of Yamna c 3300 3200 expansion of Yamna across the Pontic Caspian steppe c 2700 end of Trypillia culture 14 and transformation of Yamna into Corded Ware in the contact zone east of the Carpathian mountains 3100 2600 Yamnaya expansion into the Danube Valley 15 16 17 The Yamnaya culture was defined by Vasily Gorodtsov in order to differentiate it from the Catacomb and Srubnaya cultures that existed in the area but were considered to be of a later period Due to the time interval to the Yamnaya culture and the reliance on archaeological findings debate as to its origin is ongoing 18 In 1996 Pavel Dolukhanov suggested that the emergence of the Pit Grave culture represents a social development of various different local Bronze Age cultures thus representing an expression of social stratification and the emergence of chiefdom type nomadic social structures which in turn intensified inter group contacts between essentially heterogeneous social groups 19 The origin of the Yamnaya culture continues to be debated with proposals for its origins pointing to both the Khvalynsk and Sredny Stog cultures 18 The Khvalynsk culture 4700 3800 BCE 20 middle Volga and the Don based Repin culture c 3950 3300 BCE 21 in the eastern Pontic Caspian steppe and the closely related Sredny Stog culture c 4500 3500 BCE in the western Pontic Caspian steppe preceded the Yamnaya culture 3300 2500 BCE 22 23 Further efforts to pinpoint the location came from Anthony 2007 who suggested that the Yamnaya culture 3300 2600 BCE originated in the Don Volga area at c 3400 BC 24 2 preceded by the middle Volga based Khvalynsk culture and the Don based Repin culture c 3950 3300 BCE 21 2 arguing that late pottery from these two cultures can barely be distinguished from early Yamnaya pottery 25 Earlier continuity from eneolithic but largely hunter gatherer Samara culture and influences from the more agricultural Dnieper Donets II are apparent He argues that the early Yamnaya horizon spread quickly across the Pontic Caspian steppes between c 3400 and 3200 BCE 24 The spread of the Yamnaya horizon was the material expression of the spread of late Proto Indo European across the Pontic Caspian steppes 26 The Yamnaya horizon is the visible archaeological expression of a social adjustment to high mobility the invention of the political infrastructure to manage larger herds from mobile homes based in the steppes 27 Alternatively Parpola 2015 relates both the Corded ware culture and the Yamnaya culture to the late Trypillia Tripolye culture 28 He hypothesizes that the Tripolye culture was taken over by PIE speakers by c 4000 BCE 29 and that in its final phase the Trypillian culture expanded to the steppes morphing into various regional cultures which fused with the late Sredny Stog Serednii Stih pastoralist cultures which he suggests gave rise to the Yamnaya culture 30 Dmytro Telegin viewed Sredny Stog and Yamna as one cultural continuum and considered Sredny Stog to be the genetic foundation of the Yamna 31 The Yamnaya culture was succeeded in its western range by the Catacomb culture 2800 2200 BCE in the east by the Poltavka culture 2700 2100 BCE at the middle Volga These two cultures were followed by the Srubnaya culture 18th 12th century BCE Maps of the origins of Yamnaya culture nbsp Sredny Stog culture c 4500 3500 BCE nbsp Usatovo culture c 3500 3000 BCE nbsp Khvalynsk culture c 4900 3500 BCE nbsp Early Yamnaya culture 3400 BCE according to Anthony 2007 Characteristics edit nbsp Yamnaya culture grave Volgograd OblastThe Yamnaya culture was nomadic 4 or semi nomadic with some agriculture practiced near rivers and a few fortified sites the largest of which is Mikhaylivka 32 Characteristic for the culture are the burials in pit graves under kurgans tumuli often accompanied by animal offerings Some graves contain large anthropomorphic stelae with carved human heads arms hands belts and weapons 33 The dead bodies were placed in a supine position with bent knees and covered in ochre Some kurgans contained stratified sequences of graves 34 Kurgan burials may have been rare and were perhaps reserved for special adults who were predominantly but not necessarily male 35 Status and gender are marked by grave goods and position and in some areas elite individuals are buried with complete wooden wagons 36 Grave goods are more common in eastern Yamnaya burials which are also characterized by a higher proportion of male burials and more male centred rituals than western areas 37 The Yamnaya culture had and used two wheeled carts and four wheeled wagons which are thought to have been oxen drawn at this time and there is evidence that they rode horses 38 39 For instance several Yamnaya skeletons exhibit specific characteristics in their bone morphology that may have been caused by long term horseriding 38 Metallurgists and other craftsmen are given a special status in Yamnaya society and metal objects are sometimes found in large quantities in elite graves New metalworking technologies and weapon designs are used 36 Stable isotope ratios of Yamna individuals from the Dnipro Valley suggest the Yamnaya diet was terrestrial protein based with insignificant contribution from freshwater or aquatic resources 40 Anthony speculates that the Yamnaya ate meat milk yogurt cheese and soups made from seeds and wild vegetables and probably consumed mead 41 Mallory and Adams suggest that Yamnaya society may have had a tripartite structure of three differentiated social classes although the evidence available does not demonstrate the existence of specific classes such as priests warriors and farmers 42 Gallery edit nbsp Corded ware pot nbsp Daggers arrowheads and bone artefacts nbsp Yamanaya decorative artifacts nbsp The Kernosivsky idol late Yamnaya nbsp Western Yamnaya artefacts nbsp Illustration of a Yamnaya wagon nbsp Metal artefacts nbsp Metal artefacts nbsp Northern Caucasus Early Bronze Age artifacts 3rd millennium BCE nbsp Ceramic vessel nbsp Cis Ural Yamnaya artefacts and burials nbsp Horses were domesticated on the Pontic Caspian steppe 43 Archaeogenetics editFurther information Western Steppe Herders nbsp Main genetic ancestry of Western Steppe Herders Yamnaya pastoralists a confluence of Eastern Hunter Gatherers EHG and Caucasus Hunter Gatherers CHG 44 nbsp Admixture proportions of Yamnaya populations They combined Eastern Hunter Gatherer EHG Caucasian Hunter Gatherer CHG Anatolian Neolithic and Western Hunter Gatherer WHG ancestry 45 According to Jones et al 2015 and Haak et al 2015 autosomal tests indicate that the Yamnaya people were the result of a genetic admixture between two different hunter gatherer populations distinctive Eastern Hunter Gatherers EHG from Eastern Europe with high affinity to the Mal ta Buret culture or other closely related people from Siberia 10 and a population of Caucasus hunter gatherers CHG who probably arrived from the Caucasus 46 7 or Iran 47 Each of those two populations contributed about half the Yamnaya DNA 11 7 This admixture is referred to in archaeogenetics as Western Steppe Herder WSH ancestry Admixture between EHGs and CHGs is believed to have occurred on the eastern Pontic Caspian steppe starting around 5 000 BCE while admixture with Early European Farmers EEF happened in the southern parts of the Pontic Caspian steppe sometime later More recent genetic studies have found that the Yamnaya were a mixture of EHGs CHGs and to a lesser degree Anatolian farmers and Levantine farmers but not EEFs from Europe due to lack of WHG DNA in the Yamnaya This occurred in two distinct admixture events from West Asia into the Pontic Caspian steppe 48 49 Haplogroup R1b specifically the Z2103 suBCElade of R1b L23 is the most common Y DNA haplogroup found among the Yamnaya specimens This haplogroup is rare in Western Europe and mainly exists in Southeastern Europe today 50 Additionally a minority are found to belong to haplogroup I2 11 They are found to belong to a wider variety of West Eurasian mtDNA haplogroups including U T and haplogroups associated with Caucasus Hunter Gatherers and Early European Farmers 51 52 A small but significant number of Yamnaya kurgan specimens from Northern Ukraine carried the East Asian mtDNA haplogroup C4 53 54 People of the Yamnaya culture are believed to have had mostly brown eye colour light to intermediate skin and brown hair colour with some variation 55 56 Some Yamnaya individuals are believed to have carried a mutation to the KITLG gene associated with blond hair as several individuals with Steppe ancestry are later found to carry this mutation The Ancient North Eurasian Afontova Gora group who contributed significant ancestry to Western Steppe Herders are believed to be the source of this mutation 57 A study in 2015 found that Yamnaya had the highest ever calculated genetic selection for height of any of the ancient populations tested 58 59 It has been hypothesized that an allele associated with lactase persistence conferring lactose tolerance into adulthood was brought to Europe from the steppe by Yamnaya related migrations 60 61 62 63 A 2022 study by Lazaridis et al found that the typical phenotype among the Yamnaya population was brown eyes brown hair and intermediate skin colour None of their Yamnaya samples were predicted to have either blue eyes or blond hair in contrast with later Steppe groups in Russia and Central Asia as well as the Bell Beaker culture in Europe who did carry these phenotypes in high proportions 64 The geneticist David Reich has argued that the genetic data supports the likelihood that the people of the Yamnaya culture were a single genetically coherent group who were responsible for spreading many Indo European languages 65 Reich s group recently suggested that the source of Anatolian and Indo European subfamilies of the Proto Indo European PIE language may have been in west Asia and the Yamna were responsible for the dissemination of the latter 9 Reich also argues that the genetic evidence shows that Yamnaya society was an oligarchy dominated by a small number of elite males 66 The genetic evidence for the extent of the role of the Yamnaya culture in the spread of Indo European languages has been questioned by Russian archaeologist Leo Klejn 67 and Balanovsky et al 68 who note a lack of male haplogroup continuity between the people of the Yamnaya culture and the contemporary populations of Europe Klejn has also suggested that the autosomal evidence does not support a Yamnaya migration arguing that Western Steppe Herder ancestry in both contemporary and Bronze Age samples is lowest around the Danube in Hungary near the western limits of the Yamnaya culture and highest in Northern Europe which Klejn argues is the opposite of what would be expected if the geneticists hypothesis is correct 69 Language edit nbsp Illustration of the closely related Afanasievo cultureMarija Gimbutas identified the Yamnaya culture with the late Proto Indo Europeans PIE in her Kurgan hypothesis In the view of David Anthony the Pontic Caspian steppe is the strongest candidate for the Urheimat original homeland of the Proto Indo European language citing evidence from linguistics and genetics 10 70 which suggests that the Yamnaya culture may be the homeland of the Indo European languages with the possible exception of the Anatolian languages 71 72 On the other hand Colin Renfrew has argued for a Near Eastern origin of the earliest Indo European speakers 73 According to David W Anthony the genetic evidence suggests that the leading clans of the Yamnaya were of EHG Eastern European hunter gatherer and WHG Western European hunter gatherer paternal origin 74 and implies that the Indo European languages were the result of a dominant language spoken by EHGs that absorbed Caucasus like elements in phonology morphology and lexicon 75 It has also been suggested that the PIE language evolved through trade interactions in the circum Pontic area in the 4th millennium BCE mediated by the Yamna predecessors in the North Pontic steppe 76 Guus Kroonen et al 2022 found that the basal Indo European stage also known as Indo Anatolian or Pre Proto Indo European language largely but not totally lacked agricultural related vocabulary and only the later core Indo European languages saw an increase in agriculture associated words According to them this fits a homeland of early core Indo European within the westernmost Yamnaya horizon around and west of the Dnieper while its basal stage Indo Anatolian may have originated in the Sredny Stog culture as opposed to the eastern Yamnaya horizon The Corded Ware culture may have acted as major source for the spread of later Indo European languages including Indo Iranian while Tocharian languages may have been mediated via the Catacomb culture They also argue that this new data contradicts a possible earlier origin of Pre Proto Indo European among agricultural societies South of the Caucasus rather this may support a scenario of linguistic continuity of local non mobile herders in the Lower Dnieper region and their genetic persistence after their integration into the successive and expansive Yamnaya horizon Furthermore the authors mention that this scenario can explain the difference in paternal haplogroup frequency between the Yamnaya and Corded Ware cultures while both sharing similar autosomal DNA ancestry 77 Yamnaya related migrations editSee also Indo European migrations nbsp Scheme of Indo European dispersals from a Yamanaya Western Steppe Herders homeland c 4000 to 1000 BCE according to the widely held Steppe hypothesis Western Europe edit See also Corded Ware culture Genetic studies have found that Yamnaya autosomal characteristics are very close to the Corded Ware culture people with up to 75 Yamnaya like ancestry in the DNA of Corded Ware skeletons from Central and Eastern Europe 78 Yamnaya related ancestry is found in the DNA of modern Central and Northern Europeans c 38 8 50 4 and is also found in lower levels in present day Southern Europeans c 18 5 32 6 Sardinians c 2 4 7 1 and Sicilians c 5 9 11 6 79 70 12 However according to Heyd et al 2023 the specific paternal DNA haplogroup that is most commonly found in male Yamnaya specimens cannot be found in modern Western Europeans or in males from the nearby Corded Ware culture This makes it unlikely that the Corded Ware culture can be directly descended from the Yamnaya culture at least along the paternal line 80 Autosomal tests also indicate that the Yamnaya are the vector for Ancient North Eurasian admixture into Europe 10 Ancient North Eurasian is the name given in literature to a genetic component that represents descent from the people of the Mal ta Buret culture 10 or a population closely related to them That genetic component is visible in tests of the Yamnaya people 10 as well as modern day Europeans 81 Eastern Europe and Finland edit nbsp According to Allentoft 2015 the Sintashta culture probably derived from the Corded Ware Culture In the Baltic Jones et al 2017 found that the Neolithic transition the passage from a hunter gatherer economy to a farming based economy coincided with the arrival en masse of individuals with Yamnaya like ancestry This is different from what happened in Western and Southern Europe where the Neolithic transition was caused by a population that came from Anatolia with Pontic steppe ancestry being detected from only the late Neolithic onward 82 Per Haak et al 2015 the Yamnaya contribution in the modern populations of Eastern Europe ranges from 46 8 among Russians to 42 8 in Ukrainians Finland has the highest Yamnaya contributions in all of Europe 50 4 83 e Central and South Asia edit See also Sintashta culture nbsp Map of the approximate maximal extent of the Andronovo culture The formative Sintashta Petrovka culture is shown in darker red The location of the earliest spoke wheeled chariot finds is indicated in purple Adjacent and overlapping cultures Afanasevo Srubna Bactria Margiana Culture are shown in green nbsp Archaeological cultures associated with Indo Iranian migrations and Indo Aryan migrations after EIEC The Andronovo BMAC and Yaz cultures have often been associated with Indo Iranian migrations The GGC Cemetery H Copper Hoard and PGW cultures are candidates for cultures associated with Indo Aryan migrations Studies also point to the strong presence of Yamnaya descent in the current nations of South Asia especially in groups that are referred to as Indo Aryans 84 85 According to Pathak et al 2018 the North Western Indian amp Pakistani populations PNWI showed significant Middle Late Bronze Age Steppe Steppe MLBA ancestry along with Yamnaya Early Middle Bronze Age Steppe EMBA ancestry but the Indo Europeans of Gangetic Plains and Dravidian people only showed significant Yamnaya Steppe EMBA ancestry and no Steppe MLBA The study also noted that ancient south Asian samples had significantly higher Steppe MLBA than Steppe EMBA or Yamnaya The study identified the Rors as the population in South Asia with the highest proportion of Steppe ancestry 85 Lazaridis et al 2016 estimated 6 5 50 2 steppe related admixture in South Asians though the proportion of Steppe ancestry varies widely across ethnic groups 47 f According to Narasimhan et al 2019 the Yamnaya related ancestry termed Western Steppe EMBA that reached central and south Asia was not the initial expansion from the steppe to the east but a secondary expansion that involved a group possessing 67 Western Steppe EMBA ancestry and 33 ancestry from the European cline This group included people similar to that of Corded Ware Srubnaya Petrovka and Sintashta Moving further east in the central steppe it acquired 9 ancestry from a group of people that possessed West Siberian Hunter Gatherer ancestry thus forming the Central Steppe MLBA cluster which is the primary source of steppe ancestry in South Asia contributing up to 30 of the ancestry of the modern groups in the region 84 According to Unterlander et al 2017 all Iron Age Scythian Steppe nomads can best be described as a mixture of Yamnaya related ancestry and an East Asian related component which most closely corresponds to the modern North Siberian Nganasan people of the lower Yenisey River to varying degrees but generally higher among Eastern Scythians 86 See also editKurgan Kurgan stelae Butmir culture Vinca culture Beaker culture Baden culture Botai culture Khvalynsk culture Mamai Hora Samara culture Sintashta culture Yersinia pestis Proto Indo Europeans Indigenous AryanismNotes edit ˈ j ae m n aɪ e YAM ny ye Russian Yamnaya kultura romanized Yamnaya kultura pronounced ˈjamneje kʊlʲˈture from yama pit hole ˈ j ae m n e YAM ne Ukrainian Yamna kultura romanized Jamna kuľtura Ukrainian pronunciation ˈjamnɐ kʊlʲˈturɐ lit culture of pits Yamnayan cultural aspects for example were horse riding burial styles and to some extent the pastoralist economy The Eastern European hunter gatherers were themselves mostly descended from ancient North Eurasians related to the palaeolithic Mal ta Buret culture Per Haak et al 2015 adding a north Siberian people as a fourth reference population improves residuals for northeastern European populations This accounts for the higher than expected Yamnaya contribution and brings it down to expected levels 67 8 50 4 in Finns 64 9 46 8 in Russians Lazaridis et al 2016 Supplementary Information Table S9 1 Kalash 50 2 Tiwari Brahmins 44 1 Gujarati four samples 46 1 to 27 5 Pathan 44 6 Burusho 42 5 Sindhi 37 7 Punjabi 32 6 Balochi 32 4 Brahui 30 2 Lodhi 29 3 Bengali 24 6 Vishwabhramin 20 4 Makrani 19 2 Mala 18 4 Kusunda 8 9 Kharia 6 5 References edit a b c Allentoft 2015 a b c Morgunova amp Khokhlova 2013 Shishlina Natalia I 11 May 2023 Kristiansen Kristian Kroonen Guus Willerslev Eske eds Yamnaya Pastoralists in the Eurasian Desert Steppe Zone New Perspectives on Mobility The Indo European Puzzle Revisited 1 ed Cambridge University Press pp 34 41 doi 10 1017 9781009261753 006 ISBN 978 1 00 926175 3 retrieved 2023 05 23 a b c Anthony David W 11 May 2023 Kristiansen Kristian Kroonen Guus Willerslev Eske eds The Yamnaya Culture and the Invention of Nomadic Pastoralism in the Eurasian Steppes The Indo European Puzzle Revisited 1 ed Cambridge University Press pp 13 33 doi 10 1017 9781009261753 005 ISBN 978 1 00 926175 3 retrieved 2023 05 13 Novembre 2015 evidence to support theories of a back migration from Corded Ware related populations that contributed to the origins of the Sintashta culture in the Urals and their descendants the Andronovo Hermes Taylor R Tishkin Alexey A Kosintsev Pavel A Stepanova Nadezhda F Krause Kyora Ben Makarewicz Cheryl A 1 December 2020 Mitochondrial DNA of domesticated sheep confirms pastoralist component of Afanasievo subsistence economy in the Altai Mountains 3300 2900 cal BC Archaeological Research in Asia 24 100232 doi 10 1016 j ara 2020 100232 ISSN 2352 2267 S2CID 225136827 a b c Europe s fourth ancestral tribe uncovered BBC 16 November 2015 Wang Chuan Chao Reinhold Sabine Kalmykov Alexey Wissgott Antje Brandt Guido Jeong Choongwon Cheronet Olivia Ferry Matthew Harney Eadaoin Keating Denise Mallick Swapan 4 February 2019 Ancient human genome wide data from a 3000 year interval in the Caucasus corresponds with eco geographic regions Nature Communications 10 1 590 Bibcode 2019NatCo 10 590W doi 10 1038 s41467 018 08220 8 ISSN 2041 1723 PMC 6360191 PMID 30713341 a b Lazaridis Iosif Alpaslan Roodenberg Songul Acar Ayse Acikkol Aysen Agelarakis Anagnostis Aghikyan Levon Akyuz Ugur Andreeva Desislava Andrijasevic Gojko Antonovic Dragana Armit Ian Atmaca Alper Avetisyan Pavel Aytek Ahmet Ihsan Bacvarov Krum 26 August 2022 The genetic history of the Southern Arc A bridge between West Asia and Europe Science 377 6609 eabm4247 doi 10 1126 science abm4247 ISSN 0036 8075 PMC 10064553 PMID 36007055 S2CID 251843620 a b c d e f Haak et al 2015 a b c Mathieson et al 2015 a b Gibbons Ann 10 June 2015 Nomadic herders left a strong genetic mark on Europeans and Asians Science AAAS Nikitin Alexey G Ivanova Svetlana Kiosak Dmytro Badgerow Jessica Pashnick Jeff June 2017 Subdivisions of haplogroups U and C encompass mitochondrial DNA lineages of Eneolithic Early Bronze Age Kurgan populations of western North Pontic steppe Journal of Human Genetics 62 6 605 613 doi 10 1038 jhg 2017 12 ISSN 1435 232X PMID 28148921 S2CID 7459815 Nikitin Alexey G Potekhina Inna Rohland Nadin Mallick Swapan Reich David Lillie Malcolm 24 February 2017 Mitochondrial DNA analysis of eneolithic trypillians from Ukraine reveals neolithic farming genetic roots PLOS ONE 12 2 e0172952 Bibcode 2017PLoSO 1272952N doi 10 1371 journal pone 0172952 ISSN 1932 6203 PMC 5325568 PMID 28235025 Anthony 2007 p 300 370 Nordgvist amp Heyd 2020 Mallory 1999 a b Mallory 1999 p 215 Dolukhanov 1996 p 94 Anthony 2007 p 182 a b Anthony 2007 p 275 Anthony 2007 p 300 Mallory 1999 p 210 211 a b Anthony 2007 p 321 Anthony 2007 pp 274 277 317 320 Anthony 2007 pp 301 302 Anthony 2007 p 303 Parpola 2015 p 49 Parpola 2015 p 45 Parpola 2015 p 47 Telegin D Y 1973 Serednʹo stohivsʹka kulʹtura epokhy midi Kyiv Naukova Dumka p 147 Mallory 1997 p 212 Anthony 2007 p 339 Anthony 2007 p 319 Anthony David The Horse the Wheel and Language pp 328 329 OCLC 1102387902 a b J Harrison Richard The transformation of Europe in the third millennium BC the example of Le Petit Chasseur I II Sion Valais Switzerland p 196 OCLC 718304072 a href Template Cite book html title Template Cite book cite book a CS1 maint multiple names authors list link Anthony 2007 p 305 a b Trautmann Martin 2023 First bioanthropological evidence for Yamnaya horsemanship Science Advances 9 9 eade2451 Bibcode 2023SciA 9E2451T doi 10 1126 sciadv ade2451 hdl 10138 356326 PMID 36867690 S2CID 257334477 P Mallory J 2003 1989 In search of the Indo Europeans language archaeology and myth Thames and Hudson p 213 ISBN 0 500 27616 1 OCLC 886668216 a href Template Cite book html title Template Cite book cite book a CS1 maint multiple names authors list link Gerling Claudia 1 July 2015 Prehistoric Mobility and Diet in the West Eurasian Steppes 3500 to 300 BC An Isotopic Approach De Gruyter doi 10 1515 9783110311211 ISBN 978 3 11 031121 1 Anthony 2007 p 430 J P Mallory Douglas Q Adams eds 1997 Encyclopedia of Indo European culture London Fitzroy Dearborn p 653 ISBN 1 884964 98 2 OCLC 37931209 Librado Pablo 2021 The origins and spread of domestic horses from the Western Eurasian steppes Nature 598 7882 634 640 Bibcode 2021Natur 598 634L doi 10 1038 s41586 021 04018 9 PMC 8550961 PMID 34671162 Hanel Andrea Carlberg Carsten 3 July 2020 Skin colour and vitamin D An update Experimental Dermatology 29 9 864 875 doi 10 1111 exd 14142 PMID 32621306 Wang Chuan Chao Reinhold Sabine Kalmykov Alexey 4 February 2019 Ancient human genome wide data from a 3000 year interval in the Caucasus corresponds with eco geographic regions Nature Communications 10 1 590 Bibcode 2019NatCo 10 590W doi 10 1038 s41467 018 08220 8 ISSN 2041 1723 PMC 6360191 PMID 30713341 Jones et al 2015 a b Lazaridis et al 2016 Lazaridis Iosif Alpaslan Roodenberg Songul Acar Ayse Acikkol Aysen Agelarakis Anagnostis Aghikyan Levon Akyuz Ugur Andreeva Desislava Andrijasevic Gojko Antonovic Dragana Armit Ian Atmaca Alper Avetisyan Pavel Aytek Ahmet Ihsan Bacvarov Krum 26 August 2022 The genetic history of the Southern Arc A bridge between West Asia and Europe Science 377 6609 eabm4247 doi 10 1126 science abm4247 ISSN 0036 8075 PMC 10064553 PMID 36007055 S2CID 251843620 Chintalapati Manjusha Patterson Nick Moorjani Priya 30 May 2022 Perry George H ed The spatiotemporal patterns of major human admixture events during the European Holocene eLife 11 e77625 doi 10 7554 eLife 77625 ISSN 2050 084X PMC 9293011 PMID 35635751 Balanovsky 2017 p 437 Wang 2019 Eske Allentoft Morten E Pokutta Dalia Willerslev 2015 Population genomics of Bronze Age Eurasia University of Copenhagen Denmark OCLC 1234973657 Pilipenko Aleksandr S Trapezov Rostislav O Cherdantsev Stepan V Babenko Vladimir N Nesterova Marina S Pozdnyakov Dmitri V Molodin Vyacheslav I Polosmak Natalia V 20 September 2018 Maternal genetic features of the Iron Age Tagar population from Southern Siberia 1st millennium BC PLOS ONE Public Library of Science PLoS 13 9 e0204062 Bibcode 2018PLoSO 1304062P doi 10 1371 journal pone 0204062 ISSN 1932 6203 PMC 6147448 PMID 30235269 Nikitin Alexey G Ivanova Svetlana Kiosak Dmytro Badgerow Jessica Pashnick Jeff 2 February 2017 Subdivisions of haplogroups U and C encompass mitochondrial DNA lineages of Eneolithic Early Bronze Age Kurgan populations of western North Pontic steppe Journal of Human Genetics Springer Science and Business Media LLC 62 6 605 613 doi 10 1038 jhg 2017 12 ISSN 1434 5161 PMID 28148921 S2CID 7459815 In the 12 successfully haplotyped specimens 75 of mtDNA lineages consisted of west Eurasian haplogroup U and its U4 and U5 sublineages Furthermore we identified a subgroup of east Eurasian haplogroup C in two representatives of the Yamna culture in one of the studied kurgans Gibbons A 24 July 2015 Revolution in human evolution Science 349 6246 362 366 Bibcode 2015Sci 349 362G doi 10 1126 science 349 6246 362 ISSN 0036 8075 PMID 26206910 Hanel Andrea Carlberg Carsten September 2020 Skin colour and vitamin D An update Experimental Dermatology 29 9 864 875 doi 10 1111 exd 14142 ISSN 0906 6705 PMID 32621306 Mathieson Iain Alpaslan Roodenberg Songul Posth Cosimo Szecsenyi Nagy Anna Rohland Nadin Mallick Swapan Olalde Inigo Broomandkhoshbacht Nasreen Candilio Francesca Cheronet Olivia Fernandes Daniel March 2018 The genomic history of southeastern Europe Nature 555 7695 197 203 Bibcode 2018Natur 555 197M doi 10 1038 nature25778 ISSN 1476 4687 PMC 6091220 PMID 29466330 Heyd Volker April 2017 Kossinna s smile Antiquity 91 356 348 359 doi 10 15184 aqy 2017 21 hdl 10138 255652 ISSN 0003 598X S2CID 164376362 Mathieson Iain Lazaridis Iosif Rohland Nadin Mallick Swapan Patterson Nick Roodenberg Songul Alpaslan Harney Eadaoin Stewardson Kristin Fernandes Daniel Novak Mario Sirak Kendra 24 December 2015 Genome wide patterns of selection in 230 ancient Eurasians Nature 528 7583 499 503 Bibcode 2015Natur 528 499M doi 10 1038 nature16152 ISSN 0028 0836 PMC 4918750 PMID 26595274 Segurel Laure 2020 Why and when was lactase persistence selected for Insights from Central Asian herders and ancient DNA PLOS Biology 18 6 e3000742 doi 10 1371 journal pbio 3000742 PMC 7302802 PMID 32511234 Furthermore ancient DNA studies found that the LP mutation was absent or very rare in Europe until the end of the Bronze Age 26 29 and appeared first in individuals with steppe ancestry 19 20 Thus it was proposed that the mutation originated in Yamnaya associated populations and arrived later in Europe by migration of these steppe herders Callaway Ewen DNA data explosion lights up the Bronze Age Nature the 101 sequenced individuals the Yamnaya were most likely to have the DNA variation responsible for lactose tolerance hinting that the steppe migrants might have eventually introduced the trait to Europe Furholt Martin 2018 Massive Migrations The Impact of Recent DNA Studies on our View of Third Millennium Europe European Journal of Archaeology 21 2 159 191 doi 10 1017 eaa 2017 43 For example one lineage could have a biological evolutionary advantage over the other Allentoft et al 2015 171 have found a remarkably high rate of lactose tolerance among individuals connected to Yamnaya and to Corded Ware as opposed to the majority of Late Neolithic individuals Saag L 2020 Human Genetics Lactase Persistence in a Battlefield Current Biology 30 21 R1311 R1313 doi 10 1016 j cub 2020 08 087 PMID 33142099 S2CID 226229587 Lazaridis et al 2022c Reich David 2018 Who we are and how we got here ancient DNA and the new science of the human past Oxford United Kingdom p 121 ISBN 978 0 19 882125 0 OCLC 1006478846 a href Template Cite book html title Template Cite book cite book a CS1 maint location missing publisher link Reich David 2018 Who we are and how we got here ancient DNA and the new science of the human past Oxford United Kingdom p 239 ISBN 978 0 19 882125 0 OCLC 1006478846 a href Template Cite book html title Template Cite book cite book a CS1 maint location missing publisher link Klejn Leo 2017 The Steppe Hypothesis of Indo European Origins Remains to be Proven Acta Archaeologica 88 1 193 204 doi 10 1111 j 1600 0390 2017 12184 x Balanovsky O 2017 Genetic differentiation between upland and lowland populations shapes the Y chromosomal landscape of West Asia Human Genetics 136 4 437 450 doi 10 1007 s00439 017 1770 2 PMID 28281087 S2CID 3735168 The ancient Yamnaya samples are located on the eastern R GG400 branch of haplogroup R1b L23 showing that the paternal descendants of the Yamnaya still live in the Pontic steppe and that the ancient Yamnaya population was not an important source of paternal lineages in present day West Europeans Klejn 2017 p 201 In the tables presented in the article by Reichs team Haak et al 2015 the genetic pool connecting the Yamnaya culture with the Corded Ware people is shown to be more intense in Northern Europe Norway and Sweden and decreases gradually from the North to the South Fig 6 It is weakest around the Danube in Hungary i e areas neighbouring the western branch of the Yamnaya culture This is the reverse image to what the proposed hypothesis by the geneticists would lead us to expect It is true that this gradient is traced back from the contemporary materials but it was already present during the Bronze Age a b Zimmer Carl 10 June 2015 DNA Deciphers Roots of Modern Europeans New York Times Retrieved 2020 12 12 Olsen Birgit A Olander Thomas Kristiansen Kristian 23 August 2019 Tracing the Indo Europeans Oxbow Books pp 1 6 doi 10 2307 j ctvmx3k2h 6 ISBN 978 1 78925 273 6 S2CID 202354040 Olsen Birgit A 11 May 2023 Kristiansen Kristian Kroonen Guus Willerslev Eske eds Marriage Strategies and Fosterage among the Indo Europeans A Linguistic Perspective The Indo European Puzzle Revisited 1 ed Cambridge University Press pp 296 302 doi 10 1017 9781009261753 027 ISBN 978 1 00 926175 3 retrieved 2023 05 13 Schneider Thomas 2023 Language Contact in Ancient Egypt LIT Verlag Munster p 110 113 ISBN 978 3 643 91507 8 Anthony 2019b p 36 Anthony 2019a p 1 19 Nikitin Alexey Ivanova Svetlana 28 November 2022 Long distance exchanges along the Black Sea coast in the Eneolithic and the steppe genetic ancestry problem Cambridge Open Engage doi 10 33774 coe 2022 7m315 Kroonen Guus Jakob Anthony Palmer Axel I van Sluis Paulus Wigman Andrew 12 October 2022 Indo European cereal terminology suggests a Northwest Pontic homeland for the core Indo European languages PLOS ONE 17 10 e0275744 Bibcode 2022PLoSO 1775744K doi 10 1371 journal pone 0275744 ISSN 1932 6203 PMC 9555676 PMID 36223379 Stoneking Mark Arias Leonardo Liu Dang Oliveira Sandra Pugach Irina Rodriguez Jae Joseph Russell B 24 January 2023 Genomic perspectives on human dispersals during the Holocene Proceedings of the National Academy of Sciences 120 4 e2209475119 Bibcode 2023PNAS 12009475S doi 10 1073 pnas 2209475119 ISSN 0027 8424 PMC 9942792 PMID 36649433 Haak et al 2015 pp 121 124 Kristiansen Kristian Kroonen Guus Willerslev Eske 11 May 2023 The Indo European Puzzle Revisited Integrating Archaeology Genetics and Linguistics Cambridge University Press p 70 76 ISBN 978 1 009 26174 6 Lazaridis et al 2014 Jones et al 2017 Haak et al 2015 pp 121 122 a b Narasimhan et al 2019 a b Pathak et al 2018 Unterlander et al 2017Genomic inference reveals that Scythians in the east and the west of the steppe zone can best be described as a mixture of Yamnaya related ancestry and an East Asian component Demographic modelling suggests independent origins for eastern and western groups with ongoing gene flow between them plausibly explaining the striking uniformity of their material culture We also find evidence that significant gene flow from east to west Eurasia must have occurred early during the Iron Age and The blend of EHG European hunter gatherer and Caucasian elements in carriers of the Yamnaya culture was formed on the European steppe and exported into Central Asia and Siberia We therefore considered an alternative model in which we treat them as a mix of Yamnaya and the Han Supplementary Table 25 This model fits all of the Iron Age Scythian groups consistent with these groups having ancestry related to East Asians not found in the other populations Alternatively the Iron Age Scythian groups can also be modelled as a mix of Yamnaya and the north Siberian Nganasan Supplementary Note 2 Supplementary Table 26 Sources edit Allentoft Morten E et al 2015 Population genomics of Bronze Age Eurasia Nature 522 7555 167 172 Bibcode 2015Natur 522 167A doi 10 1038 nature14507 PMID 26062507 S2CID 4399103 Anthony David W 2007 The Horse the Wheel and Language How Bronze Age Riders from the Eurasian Steppes Shaped the Modern World Princeton University Press ISBN 978 0 691 05887 0 Anthony David 2017 Archaeology and Language Why Archaeologists Care About the Indo European Problem In Crabtree P J Bogucki P eds European Archaeology as Anthropology Essays in Memory of Bernard Wailes Anthony David Spring Summer 2019a Archaeology Genetics and Language in the Steppes A Comment on Bomhard Journal of Indo European Studies 47 1 2 Retrieved 2020 01 09 Anthony David W 2019b Ancient DNA Mating Networks and the Anatolian Split In Serangeli Matilde Olander Thomas eds Dispersals and Diversification Linguistic and Archaeological Perspectives on the Early Stages of Indo European BRILL pp 21 54 ISBN 978 9004416192 Cassidy LM Martiniano R Murphy EM Teasdale MD Mallory J Hartwell B Bradley DG 2016 Neolithic and Bronze Age migration to Ireland and establishment of the insular Atlantic genome PNAS 113 2 368 373 Bibcode 2016PNAS 113 368C doi 10 1073 pnas 1518445113 PMC 4720318 PMID 26712024 Dolukhanov Pavel M 1996 The Early Slavs Eastern Europe from the Initial Settlement to the Kievan Rus New York Longman ISBN 0 582 23627 4 Fortson Benjamin W 2004 Indo European Language and Culture An Introduction Blackwell Publishing Gallego Llorente M Connell S Jones ER Merrett DC Jeon Y Eriksson A et al 2016 The genetics of an early Neolithic pastoralist from the Zagros Iran Scientific Reports 6 31326 Bibcode 2016NatSR 631326G doi 10 1038 srep31326 PMC 4977546 PMID 27502179 Haak W Lazaridis I Patterson N Rohland N Mallick S Llamas B et al 2015 Massive migration from the steppe was a source for Indo European languages in Europe Nature 522 7555 207 211 arXiv 1502 02783 Bibcode 2015Natur 522 207H bioRxiv 10 1101 013433 doi 10 1038 nature14317 PMC 5048219 PMID 25731166 Jeong Choongwon Balanovsky Oleg Lukianova Elena Kahbatkyzy Nurzhibek et al 29 April 2019 The genetic history of admixture across inner Eurasia languages in Europe Nature Ecology and Evolution Nature Research 3 6 966 976 doi 10 1038 s41559 019 0878 2 hdl 10871 36562 PMC 6542712 PMID 31036896 Jones Eppie R Gonzalez Fortes Gloria Connell Sarah Siska Veronika et al 2015 Upper Palaeolithic genomes reveal deep roots of modern Eurasians Nature Communications 6 8912 Bibcode 2015NatCo 6 8912J doi 10 1038 ncomms9912 PMC 4660371 PMID 26567969 Jones ER Zarina G Moiseyev V Lightfoot E Nigst PR Manica A Pinhasi R Bradley DG 2017 The Neolithic transition in the Baltic was not driven by admixture with early European farmers Current Biology 27 4 576 582 doi 10 1016 j cub 2016 12 060 PMC 5321670 PMID 28162894 Kuzmina Elena E 2007 Mallory J P ed The Origin of the Indo Iranians BRILL ISBN 978 9004160545 Lazaridis Iosif Patterson Nick Mittnik Alissa Renaud Gabriel et al 2014 Ancient human genomes suggest three ancestral populations for present day Europeans Nature 513 7518 409 413 arXiv 1312 6639 Bibcode 2014Natur 513 409L doi 10 1038 nature13673 PMC 4170574 PMID 25230663 Lazaridis I Nadel D Rollefson G Merrett DC Rohland N Mallick S et al 16 June 2016 The genetic structure of the world s first farmers bioRxiv 10 1101 059311 Supplementary Information Lazaridis I Nadel D Rollefson G Merrett DC Rohland N Mallick S et al 25 July 2016 Genomic insights into the origin of farming in the ancient Near East Nature published August 2016 536 7617 419 424 Bibcode 2016Natur 536 419L doi 10 1038 nature19310 PMC 5003663 PMID 27459054 Lazaridis Iosif Alpaslan Roodenberg Songul Acar Ayse Acikkol Aysen Agelarakis Anagnostis Aghikyan Levon Akyuz Ugur Andreeva Desislava Andrijasevic Gojko Antonovic Dragana Armit Ian Atmaca Alper Avetisyan Pavel Aytek Ahmet Ihsan Bacvarov Krum 26 August 2022 A genetic probe into the ancient and medieval history of Southern Europe and West Asia Science 377 6609 940 951 Bibcode 2022Sci 377 940L doi 10 1126 science abq0755 ISSN 0036 8075 PMC 10019558 PMID 36007020 S2CID 251844202 Mallory J P 1997 Yamna Culture Encyclopedia of Indo European Culture Fitzroy Dearborn Mallory J P 1999 In Search of the Indo Europeans Language Archaeology and Myth reprint ed London Thames amp Hudson ISBN 978 0 500 27616 7 Martiniano R et al 2017 The population genomics of archaeological transition in west Iberia Investigation of ancient substructure using imputation and haplotype based methods PLOS Genet 13 7 e1006852 doi 10 1371 journal pgen 1006852 PMC 5531429 PMID 28749934 Mathieson Iain et al 10 October 2015 Eight thousand years of natural selection in Europe bioRxiv 10 1101 016477 Mathieson Iain et al 23 November 2015 Genome wide patterns of selection in 230 ancient Eurasians Nature published 24 December 2015 528 7583 499 503 Bibcode 2015Natur 528 499M doi 10 1038 nature16152 PMC 4918750 PMID 26595274 Mathieson Iain et al 21 February 2018 The Genomic History of Southeastern Europe Nature Nature Research 555 7695 197 203 Bibcode 2018Natur 555 197M doi 10 1038 nature25778 PMC 6091220 PMID 29466330 Morgunova Nina Khokhlova Olga 2013 Chronology and Periodization of the Pit Grave Culture in the Area Between the Volga and Ural Rivers Based on 14C Dating and Paleopedological Research Radiocarbon 55 2 3 1286 1296 doi 10 2458 azu js rc 55 16087 ISSN 0033 8222 Narasimhan VM Patterson N Moorjani P Rohland N et al 2019 The formation of human populations in South and Central Asia Science 365 6457 eaat7487 doi 10 1126 science aat7487 PMC 6822619 PMID 31488661 Nordgvist Kerkko Heyd Volker 2020 The Forgotten Child of the Wider Corded Ware Family Russian Fatyanovo Culture in Context Proceedings of the Prehistoric Society 86 65 93 doi 10 1017 ppr 2020 9 S2CID 228923806 Novembre John 11 June 2015 Ancient DNA steps into the language debate Nature 522 7555 164 165 doi 10 1038 522164a PMID 26062506 S2CID 205085294 Parpola Asko 2015 The Roots of Hinduism Oxford University Press Pashnick Jeff August 2014 Genetic Analysis of Ancient Human Remains from the Early Bronze Age Cultures of the North PonticSteppe Region Masters Theses thesis Vol 737 Grand Valley State University Retrieved 2020 01 12 Pathak AK Kadian A Kushniarevich A Montinaro F Mondal M Ongaro L et al 6 December 2018 The Genetic Ancestry of Modern Indus Valley Populations from Northwest India The American Journal of Human Genetics 103 6 918 929 doi 10 1016 j ajhg 2018 10 022 PMC 6288199 PMID 30526867 Unterlander Martina Palstra Friso Lazaridis Iosif Pilipenko Aleksandr Hofmanova Zuzana Gross Melanie et al 2017 Ancestry and demography and descendants of Iron Age nomads of the Eurasian Steppe Nature Communications 8 14615 Bibcode 2017NatCo 814615U doi 10 1038 ncomms14615 ISSN 2041 1723 PMC 5337992 PMID 28256537 Wang Chuan Chao 4 February 2019 Ancient human genome wide data from a 3000 year interval in the Caucasus corresponds with eco geographic regions Eurasia Nature Communications Nature Research 10 1 590 Bibcode 2019NatCo 10 590W bioRxiv 10 1101 322347 doi 10 1038 s41467 018 08220 8 PMC 6360191 PMID 30713341 Wilde S Timpson A Kirsanow K Kaiser E Kayser M Unterlander M et al 2014 Direct evidence for positive selection of skin hair and eye pigmentation in Europeans during the last 5 000 y Proceedings of the National Academy of Sciences of the United States of America 111 13 4832 4837 Bibcode 2014PNAS 111 4832W doi 10 1073 pnas 1316513111 PMC 3977302 PMID 24616518 External links edit nbsp Wikimedia Commons has media related to Yamna culture Genetic study revives debate on origin and expansion of Indo European Languages Science Daily March 2015 First Horse Warriors PBS org NOVA series 15 May 2019 Retrieved from https en wikipedia org w index php title Yamnaya culture amp oldid 1206647116, wikipedia, wiki, book, books, library,

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