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Wikipedia

Lilium

Lilium (/ˈlɪliəm/ LIL-ee-əm)[3] is a genus of herbaceous flowering plants growing from bulbs, all with large prominent flowers. They are the true lilies. Lilies are a group of flowering plants which are important in culture and literature in much of the world. Most species are native to the northern hemisphere and their range is temperate climates and extends into the subtropics. Many other plants have "lily" in their common names, but do not belong to the same genus and are therefore not true lilies.

Lilium
Lilium candidum
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Order: Liliales
Family: Liliaceae
Subfamily: Lilioideae
Tribe: Lilieae
Genus: Lilium
L.[1]
Type species
Lilium candidum
Species

List of Lilium species

Synonyms[1]
  • Lirium Scop., nom. illeg.
  • Martagon Wolf
  • Martagon (Rchb.) Opiz, nom. illeg.
  • Nomocharis Franch.

Description

 
Lilium longiflorum flower – 1. stigma, 2. style, 3. anthers, 4. filament, 5. tepal

Lilies are tall perennials ranging in height from 2–6 ft (60–180 cm). They form naked or tunicless scaly underground bulbs which are their organs of perennation. In some North American species the base of the bulb develops into rhizomes, on which numerous small bulbs are found. Some species develop stolons.[4] Most bulbs are buried deep in the ground, but a few species form bulbs near the soil surface. Many species form stem-roots. With these, the bulb grows naturally at some depth in the soil, and each year the new stem puts out adventitious roots above the bulb as it emerges from the soil. These roots are in addition to the basal roots that develop at the base of the bulb, a number of species also produce contractile roots that move the bulbs deeper into the soil.[5]

 
Lily, petal

The flowers are large, often fragrant, and come in a wide range of colors including whites, yellows, oranges, pinks, reds and purples. Markings include spots and brush strokes. The plants are late spring- or summer-flowering. Flowers are borne in racemes or umbels at the tip of the stem, with six tepals spreading or reflexed, to give flowers varying from funnel shape to a "Turk's cap". The tepals are free from each other, and bear a nectary at the base of each flower. The ovary is 'superior', borne above the point of attachment of the anthers. The fruit is a three-celled capsule.[6]

 
Stamen of lilium

Seeds ripen in late summer. They exhibit varying and sometimes complex germination patterns, many adapted to cool temperate climates.

Most cool temperate species are deciduous and dormant in winter in their native environment. But a few species native to areas with hot summers and mild winters (Lilium candidum, Lilium catesbaei, Lilium longiflorum) lose their leaves and enter a short dormant period in summer or autumn, sprout from autumn to winter, forming dwarf stems bearing a basal rosette of leaves until, after they have received sufficient chilling, the stem begins to elongate in warming weather.

 
Lilium candidum seeds

The basic chromosome number is twelve (n=12).[7]

Taxonomy

Taxonomical division in sections follows the classical division of Comber,[8] species acceptance follows the World Checklist of Selected Plant Families,[9] the taxonomy of section Pseudolirium is from the Flora of North America,[10] the taxonomy of Section Liriotypus is given in consideration of Resetnik et al. 2007,[11] the taxonomy of Chinese species (various sections) follows the Flora of China[12] and the taxonomy of Section Sinomartagon follows Nishikawa et al.[13] as does the taxonomy of Section Archelirion.[14]

The World Checklist of Selected Plant Families, as of January 2014, considers Nomocharis a separate genus in its own right,[15] however some authorities consider Nomocharis to be embedded within Lilium, rather than treat it as a separate genus.[16][17] The Sinomartagon are divided in 3 paraphyletic groups, while the Leucolirion are divided in 2 paraphyletic groups.[18]

There are seven sections:

  • Martagon
  • Pseudolirium
  • Liriotypus
  • Archelirion
  • Sinomartagon
  • Leucolirion
  • Daurolirion

There are 111 species counted in this genus.[19] For a full list of accepted species[1] with their native ranges, see List of Lilium species.

Some species formerly included within this genus have now been placed in other genera. These genera include Cardiocrinum, Notholirion, Nomocharis and Fritillaria.

Etymology

The botanic name Lilium is the Latin form and is a Linnaean name. The Latin name is derived from the Greek word λείριον leírion, generally assumed to refer to true, white lilies as exemplified by the Madonna lily.[23][24][25] The word was borrowed from Coptic (dial. Fayyumic) hleri, from standard hreri, from Demotic hrry, from Egyptian hrṛt "flower".[citation needed] Meillet maintains that both the Egyptian and the Greek word are possible loans from an extinct, substratum language of the Eastern Mediterranean.[citation needed] Ancient Greek: κρῖνον, krīnon, was used by the Greeks, albeit for lilies of any color.[26]

The term "lily" has in the past been applied to numerous flowering plants, often with only superficial resemblance to the true lily, including water lily, fire lily, lily of the Nile, calla lily, trout lily, kaffir lily, cobra lily, lily of the valley, daylily, ginger lily, Amazon lily, leek lily, Peruvian lily, and others. All English translations of the Bible render the Hebrew shūshan, shōshan, shōshannā as "lily", but the "lily among the thorns" of Song of Solomon, for instance, may be the honeysuckle.[27]

Distribution and habitat

The range of lilies in the Old World extends across much of Europe, across most of Asia to Japan, south to India, and east to Indochina and the Philippines. In the New World they extend from southern Canada through much of the United States. They are commonly adapted to either woodland habitats, often montane, or sometimes to grassland habitats. A few can survive in marshland and epiphytes are known in tropical southeast Asia. In general they prefer moderately acidic or lime-free soils.

Ecology

Lilies are used as food plants by the larvae of some Lepidoptera species including the Dun-bar.

The proliferation of deer (e.g. Odocoileus virginianus) in North America, mainly due to factors such as the elimination of large predators for human safety, is responsible there for a downturn in lily populations in the wild and is a threat to garden lilies as well.[28] Fences as high as 8 feet may be required to prevent them from consuming the plants, an impractical solution for most wild areas.[29]

Cultivation

Many species are widely grown in the garden in temperate, sub-tropical and tropical regions.[30] Numerous ornamental hybrids have been developed. They are used in herbaceous borders, woodland and shrub plantings, and as patio plants. Some lilies, especially Lilium longiflorum, form important cut flower crops or potted plants. These are forced to flower outside of the normal flowering season for particular markets; for instance, Lilium longiflorum for the Easter trade, when it may be called the Easter lily.

Lilies are usually planted as bulbs in the dormant season. They are best planted in a south-facing (northern hemisphere), slightly sloping aspect, in sun or part shade, at a depth 2½ times the height of the bulb (except Lilium candidum which should be planted at the surface). Most prefer a porous, loamy soil, and good drainage is essential. Most species bloom in July or August (northern hemisphere). The flowering periods of certain lily species begin in late spring, while others bloom in late summer or early autumn.[31] They have contractile roots which pull the plant down to the correct depth, therefore it is better to plant them too shallowly than too deep. A soil pH of around 6.5 is generally safe. Most grow best in well-drained soils, and plants are watered during the growing season. Some species and cultivars have strong wiry stems, but those with heavy flower heads are staked to stay upright.[32][33]

Awards

The following lily species and cultivars currently hold the Royal Horticultural Society's Award of Garden Merit (confirmed 2017):[34]

 
'Golden Splendor'

Classification of garden forms

Numerous forms, mostly hybrids, are grown for the garden. They vary according to the species and interspecific hybrids that they derived from, and are classified in the following broad groups:[40][41][42]

Asiatic hybrids (Division I)

These are derived from hybrids between species in Lilium section Sinomartagon.[43][44]
They are derived from central and East Asian species and interspecific hybrids, including Lilium amabile, Lilium bulbiferum, Lilium callosum, Lilium cernuum, Lilium concolor, Lilium dauricum, Lilium davidii, Lilium × hollandicum, Lilium lancifolium (syn. Lilium tigrinum), Lilium lankongense, Lilium leichtlinii, Lilium × maculatum, Lilium pumilum, Lilium × scottiae, Lilium wardii and Lilium wilsonii.
These are plants with medium-sized, upright or outward facing flowers, mostly unscented. There are various cultivars such as Lilium 'Cappuccino', Lilium 'Dimension', Lilium 'Little Kiss' and Lilium 'Navona'.[45]
  • Dwarf (Patio, Border) varieties are much shorter, c.36–61 cm in height and were designed for containers.[46] They often bear the cultivar name 'Tiny', such as the 'Lily Looks' series, e.g. 'Tiny Padhye',[47] 'Tiny Dessert'.[48]

Martagon hybrids (Division II)

These are based on Lilium dalhansonii, Lilium hansonii, Lilium martagon, Lilium medeoloides, and Lilium tsingtauense.
The flowers are nodding, Turk's cap style (with the petals strongly recurved).

Candidum (Euro-Caucasian) hybrids (Division III)

This includes mostly European species: Lilium candidum, Lilium chalcedonicum, Lilium kesselringianum, Lilium monadelphum, Lilium pomponium, Lilium pyrenaicum and Lilium × testaceum.

American hybrids (Division IV)

These are mostly taller growing forms, originally derived from Lilium bolanderi, Lilium × burbankii, Lilium canadense, Lilium columbianum, Lilium grayi, Lilium humboldtii, Lilium kelleyanum, Lilium kelloggii, Lilium maritimum, Lilium michauxii, Lilium michiganense, Lilium occidentale, Lilium × pardaboldtii, Lilium pardalinum, Lilium parryi, Lilium parvum, Lilium philadelphicum, Lilium pitkinense, Lilium superbum, Lilium ollmeri, Lilium washingtonianum, and Lilium wigginsii.
Many are clump-forming perennials with rhizomatous rootstocks.

Longiflorum hybrids (Division V)

These are cultivated forms of this species and its subspecies.
They are most important as plants for cut flowers, and are less often grown in the garden than other hybrids.

Trumpet lilies (Division VI), including Aurelian hybrids (with L. henryi)

This group includes hybrids of many Asiatic species and their interspecific hybrids, including Lilium × aurelianense, Lilium brownii, Lilium × centigale, Lilium henryi, Lilium × imperiale, Lilium × kewense, Lilium leucanthum, Lilium regale, Lilium rosthornii, Lilium sargentiae, Lilium sulphureum and Lilium × sulphurgale.
The flowers are trumpet shaped, facing outward or somewhat downward, and tend to be strongly fragrant, often especially night-fragrant.

Oriental hybrids (Division VII)

These are based on hybrids within Lilium section Archelirion,[43][44] specifically Lilium auratum and Lilium speciosum, together with crossbreeds from several species native to Japan, including Lilium nobilissimum, Lilium rubellum, Lilium alexandrae, and Lilium japonicum.
They are fragrant, and the flowers tend to be outward facing. Plants tend to be tall, and the flowers may be quite large. The whole group are sometimes referred to as "stargazers" because many of them appear to look upwards. (For the specific cultivar, see Lilium 'Stargazer'.)

Other hybrids (Division VIII)

Includes all other garden hybrids.

Species (Division IX)

All natural species and naturally occurring forms are included in this group.

The flowers can be classified by flower aspect and form:[49]

  • Flower aspect:
  • a up-facing
  • b out-facing
  • c down-facing
  • Flower form:
  • a trumpet-shaped
  • b bowl-shaped
  • c flat (or with tepal tips recurved)
  • d tepals strongly recurved (with the Turk's cap form as the ultimate state)

Many newer commercial varieties are developed by using new technologies such as ovary culture and embryo rescue.[50]

Pests and diseases

 
Scarlet lily beetles, Oxfordshire, UK

Aphids may infest plants. Leatherjackets feed on the roots. Larvae of the Scarlet lily beetle can cause serious damage to the stems and leaves. The scarlet beetle lays its eggs and completes its life cycle only on true lilies (Lilium) and fritillaries (Fritillaria).[51] Oriental, rubrum, tiger and trumpet lilies as well as Oriental trumpets (orienpets) and Turk's cap lilies and native North American Lilium species are all vulnerable, but the beetle prefers some types over others. The beetle could also be having an effect on native Canadian species and some rare and endangered species found in northeastern North America.[52] Daylilies (Hemerocallis, not true lilies) are excluded from this category. Plants can suffer from damage caused by mice, deer and squirrels. Slugs,[53] snails and millipedes attack seedlings, leaves and flowers.

Brown spots on damp leaves may signal an infection of Botrytis elliptica, also known as Lily blight, lily fire, and botrytis leaf blight.[54] Various viral diseases can cause mottling of leaves and stunting of growth, including lily curl stripe, ringspot, and lily rosette virus.[55]

Propagation and growth

Lilies can be propagated in several ways;

  • by division of the bulbs
  • by growing-on bulbils which are adventitious bulbs formed on the stem
  • by scaling, for which whole scales are detached from the bulb and planted to form a new bulb
  • by seed; there are many seed germination patterns, which can be complex
  • by micropropagation techniques (which include tissue culture);[56] commercial quantities of lilies are often propagated in vitro and then planted out to grow into plants large enough to sell. A highly efficient technique for multiple shoot and propagule formation was given by Yadav et al., in 2013.[57]

Plant grow regulators (PGRs) are used to limit the height of lilies, especial those sold as potted plants; commonly used chemicals include ancymidol, fluprimidol, paclobutrazol, and uni-conazole, which are applied to the foliage and retard the biosynthesis of gibberellins, a class of plant hormones responsible for stem growth.[58]

Research

A comparison of meiotic crossing-over (recombination) in lily and mouse led, in 1977, to the conclusion that diverse eukaryotes share a common pattern of meiotic crossing-over.[59] Lilium longiflorum has been used for studying aspects of the basic molecular mechanism of genetic recombination during meiosis.[60][61]

Toxicity

Some Lilium species are toxic to cats. This is known to be so especially for Lilium longiflorum, though other Lilium and the unrelated Hemerocallis can also cause the same symptoms.[62][63][64][65] The true mechanism of toxicity is undetermined, but it involves damage to the renal tubular epithelium (composing the substance of the kidney and secreting, collecting, and conducting urine), which can cause acute kidney failure.[65] Veterinary help should be sought, as a matter of urgency, for any cat that is suspected of eating any part of a lily – including licking pollen that may have brushed onto its coat.[66]

Culinary uses

Chinese cuisine

Lily bulbs are starchy and edible as root vegetables, though bulbs of some species may be too bitter to eat.[67]

Lilium brownii var. viridulum, known as 百合 (pak hop; pinyin: bǎi hé; Cantonese Yale: baak hap; lit. 'hundred united'), is one of the most prominent edible lilies in China. Its bulbs are large in size and not bitter. They were even exported and sold in the San Francisco Chinatown in the 19th century, available both fresh and dry.[67] A landrace called 龍牙百合 (pinyin: lóng yá bǎi hé; lit. 'dragon-tooth lily') mainly cultivated in Hunan and Jiangxi is especially renowned for its good-quality bulbs.[68]

L. lancifolium (Chinese: 卷丹; pinyin: juǎn dān; lit. 'reflexed red') is widely cultivated in China, especially in Yixing, Huzhou and Longshan. Its bulbs are slightly bitter.[68]

L. davidii var. unicolor (Chinese: 蘭州百合; lit. 'Lanzhou lily') is mainly cultivated in Lanzhou and its bulbs are valued for sweetness.[68]

Other edible Chinese lilies include L. brownii var. brownii, L. davidii var. davidii, L. concolor, L. pensylvanicum, L. distichum, L. martagon var. pilosiusculum, L. pumilum, L. rosthornii and L. speciosum var. gloriosoides.[69] Researchers have also explored the possibility of using ornamental cultivars as edible lilies.[72]

The dried bulbs are commonly used in the south to flavor soup.[citation needed] They may be reconstituted and stir-fried, grated and used to thicken soup, or processed to extract starch.[citation needed] Their texture and taste draw comparisons with the potato, although the individual bulb scales are much smaller.[citation needed]

The commonly marketed "lily" flower buds, called 金针菜 (kam cham tsoi;[73] pinyin: jīn zhēn cài; Cantonese Yale: gāmjām choi; lit. 'gold needle vegetable') in Chinese cuisine, are actually from daylilies, Hemerocallis citrina,[74] or possibly H. fulva.[a][73] Flowers of the H. graminea and Lilium bulbiferum were reported to have been eaten as well, but samples provided by the informant were strictly daylilies and did not include L. bulbiferum.[b][75]

Lily flowers and bulbs are eaten especially in the summer, for their perceived ability to reduce internal heat.[76] A 19th century English source reported that "Lily flowers are also said to be efficacious in pulmonary affections, and to have tonic properties".[75]

Asiatic lily cultivars are also imported from the Netherlands; the seedling bulbs must be imported from the Netherlands every year.[77][78][79]

The parts of Lilium species which are officially listed as food material in Taiwan are the flower and bulbs of Lilium lancifolium, Lilium brownii var. viridulum, Lilium pumilum and Lilium candidum.[80]

Japanese cuisine

 
Loose scales of lily bulb in a donburi bowl dish

The lily bulb or yuri-ne is sometimes used in Japanese cuisine.[c][81] It may be most familiar in the present day as an occasional ingredient (, gu) in the chawan-mushi (savoury egg custard),[82] where a few loosened scales of this optional ingredient are found embedded in the "hot pudding" of each serving.[83][84] It could also be used as an ingredient in a clear soup or suimono [ja].[85][86]

The boiled bulb may also be strained[d] into purée for use, as in the sweetened kinton,[87][88] or chakin-shibori.[88][89][e]

Yokan

There is also the yuri-yōkan, one recipe of which calls for combining measures of yuri starch with agar dissolved in water and sugar.[91] This was a specialty of Hamada, Shimane,[92] and the shop Kaisei-dō (開盛堂) established in 1885 became famous for it.[93][94] Because a certain Viscount Jimyōin wrote a waka poem about the confection which mentioned hime-yuri "princess lily",[f] one source stated that the hime-yuri (usually taken to mean L. concolor) had to have been used,[93] but another source points out that the city of Hamada lies back to back with across a mountain range with Fuchu, Hiroshima which is renowned for its production of yama-yuri (L. auratum).[90][g]

Species used

Current Japanese governmental sources (c. 2005) list the following lily species as prominent in domestic consumption:[98][99] the oni yuri or tiger lily Lilium lancifolium, the kooni yuri Lilium leichtlinii var. maximowiczii,[h] and the gold-banded white yama-yuri L. auratum.

But Japanese sources c. 1895–1900,[95][100] give a top-three list which replaces kooni yuri with the sukashi-yuri (透かし百合, lit. "see-through lily", L. maculatum) named from the gaps between the tepals.[101][102]

There is uncertainty regarding which species is meant by the hime-yuri used as food, because although this is usually the common name for L. concolor in most up-to-date literature,[103] it used to ambiguously referred to the tiger lily as well, c. 1895–1900.[95] The non-tiger-lily himeyuri is certainly described as quite palatable in the literature at the time, but the extent of exploitation could not have been as significant.[i]

North America

The flower buds and roots of Lilium canadense are traditionally gathered and eaten by North American indigenous peoples.[104] Coast Salish, Nuu-chah-nulth and most western Washington peoples steam, boil or pit-cook the bulbs of Lilium columbianum. Bitter or peppery-tasting, they were mostly used as a flavoring, often in soup with meat or fish.[105]

Medicinal uses

Traditional Chinese medicine list the use of the following: 野百合 Lilium brownii, 百合 Lilium brownii var. viridulum, 渥丹 Lilium concolor, 毛百合 Lilium dauricum, 卷丹 Lilium lancifolium, 山丹 Lilium pumilum, 南川百合 Lilium rosthornii, 药百合Lilium speciosum var. gloriosoides, 淡黄花百合 Lilium sulphureum.[106][107]

In Taiwan, governmental publications list Lilium lancifolium Thunb., Lilium brownii var. viridulum Baker, Lilium pumilum DC.[108]

In the kanpō or Chinese medicine as practiced in Japan, the official Japanese governmental pharmacopeia Nihon yakkyokuhō (日本薬局方) includes the use of lily bulb (known as byakugō (ビャクゴウ 百合) in traditional pharmacological circles), listing the use of the following species: Lilium lancifolium, Lilium brownii, Lilium brownii var. colchesteri, Lilium pumilum[109] The scales flaked off from the bulbs are used, usually steamed.[109]

In South Korea, the lilium species which are officially listed for medicinal use are 참나리 Lilium lancifolium Thunberg; 당나리 Lilium brownii var. viridulun Baker.[110][111]

In culture

Symbolism

In the Victorian language of flowers, lilies portray love, ardor, and affection for your loved ones, while orange lilies stand for happiness, love, and warmth.[112]

Lilies are the flowers most commonly used at funerals, where they symbolically signify that the soul of the deceased has been restored to the state of innocence.[113]

Lilium formosanum, or Taiwanese lily, is called "the flower of broken bowl" (Chinese: 打碗花) by the elderly members of the Hakka ethnic group. They believe that because this lily grows near bodies of clean water, harming the lily may damage the environment, just like breaking the bowls that people rely on.[114] An alternative explanation is that parents convince children into not taking the lily by convincing the children that their dinner bowls may break if they destroy this flower.

In Western Christianity, Madonna lily or Lilium candidum has been associated with the Virgin Mary since at least the Medieval Era. Medieval and Renaissance depictions of the Virgin Mary, especially at the Annunciation, often show her with these flowers. Madonna lilies are also commonly included in depictions of Christ's resurrection. Lilium longiflorum, the Easter lily, is a symbol of Easter, and Lilium candidum, the Madonna lily, carries a great deal of symbolic value in many cultures. See the articles for more information.

Heraldry

 
A stylized lily flower in the coat of arms of Pohja

The fleur-de-lis, associated primarily with French royalty, is a stylized lily flower.

Lilium bulbiferum has long been recognised as a symbol of the Orange Order in Northern Ireland.[115]

Lilium mackliniae is the state flower of Manipur. Lilium michauxii, the Carolina lily, is the official state flower of North Carolina. Idyllwild, California, hosts the Lemon Lily Festival, which celebrates Lilium parryi.[116] Lilium philadelphicum is the floral emblem of Saskatchewan province in Canada, and is on the flag of Saskatchewan.[117][118][119]

Other plants referred to as lilies

Lily of the valley, flame lilies, daylilies, and water lilies are symbolically important flowers commonly referred to as lilies, but they are not in the genus Lilium.

See also

Explanatory notes

  1. ^ Blasdale cites Bretschneider (1889), but in Bretschneider (1875), "Notes on Chinese Mediaeval Travellers to the West", p. 123, first gives the Chinese name for H. fulva as "kïm châm hōa" as according to João de Loureiro, while he himself only recognized its name as "kin huang hua" 金黃花 or as [黃花菜]; huang-hua ts'ai; 'yellow-flower vegetable' as they were called by Beijing merchants.
  2. ^ The informant, Pelham L. Warren, consul at Taiwan was presumably providing imports from China (main port Hankow) or Japan.
  3. ^ "not a common food" (Shizuo Tsuji [ja]).
  4. ^ The term uragoshi [ja] "straining" orthodoxically means using the "uragoshi-ki", traditionally a sieve with a fine mesh of horse-hair instead of metal wire.
  5. ^ These could refer to essentially the same thing, except for slight difference in texture and appearance. The yuri-kinton has been described as "ogura an (sweet adzuki bean paste) core surrounded with stipples (soboro) of strained lily bulb and white adzuki (shiroazuki or shiroshōzu).[90] A recipe for lily bulb dumplings or chakin-shibori calls for wrapping adzuki bean paste with lily bulb mashed into purée, then wrapping it in a cloth and wringing the dumpling into a ball shape.[89]
  6. ^ Jimyōin Motoaki [ja] b. 1865 was a viscount and poet. So was his son Motonori.
  7. ^ And as discussed below, this yama-yuri was also called "hime-yuri" in earlier days.[95]
  8. ^ The kooni yuri (小鬼百合, "lesser ogre lily").
  9. ^ That is, not in the top three of this period.[95]

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Citations
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Bibliography
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External links

  • The Plant List
  • North American Lily Society
  • Royal Horticultural Society Lily Group
  • 1 2 3 Time-lapse videos
  • THE GENUS LILIUM
  • "Lilium" at the Encyclopedia of Life  
  • Lily perenialization, Flower Bulb Research Program, Department of Horticulture, Cornell University
  • .

Flora

  • Flora Europaea: Lilium
  • Flora of China: Lilium
  • Flora of Nepal: Lilium species list
  • Flora of North America: Lilium

lilium, lily, lilies, redirect, here, other, uses, disambiguation, lily, disambiguation, lilies, disambiguation, other, plants, called, lilies, list, plants, known, lily, genus, herbaceous, flowering, plants, growing, from, bulbs, with, large, prominent, flowe. Lily and Lilies redirect here For other uses see Lilium disambiguation Lily disambiguation and Lilies disambiguation For other plants called lilies see List of plants known as lily Lilium ˈ l ɪ l i e m LIL ee em 3 is a genus of herbaceous flowering plants growing from bulbs all with large prominent flowers They are the true lilies Lilies are a group of flowering plants which are important in culture and literature in much of the world Most species are native to the northern hemisphere and their range is temperate climates and extends into the subtropics Many other plants have lily in their common names but do not belong to the same genus and are therefore not true lilies LiliumLilium candidumScientific classificationKingdom PlantaeClade TracheophytesClade AngiospermsClade MonocotsOrder LilialesFamily LiliaceaeSubfamily LilioideaeTribe LilieaeGenus LiliumL 1 Type speciesLilium candidumL 2 SpeciesList of Lilium speciesSynonyms 1 Lirium Scop nom illeg Martagon WolfMartagon Rchb Opiz nom illeg Nomocharis Franch Contents 1 Description 2 Taxonomy 2 1 Etymology 3 Distribution and habitat 4 Ecology 5 Cultivation 5 1 Awards 5 2 Classification of garden forms 5 2 1 Asiatic hybrids Division I 5 2 2 Martagon hybrids Division II 5 2 3 Candidum Euro Caucasian hybrids Division III 5 2 4 American hybrids Division IV 5 2 5 Longiflorum hybrids Division V 5 2 6 Trumpet lilies Division VI including Aurelian hybrids with L henryi 5 2 7 Oriental hybrids Division VII 5 2 8 Other hybrids Division VIII 5 2 9 Species Division IX 5 3 Pests and diseases 5 4 Propagation and growth 6 Research 7 Toxicity 8 Culinary uses 8 1 Chinese cuisine 8 2 Japanese cuisine 8 2 1 Yokan 8 2 2 Species used 8 3 North America 9 Medicinal uses 10 In culture 10 1 Symbolism 10 2 Heraldry 10 3 Other plants referred to as lilies 11 See also 12 Explanatory notes 13 References 14 External links 14 1 FloraDescription Edit Lilium longiflorum flower 1 stigma 2 style 3 anthers 4 filament 5 tepal Lilies are tall perennials ranging in height from 2 6 ft 60 180 cm They form naked or tunicless scaly underground bulbs which are their organs of perennation In some North American species the base of the bulb develops into rhizomes on which numerous small bulbs are found Some species develop stolons 4 Most bulbs are buried deep in the ground but a few species form bulbs near the soil surface Many species form stem roots With these the bulb grows naturally at some depth in the soil and each year the new stem puts out adventitious roots above the bulb as it emerges from the soil These roots are in addition to the basal roots that develop at the base of the bulb a number of species also produce contractile roots that move the bulbs deeper into the soil 5 Lily petal The flowers are large often fragrant and come in a wide range of colors including whites yellows oranges pinks reds and purples Markings include spots and brush strokes The plants are late spring or summer flowering Flowers are borne in racemes or umbels at the tip of the stem with six tepals spreading or reflexed to give flowers varying from funnel shape to a Turk s cap The tepals are free from each other and bear a nectary at the base of each flower The ovary is superior borne above the point of attachment of the anthers The fruit is a three celled capsule 6 Stamen of lilium Seeds ripen in late summer They exhibit varying and sometimes complex germination patterns many adapted to cool temperate climates Most cool temperate species are deciduous and dormant in winter in their native environment But a few species native to areas with hot summers and mild winters Lilium candidum Lilium catesbaei Lilium longiflorum lose their leaves and enter a short dormant period in summer or autumn sprout from autumn to winter forming dwarf stems bearing a basal rosette of leaves until after they have received sufficient chilling the stem begins to elongate in warming weather Lilium candidum seeds The basic chromosome number is twelve n 12 7 Taxonomy EditTaxonomical division in sections follows the classical division of Comber 8 species acceptance follows the World Checklist of Selected Plant Families 9 the taxonomy of section Pseudolirium is from the Flora of North America 10 the taxonomy of Section Liriotypus is given in consideration of Resetnik et al 2007 11 the taxonomy of Chinese species various sections follows the Flora of China 12 and the taxonomy of Section Sinomartagon follows Nishikawa et al 13 as does the taxonomy of Section Archelirion 14 The World Checklist of Selected Plant Families as of January 2014 update considers Nomocharis a separate genus in its own right 15 however some authorities consider Nomocharis to be embedded within Lilium rather than treat it as a separate genus 16 17 The Sinomartagon are divided in 3 paraphyletic groups while the Leucolirion are divided in 2 paraphyletic groups 18 There are seven sections Martagon Pseudolirium Liriotypus Archelirion Sinomartagon Leucolirion DaurolirionThere are 111 species counted in this genus 19 For a full list of accepted species 1 with their native ranges see List of Lilium species Picture Section Sub Section Botanical name common name Martagon Lilium distichum Martagon Lilium hansonii Martagon Lilium martagon Martagon or Turk s cap lily Martagon Lilium medeoloides Martagon Lilium tsingtauense Pseudolirium 2a Lilium bolanderi Bolander s Lily Pseudolirium 2a Lilium puberulum Pseudolirium 2a Lilium kelloggii Pseudolirium 2a Lilium rubescens Pseudolirium 2a Lilium washingtonianum Washington Lily Shasta Lily or Mt Hood Lily Pseudolirium 2b Lilium kelleyanum Pseudolirium 2b Lilium maritimum Pseudolirium 2b Lilium occidentale Pseudolirium 2b Lilium pardalinum Panther or Leopard lily Pseudolirium 2b Lilium parryi Pseudolirium 2b Lilium parvum Sierra tiger lily or Alpine lily Pseudolirium 2c Lilium canadense Canada Lily or Meadow Lily Pseudolirium 2c Lilium grayi Pseudolirium 2c Lilium iridollae Pseudolirium 2c Lilium michiganense Michigan Lily Pseudolirium 2c Lilium michauxii Carolina Lily Pseudolirium 2c Lilium superbum Swamp lily or American tiger lily Pseudolirium 2c Lilium pyrophilum Sandhills Lily 20 Pseudolirium 2d Lilium catesbaei Pseudolirium 2d Lilium philadelphicum Wood lily Philadelphia lily or prairie lily Liriotypus 3a Lilium candidum Madonna lily Liriotypus 3b Lilium albanicum Liriotypus 3b Lilium bosniacum Lilium carniolicum var bosniacum Liriotypus 3b Lilium chalcedonicum Liriotypus 3b Lilium carniolicum Liriotypus 3b Lilium ciliatumLiriotypus 3b Lilium heldreichii Liriotypus 3b Lilium jankae Liriotypus 3b Lilium pomponium Turban lily Liriotypus 3b Lilium ponticum Liriotypus 3b Lilium pyrenaicum Liriotypus 3c Lilium akkusianum Liriotypus 3c Lilium kesselringianum Liriotypus 3c Lilium monadelphum Liriotypus 3c Lilium rhodopeum Liriotypus 3c Lilium szovitsianum Polish Lily Liriotypus 3c Lilium polyphyllum Liriotypus 3c Lilium ledebourii Liriotypus 3d Lilium bulbiferum Orange Lily or Fire Lily Archelirion 4a Lilium speciosum Japanese lily Archelirion 4b Lilium auratum Golden rayed lily of Japan or Goldband lily Archelirion 4c Lilium alexandrae Archelirion 4c Lilium japonicum Archelirion 4c Lilium nobilissimum Archelirion 4d Lilium brownii Archelirion 4d Lilium rubellum Archelirion 4d Lilium platyphyllum Sinomartagon 5a Lilium davidii Sinomartagon 5a Lilium duchartrei Sinomartagon 5a Lilium henryi Tiger Lily or Henry s lily Sinomartagon 5a Lilium lancifolium Tiger Lily often known as L tigrinum Sinomartagon 5a Lilium lankongense Sinomartagon 5a Lilium leichtlinii Sinomartagon 5a Lilium papilliferum Sinomartagon 5a Lilium rosthornii Sinomartagon 5b Lilium amabile Sinomartagon 5b Lilium callosum Sinomartagon 5b Lilium cernuum Sinomartagon 5b Lilium concolor Morning Star Lily Sinomartagon 5b Lilium fargesii Sinomartagon 5b Lilium pumilum Coral Lily Low Lily or Siberian LilySinomartagon 5b Lilium xanthellum Sinomartagon 5c Lilium amoenumSinomartagon 5c Lilium arboricola Sinomartagon 5c Lilium bakerianum Sinomartagon 5c Lilium euxanthumSinomartagon 5c Lilium henrici Sinomartagon 5c Lilium lophophorum Sinomartagon 5c Lilium mackliniae Siroi Lily Sinomartagon 5c Lilium majoense Sinomartagon 5c Lilium nanum Sinomartagon 5c Lilium nepalense Sinomartagon 5c Lilium oxypetalumSinomartagon 5c Lilium paradoxum Sinomartagon 5c Lilium poilanei Sinomartagon 5c Lilium primulinumSinomartagon 5c Lilium sempervivoideumSinomartagon 5c Lilium sherriffiae Sinomartagon 5c Lilium soulieiSinomartagon 5c Lilium stewartianum Sinomartagon 5c Lilium taliense Sinomartagon 5c Lilium wardiiSinomartagon 5 Lilium brevistylum Sinomartagon 5 Lilium lijiangenseSinomartagon 5 Lilium anhuienseSinomartagon 5 Lilium eupetes Sinomartagon 5 Lilium habaenseSinomartagon 5 Lilium huidongenseSinomartagon 5 Lilium jinfushanenseSinomartagon 5 Lilium matangenseSinomartagon 5 Lilium medogenseSinomartagon 5 Lilium pinifoliumSinomartagon 5 Lilium pyiSinomartagon 5 Lilium saccatumSinomartagon 5 Lilium tianschanicumSinomartagon 5 Lilium floridum Leucolirion 6a Lilium leucanthum Leucolirion 6a Lilium regale Leucolirion 6a Lilium sargentiae Leucolirion 6a Lilium sulphureum Leucolirion 6a Lilium wenshanenseLeucolirion 6b Lilium anhuiense Leucolirion 6b Lilium formosanum Leucolirion 6b Lilium longiflorum Easter LilyLeucolirion 6b Lilium neilgherrense Leucolirion 6b Lilium philippinense Benguet lily 21 22 Leucolirion 6b Lilium wallichianumLeucolirion 6b Lilium zairii Leucolirion 6b Lilium puerense Daurolirion Lilium dauricum Daurolirion Lilium maculatum Daurolirion Lilium pensylvanicumLilium eupetes Lilium armenum Lilium bosniacum Lilium columbianum Lilium debile Lilium humboldtiiLilium rockiiSome species formerly included within this genus have now been placed in other genera These genera include Cardiocrinum Notholirion Nomocharis and Fritillaria Etymology Edit The botanic name Lilium is the Latin form and is a Linnaean name The Latin name is derived from the Greek word leirion leirion generally assumed to refer to true white lilies as exemplified by the Madonna lily 23 24 25 The word was borrowed from Coptic dial Fayyumic hleri from standard hreri from Demotic hrry from Egyptian hrṛt flower citation needed Meillet maintains that both the Egyptian and the Greek word are possible loans from an extinct substratum language of the Eastern Mediterranean citation needed Ancient Greek krῖnon krinon was used by the Greeks albeit for lilies of any color 26 The term lily has in the past been applied to numerous flowering plants often with only superficial resemblance to the true lily including water lily fire lily lily of the Nile calla lily trout lily kaffir lily cobra lily lily of the valley daylily ginger lily Amazon lily leek lily Peruvian lily and others All English translations of the Bible render the Hebrew shushan shōshan shōshanna as lily but the lily among the thorns of Song of Solomon for instance may be the honeysuckle 27 Distribution and habitat Edit Turk s cap lily growing in the Carinthian Alps The range of lilies in the Old World extends across much of Europe across most of Asia to Japan south to India and east to Indochina and the Philippines In the New World they extend from southern Canada through much of the United States They are commonly adapted to either woodland habitats often montane or sometimes to grassland habitats A few can survive in marshland and epiphytes are known in tropical southeast Asia In general they prefer moderately acidic or lime free soils Ecology EditLilies are used as food plants by the larvae of some Lepidoptera species including the Dun bar The proliferation of deer e g Odocoileus virginianus in North America mainly due to factors such as the elimination of large predators for human safety is responsible there for a downturn in lily populations in the wild and is a threat to garden lilies as well 28 Fences as high as 8 feet may be required to prevent them from consuming the plants an impractical solution for most wild areas 29 Cultivation EditMany species are widely grown in the garden in temperate sub tropical and tropical regions 30 Numerous ornamental hybrids have been developed They are used in herbaceous borders woodland and shrub plantings and as patio plants Some lilies especially Lilium longiflorum form important cut flower crops or potted plants These are forced to flower outside of the normal flowering season for particular markets for instance Lilium longiflorum for the Easter trade when it may be called the Easter lily Lilies are usually planted as bulbs in the dormant season They are best planted in a south facing northern hemisphere slightly sloping aspect in sun or part shade at a depth 2 times the height of the bulb except Lilium candidum which should be planted at the surface Most prefer a porous loamy soil and good drainage is essential Most species bloom in July or August northern hemisphere The flowering periods of certain lily species begin in late spring while others bloom in late summer or early autumn 31 They have contractile roots which pull the plant down to the correct depth therefore it is better to plant them too shallowly than too deep A soil pH of around 6 5 is generally safe Most grow best in well drained soils and plants are watered during the growing season Some species and cultivars have strong wiry stems but those with heavy flower heads are staked to stay upright 32 33 Awards Edit The following lily species and cultivars currently hold the Royal Horticultural Society s Award of Garden Merit confirmed 2017 34 Golden Splendor African Queen Group VI a 2002 H6 Casa Blanca VIIb b c 1993 H6 Fata Morgana Ia b 2002 H6 Garden Party VIIb b 2002 H6 Golden Splendor Group VIb c a 35 Lilium henryi IXc d 1993 H6 Lilium mackliniae IXc a 2012 H5 Lilium martagon Turk s cap lily IXc d 36 Lilium pardalinum leopard lily IXc d 37 Pink Perfection Group VIb a 38 Lilium regale regal lily king s lily IXb a 39 Classification of garden forms Edit Numerous forms mostly hybrids are grown for the garden They vary according to the species and interspecific hybrids that they derived from and are classified in the following broad groups 40 41 42 Asiatic hybrids Division I Edit These are derived from hybrids between species in Lilium section Sinomartagon 43 44 They are derived from central and East Asian species and interspecific hybrids including Lilium amabile Lilium bulbiferum Lilium callosum Lilium cernuum Lilium concolor Lilium dauricum Lilium davidii Lilium hollandicum Lilium lancifolium syn Lilium tigrinum Lilium lankongense Lilium leichtlinii Lilium maculatum Lilium pumilum Lilium scottiae Lilium wardii and Lilium wilsonii These are plants with medium sized upright or outward facing flowers mostly unscented There are various cultivars such as Lilium Cappuccino Lilium Dimension Lilium Little Kiss and Lilium Navona 45 Dwarf Patio Border varieties are much shorter c 36 61 cm in height and were designed for containers 46 They often bear the cultivar name Tiny such as the Lily Looks series e g Tiny Padhye 47 Tiny Dessert 48 Martagon hybrids Division II Edit These are based on Lilium dalhansonii Lilium hansonii Lilium martagon Lilium medeoloides and Lilium tsingtauense The flowers are nodding Turk s cap style with the petals strongly recurved Candidum Euro Caucasian hybrids Division III Edit This includes mostly European species Lilium candidum Lilium chalcedonicum Lilium kesselringianum Lilium monadelphum Lilium pomponium Lilium pyrenaicum and Lilium testaceum American hybrids Division IV Edit These are mostly taller growing forms originally derived from Lilium bolanderi Lilium burbankii Lilium canadense Lilium columbianum Lilium grayi Lilium humboldtii Lilium kelleyanum Lilium kelloggii Lilium maritimum Lilium michauxii Lilium michiganense Lilium occidentale Lilium pardaboldtii Lilium pardalinum Lilium parryi Lilium parvum Lilium philadelphicum Lilium pitkinense Lilium superbum Lilium ollmeri Lilium washingtonianum and Lilium wigginsii Many are clump forming perennials with rhizomatous rootstocks Longiflorum hybrids Division V Edit These are cultivated forms of this species and its subspecies They are most important as plants for cut flowers and are less often grown in the garden than other hybrids Trumpet lilies Division VI including Aurelian hybrids with L henryi Edit This group includes hybrids of many Asiatic species and their interspecific hybrids including Lilium aurelianense Lilium brownii Lilium centigale Lilium henryi Lilium imperiale Lilium kewense Lilium leucanthum Lilium regale Lilium rosthornii Lilium sargentiae Lilium sulphureum and Lilium sulphurgale The flowers are trumpet shaped facing outward or somewhat downward and tend to be strongly fragrant often especially night fragrant Oriental hybrids Division VII Edit These are based on hybrids within Lilium section Archelirion 43 44 specifically Lilium auratum and Lilium speciosum together with crossbreeds from several species native to Japan including Lilium nobilissimum Lilium rubellum Lilium alexandrae and Lilium japonicum They are fragrant and the flowers tend to be outward facing Plants tend to be tall and the flowers may be quite large The whole group are sometimes referred to as stargazers because many of them appear to look upwards For the specific cultivar see Lilium Stargazer Other hybrids Division VIII Edit Includes all other garden hybrids Species Division IX Edit All natural species and naturally occurring forms are included in this group The flowers can be classified by flower aspect and form 49 Flower aspect a up facing b out facing c down facingFlower form a trumpet shaped b bowl shaped c flat or with tepal tips recurved d tepals strongly recurved with the Turk s cap form as the ultimate state Many newer commercial varieties are developed by using new technologies such as ovary culture and embryo rescue 50 Pests and diseases Edit Scarlet lily beetles Oxfordshire UK Aphids may infest plants Leatherjackets feed on the roots Larvae of the Scarlet lily beetle can cause serious damage to the stems and leaves The scarlet beetle lays its eggs and completes its life cycle only on true lilies Lilium and fritillaries Fritillaria 51 Oriental rubrum tiger and trumpet lilies as well as Oriental trumpets orienpets and Turk s cap lilies and native North American Lilium species are all vulnerable but the beetle prefers some types over others The beetle could also be having an effect on native Canadian species and some rare and endangered species found in northeastern North America 52 Daylilies Hemerocallis not true lilies are excluded from this category Plants can suffer from damage caused by mice deer and squirrels Slugs 53 snails and millipedes attack seedlings leaves and flowers Brown spots on damp leaves may signal an infection of Botrytis elliptica also known as Lily blight lily fire and botrytis leaf blight 54 Various viral diseases can cause mottling of leaves and stunting of growth including lily curl stripe ringspot and lily rosette virus 55 Propagation and growth Edit Lilies can be propagated in several ways by division of the bulbs by growing on bulbils which are adventitious bulbs formed on the stem by scaling for which whole scales are detached from the bulb and planted to form a new bulb by seed there are many seed germination patterns which can be complex by micropropagation techniques which include tissue culture 56 commercial quantities of lilies are often propagated in vitro and then planted out to grow into plants large enough to sell A highly efficient technique for multiple shoot and propagule formation was given by Yadav et al in 2013 57 Plant grow regulators PGRs are used to limit the height of lilies especial those sold as potted plants commonly used chemicals include ancymidol fluprimidol paclobutrazol and uni conazole which are applied to the foliage and retard the biosynthesis of gibberellins a class of plant hormones responsible for stem growth 58 Research EditA comparison of meiotic crossing over recombination in lily and mouse led in 1977 to the conclusion that diverse eukaryotes share a common pattern of meiotic crossing over 59 Lilium longiflorum has been used for studying aspects of the basic molecular mechanism of genetic recombination during meiosis 60 61 Toxicity EditSome Lilium species are toxic to cats This is known to be so especially for Lilium longiflorum though other Lilium and the unrelated Hemerocallis can also cause the same symptoms 62 63 64 65 The true mechanism of toxicity is undetermined but it involves damage to the renal tubular epithelium composing the substance of the kidney and secreting collecting and conducting urine which can cause acute kidney failure 65 Veterinary help should be sought as a matter of urgency for any cat that is suspected of eating any part of a lily including licking pollen that may have brushed onto its coat 66 Culinary uses EditChinese cuisine Edit Lily bulbs are starchy and edible as root vegetables though bulbs of some species may be too bitter to eat 67 Lilium brownii var viridulum known as 百合 pak hop pinyin bǎi he Cantonese Yale baak hap lit hundred united is one of the most prominent edible lilies in China Its bulbs are large in size and not bitter They were even exported and sold in the San Francisco Chinatown in the 19th century available both fresh and dry 67 A landrace called 龍牙百合 pinyin long ya bǎi he lit dragon tooth lily mainly cultivated in Hunan and Jiangxi is especially renowned for its good quality bulbs 68 L lancifolium Chinese 卷丹 pinyin juǎn dan lit reflexed red is widely cultivated in China especially in Yixing Huzhou and Longshan Its bulbs are slightly bitter 68 L davidii var unicolor Chinese 蘭州百合 lit Lanzhou lily is mainly cultivated in Lanzhou and its bulbs are valued for sweetness 68 Other edible Chinese lilies include L brownii var brownii L davidii var davidii L concolor L pensylvanicum L distichum L martagon var pilosiusculum L pumilum L rosthornii and L speciosum var gloriosoides 69 Researchers have also explored the possibility of using ornamental cultivars as edible lilies 72 The dried bulbs are commonly used in the south to flavor soup citation needed They may be reconstituted and stir fried grated and used to thicken soup or processed to extract starch citation needed Their texture and taste draw comparisons with the potato although the individual bulb scales are much smaller citation needed The commonly marketed lily flower buds called 金针菜 kam cham tsoi 73 pinyin jin zhen cai Cantonese Yale gamjam choi lit gold needle vegetable in Chinese cuisine are actually from daylilies Hemerocallis citrina 74 or possibly H fulva a 73 Flowers of the H graminea and Lilium bulbiferum were reported to have been eaten as well but samples provided by the informant were strictly daylilies and did not include L bulbiferum b 75 Lily flowers and bulbs are eaten especially in the summer for their perceived ability to reduce internal heat 76 A 19th century English source reported that Lily flowers are also said to be efficacious in pulmonary affections and to have tonic properties 75 Asiatic lily cultivars are also imported from the Netherlands the seedling bulbs must be imported from the Netherlands every year 77 78 79 The parts of Lilium species which are officially listed as food material in Taiwan are the flower and bulbs of Lilium lancifolium Lilium brownii var viridulum Lilium pumilum and Lilium candidum 80 Japanese cuisine Edit Loose scales of lily bulb in a donburi bowl dish The lily bulb or yuri ne is sometimes used in Japanese cuisine c 81 It may be most familiar in the present day as an occasional ingredient 具 gu in the chawan mushi savoury egg custard 82 where a few loosened scales of this optional ingredient are found embedded in the hot pudding of each serving 83 84 It could also be used as an ingredient in a clear soup or suimono ja 85 86 The boiled bulb may also be strained d into puree for use as in the sweetened kinton 87 88 or chakin shibori 88 89 e Yokan Edit There is also the yuri yōkan one recipe of which calls for combining measures of yuri starch with agar dissolved in water and sugar 91 This was a specialty of Hamada Shimane 92 and the shop Kaisei dō 開盛堂 established in 1885 became famous for it 93 94 Because a certain Viscount Jimyōin wrote a waka poem about the confection which mentioned hime yuri princess lily f one source stated that the hime yuri usually taken to mean L concolor had to have been used 93 but another source points out that the city of Hamada lies back to back with across a mountain range with Fuchu Hiroshima which is renowned for its production of yama yuri L auratum 90 g Species used Edit Current Japanese governmental sources c 2005 list the following lily species as prominent in domestic consumption 98 99 the oni yuri or tiger lily Lilium lancifolium the kooni yuri Lilium leichtlinii var maximowiczii h and the gold banded white yama yuri L auratum But Japanese sources c 1895 1900 95 100 give a top three list which replaces kooni yuri with the sukashi yuri 透かし百合 lit see through lily L maculatum named from the gaps between the tepals 101 102 There is uncertainty regarding which species is meant by the hime yuri used as food because although this is usually the common name for L concolor in most up to date literature 103 it used to ambiguously referred to the tiger lily as well c 1895 1900 95 The non tiger lily himeyuri is certainly described as quite palatable in the literature at the time but the extent of exploitation could not have been as significant i North America Edit The flower buds and roots of Lilium canadense are traditionally gathered and eaten by North American indigenous peoples 104 Coast Salish Nuu chah nulth and most western Washington peoples steam boil or pit cook the bulbs of Lilium columbianum Bitter or peppery tasting they were mostly used as a flavoring often in soup with meat or fish 105 Medicinal uses EditTraditional Chinese medicine list the use of the following 野百合 Lilium brownii 百合 Lilium brownii var viridulum 渥丹 Lilium concolor 毛百合 Lilium dauricum 卷丹 Lilium lancifolium 山丹 Lilium pumilum 南川百合 Lilium rosthornii 药百合Lilium speciosum var gloriosoides 淡黄花百合 Lilium sulphureum 106 107 In Taiwan governmental publications list Lilium lancifolium Thunb Lilium brownii var viridulum Baker Lilium pumilum DC 108 In the kanpō or Chinese medicine as practiced in Japan the official Japanese governmental pharmacopeia Nihon yakkyokuhō 日本薬局方 includes the use of lily bulb known as byakugō ビャクゴウ 百合 in traditional pharmacological circles listing the use of the following species Lilium lancifolium Lilium brownii Lilium brownii var colchesteri Lilium pumilum 109 The scales flaked off from the bulbs are used usually steamed 109 In South Korea the lilium species which are officially listed for medicinal use are 참나리 Lilium lancifolium Thunberg 당나리 Lilium brownii var viridulun Baker 110 111 In culture EditSymbolism Edit In the Victorian language of flowers lilies portray love ardor and affection for your loved ones while orange lilies stand for happiness love and warmth 112 Lilies are the flowers most commonly used at funerals where they symbolically signify that the soul of the deceased has been restored to the state of innocence 113 Lilium formosanum or Taiwanese lily is called the flower of broken bowl Chinese 打碗花 by the elderly members of the Hakka ethnic group They believe that because this lily grows near bodies of clean water harming the lily may damage the environment just like breaking the bowls that people rely on 114 An alternative explanation is that parents convince children into not taking the lily by convincing the children that their dinner bowls may break if they destroy this flower In Western Christianity Madonna lily or Lilium candidum has been associated with the Virgin Mary since at least the Medieval Era Medieval and Renaissance depictions of the Virgin Mary especially at the Annunciation often show her with these flowers Madonna lilies are also commonly included in depictions of Christ s resurrection Lilium longiflorum the Easter lily is a symbol of Easter and Lilium candidum the Madonna lily carries a great deal of symbolic value in many cultures See the articles for more information Heraldry Edit A stylized lily flower in the coat of arms of Pohja The fleur de lis associated primarily with French royalty is a stylized lily flower Lilium bulbiferum has long been recognised as a symbol of the Orange Order in Northern Ireland 115 Lilium mackliniae is the state flower of Manipur Lilium michauxii the Carolina lily is the official state flower of North Carolina Idyllwild California hosts the Lemon Lily Festival which celebrates Lilium parryi 116 Lilium philadelphicum is the floral emblem of Saskatchewan province in Canada and is on the flag of Saskatchewan 117 118 119 Other plants referred to as lilies Edit Lily of the valley flame lilies daylilies and water lilies are symbolically important flowers commonly referred to as lilies but they are not in the genus Lilium See also EditLily seed germination types List of plants known as lily Lilium nanumExplanatory notes Edit Blasdale cites Bretschneider 1889 but in Bretschneider 1875 Notes on Chinese Mediaeval Travellers to the West p 123 first gives the Chinese name for H fulva as kim cham hōa as according to Joao de Loureiro while he himself only recognized its name as kin huang hua 金黃花 or as 黃花菜 huang hua ts ai yellow flower vegetable as they were called by Beijing merchants The informant Pelham L Warren consul at Taiwan was presumably providing imports from China main port Hankow or Japan not a common food Shizuo Tsuji ja The term uragoshi ja straining orthodoxically means using the uragoshi ki traditionally a sieve with a fine mesh of horse hair instead of metal wire These could refer to essentially the same thing except for slight difference in texture and appearance The yuri kinton has been described as ogura an sweet adzuki bean paste core surrounded with stipples soboro of strained lily bulb and white adzuki shiroazuki or shiroshōzu 90 A recipe for lily bulb dumplings or chakin shibori calls for wrapping adzuki bean paste with lily bulb mashed into puree then wrapping it in a cloth and wringing the dumpling into a ball shape 89 Jimyōin Motoaki ja b 1865 was a viscount and poet So was his son Motonori And as discussed below this yama yuri was also called hime yuri in earlier days 95 The kooni yuri 小鬼百合 lesser ogre lily That is not in the top three of this period 95 References EditCitations a b c Lilium World Checklist of Selected Plant Families Royal Botanic Gardens Kew Archived from the original on November 14 2017 Retrieved June 13 2014 lectotype designated by N L Britton et A Brown Ill Fl N U S ed 2 1 502 1913 lilium Merriam Webster Dictionary Batygina T B 2019 04 23 Embryology of Flowering Plants Terminology and Concepts Vol 3 Reproductive Systems CRC Press ISBN 978 0 429 52671 8 Gracie Carol 2020 04 28 Summer Wildflowers of the Northeast A Natural History Princeton University Press ISBN 978 0 691 20330 0 European Garden Flora Volume 1 Pelkonen Veli Pekka Pirttila Anna Maria 2012 Taxonomy and Phylogeny of the Genus Lilium PDF Floriculture and Ornamental Biotechnology 6 Special Issue 2 1 8 Archived from the original PDF on 2016 10 08 Retrieved 2016 07 29 Harold Comber 1949 A new classification of the genus Lilium Lily Yearbook Royal Hortic Soc London 15 86 105 Govaerts R ed Lilium World Checklist of Selected Plant Families The Board of Trustees of the Royal Botanic Gardens Kew Archived from the original on 2020 09 01 Retrieved 2013 02 03 Flora of North America Vol 26 Online Archived 2012 10 15 at the Wayback Machine Resetnik I Liber Z Satovic Z Cigic P Nikolic T 2007 Molecular phylogeny and systematics of the Lilium carniolicum group Liliaceae based on nuclear ITS sequences Plant Systematics and Evolution 265 1 2 45 58 doi 10 1007 s00606 006 0513 y S2CID 32644749 Flora of China Vol 24 eFloras org Archived 2012 09 12 at the Wayback Machine Nishikawa Tomotaro Okazaki Keiichi Arakawa Katsuro Nagamine Tsukasa 2001 Phylogenetic Analysis of Section Sinomartagon in Genus Lilium Using Sequences of the Internal Transcribed Spacer Region in Nuclear Ribosomal DNA 育種学雑誌 Breeding Science 51 1 39 46 doi 10 1270 jsbbs 51 39 Nishikawa Tomotaro Okazaki Keiichi Nagamine Tsukasa 2002 Phylogenetic Relationships among Lilium auratum Lindley L auratum var platyphyllum Baker and L rubellum Baker Based on Three Spacer Regions in Chloroplast DNA 育種学雑誌 Breeding Science 52 3 207 213 doi 10 1270 jsbbs 52 207 WCLSPF 2014 Gao et al 2011 Ronsted et al 2005 Li Juan Cai Jing Qin Huan Huan Price Megan Zhang Zhen Yu Yan Xie Deng Feng He Xing Jin Zhou Song Dong Gao Xin Fen 2022 Phylogeny Age and Evolution of Tribe Lilieae Liliaceae Based on Whole Plastid Genomes Frontiers in Plant Science 12 699226 doi 10 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original on 2017 08 25 Retrieved 2017 08 19 Pojar Jim 2004 Plants of the Pacific Northwest Coast Edmonton Lone Pine Publishing ISBN 9781551055305 中国药用植物 据 中国植物志 全书记载分析而得 Archived from the original on 2014 02 02 Retrieved 2014 01 25 中国药用植物 据 中国植物志 全书记载分析而得 Archived from the original on 2014 02 02 Retrieved 2014 01 25 Taiwan Herbal Pharmarcopeia Archived 2015 01 28 at the Wayback Machine Ministry of Health and Welfare a b The Japanese Pharmacopoeia 17th edition PDF Japanese Ministry of Health p 1906 Archived from the original PDF on 2020 01 13 Retrieved 2020 01 13 index Archived 2014 07 23 at the Wayback Machine 백합 Archived from the original on 2016 09 20 Retrieved 2020 09 01 Lilii Bulbus PDF in Korean Archived PDF from the original on 2014 02 03 Retrieved 2014 01 25 Symbolism of the Lily The Flower That is a Part of History Buzzle Archived from the original on 2016 11 26 Retrieved 2016 11 26 Meaning amp Symbolism of Lilies Archived from the original on 2018 04 06 Retrieved 2018 04 06 Liberty Times 2015 魯凱六角星 原民聖花 客庄打碗花 生態指標 in Chinese 自由電子報 Archived from the original on 2013 12 02 Design Research Group 27 June 2007 A kinder gentler image Modernism Tradition and the new Orange Order logo Reinventing the Orange Order A superhero for the 21st century Archived from the original on 3 May 2016 Retrieved 17 September 2016 Lemon Lily Festival Archived 2010 07 26 at the Wayback Machine Government House Gardens Showcase Western Red Lily Government of Saskatchewan 2005 07 21 Archived from the original on 2011 06 11 Retrieved 2008 07 09 Saskatchewan s Provincial Flower Government of Saskatchewan Archived from the original on 2011 07 28 Retrieved 2008 07 09 designated in 1941 Saskatchewan Government of Canada 2013 08 20 Archived from the original on 2015 07 21 Retrieved 2015 07 18 BibliographyGao Yun Dong Hohenegger Markus Harris AJ Zhou Song Dong He Xing Jin Wan Juan 2012 A new species in the genus Nomocharis Franchet Liliaceae evidence that brings the genus Nomocharis into Lilium Plant Systematics and Evolution 298 1 69 85 doi 10 1007 s00606 011 0524 1 ISSN 0378 2697 S2CID 16912824 Ronsted N Law S Thornton H Fay M F Chase M W 2005 Molecular phylogenetic evidence for the monophyly of Fritillaria and Lilium Liliaceae Liliales and the infrageneric classification of Fritillaria Molecular Phylogenetics and Evolution 35 3 509 527 doi 10 1016 j ympev 2004 12 023 PMID 15878122 Nomocharis World Checklist of Selected Plant Families Royal Botanic Gardens Kew archived from the original on 2020 09 01 retrieved 2015 09 14 yuri ユリ in Nihon shakai jii 日本社會事彙 in Japanese Vol 2 Keizai zasshi sha 1908 pp 2077 2083 Archived from the original on 2020 09 01 Retrieved 2020 01 16 digested from Shin shikunshi Seika en Sanjin 精花園山人 Hana yuri 花百合 in Shōkadō Shujin 1901 Shin shikunshi 新四君子 in Japanese Tokyo Mita Ikushujyo pp 63 140 archived from the original on 2020 02 10 retrieved 2020 01 16External links Edit Look up lilium in Wiktionary the free dictionary Wikiquote has quotations related to Lilies Wikimedia Commons has media related to Lilium Wikispecies has information related to Lilium The Plant List North American Lily Society Royal Horticultural Society Lily Group 1 2 3 Time lapse videos THE GENUS LILIUM Lilium at the Encyclopedia of Life Lily perenialization Flower Bulb Research Program Department of Horticulture Cornell University Crossing polygon of the genus Lilium Bulb flower production Lilies International Flower Bulb Centre Lily Picture Book International Flower Bulb CentreFlora Edit Flora Europaea Lilium Flora of China Lilium Flora of Nepal Lilium species list Flora of North America Lilium Retrieved from https en wikipedia org w index php title Lilium amp oldid 1152769597, wikipedia, wiki, book, books, library,

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