fbpx
Wikipedia

Cultivar

A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated. Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting, tissue culture, or carefully controlled seed production. Most cultivars arise from deliberate human manipulation, but some originate from wild plants that have distinctive characteristics. Cultivar names are chosen according to rules of the International Code of Nomenclature for Cultivated Plants (ICNCP), and not all cultivated plants qualify as cultivars. Horticulturists generally believe the word cultivar[nb 1] was coined as a term meaning "cultivated variety".

Osteospermum 'Pink Whirls'
A cultivar selected for its intriguing and colourful flowers

Popular ornamental plants like roses, camellias, daffodils, rhododendrons, and azaleas are commonly cultivars produced by breeding and selection or as sports, for floral colour or size, plant form, or other desirable characteristics.[1] Similarly, the world's agricultural food crops are almost exclusively cultivars that have been selected for characters such as improved yield, flavour, and resistance to disease, and very few wild plants are now used as food sources. Trees used in forestry are also special selections grown for their enhanced quality and yield of timber.

Cultivars form a major part of Liberty Hyde Bailey's broader group, the cultigen,[2] which is defined as a plant whose origin or selection is primarily due to intentional human activity.[3] A cultivar is not the same as a botanical variety,[4] which is a taxonomic rank below subspecies, and there are differences in the rules for creating and using the names of botanical varieties and cultivars. In recent times, the naming of cultivars has been complicated by the use of statutory patents for plants and recognition of plant breeders' rights.[5]

The International Union for the Protection of New Varieties of Plants (UPOV – French: Union internationale pour la protection des obtentions végétales) offers legal protection of plant cultivars to persons or organisations that introduce new cultivars to commerce. UPOV requires that a cultivar be "distinct, uniform", and "stable". To be "distinct", it must have characters that easily distinguish it from any other known cultivar. To be "uniform" and "stable", the cultivar must retain these characters in repeated propagation.

The naming of cultivars is an important aspect of cultivated plant taxonomy, and the correct naming of a cultivar is prescribed by the Rules and Recommendations of the International Code of Nomenclature for Cultivated Plants (ICNCP, commonly denominated the Cultivated Plant Code). A cultivar is given a cultivar name, which consists of the scientific Latin botanical name followed by a cultivar epithet. The cultivar epithet is usually in a vernacular language.

Etymology edit

 
Liberty Hyde Bailey (1858–1954) coined the words cultigen in 1918 and cultivar in 1923.

The word cultivar originated from the need to distinguish between wild plants and those with characteristics that arose in cultivation, presently denominated cultigens. This distinction dates to the Greek philosopher Theophrastus (370–285 BC), the "Father of Botany", who was keenly aware of this difference. Botanical historian Alan Morton noted that Theophrastus in his Historia Plantarum (Enquiry into Plants) "had an inkling of the limits of culturally induced (phenotypic) changes and of the importance of genetic constitution" (Historia Plantarum, Book 3, 2, 2 and Causa Plantarum, Book 1, 9, 3).[6]

The International Code of Nomenclature for algae, fungi, and plants uses as its starting point for modern botanical nomenclature the Latin names in Linnaeus' (1707–1778) Species Plantarum (tenth edition) and Genera Plantarum (fifth edition). In Species Plantarum, Linnaeus enumerated all plants known to him, either directly or from his extensive reading. He recognised the rank of varietas (botanical "variety", a rank below that of species and subspecies) and he indicated these varieties with letters of the Greek alphabet, such as α, β, and λ, before the varietal name, rather than using the abbreviation "var." as is the present convention. Most of the varieties that Linnaeus enumerated were of "garden" origin rather than being wild plants.[7]

In time the need to distinguish between wild plants and those with variations that had been cultivated increased. In the nineteenth century many "garden-derived" plants were given horticultural names, sometimes in Latin and sometimes in a vernacular language. From circa the 1900s, cultivated plants in Europe were recognised in the Scandinavian, Germanic, and Slavic literature as stamm or sorte,[8] but these words could not be used internationally because, by international agreement, any new denominations had to be in Latin.[9] In the twentieth century an improved international nomenclature was proposed for cultivated plants.[10]

Liberty Hyde Bailey of Cornell University in New York, United States created the word cultivar in 1923 when he wrote that:

The cultigen is a species, or its equivalent, that has appeared under domestication – the plant is cultigenous. I now propose another name, cultivar, for a botanical variety, or for a race subordinate to species, that has originated under cultivation; it is not necessarily, however, referable to a recognized botanical species. It is essentially the equivalent of the botanical variety except in respect to its origin.[2]

In that essay, Bailey used only the rank of species for the cultigen, but it was obvious to him that many domesticated plants were more like botanical varieties than species, and that realization appears to have motivated the suggestion of the new category of cultivar.

Bailey created the word cultivar. It is generally assumed to be a blend of cultivated and variety but Bailey never explicitly stated the etymology and it has been suggested that the word is actually a blend of cultigen and variety.[11] The neologism cultivar was promoted as "euphonious" and "free from ambiguity".[8][nb 2] The first Cultivated Plant Code of 1953 subsequently commended its use, and by 1960 it had achieved common international acceptance.[12]

Cultigens edit

 
Bread wheat, Triticum aestivum, is considered a cultigen, and is a distinct species from other wheats according to the biological species concept. Many different cultivars have been created within this cultigen. Many other cultigens are not considered to be distinct species, and can be denominated otherwise.

The words cultigen and cultivar may be confused with each other. A cultigen is any plant that is deliberately selected for or altered in cultivation, as opposed to an indigen; the Cultivated Plant Code states that cultigens are "maintained as recognisable entities solely by continued propagation".[13] Cultigens can have names at any of many taxonomic ranks, including those of grex, species, cultivar group, variety, form, and cultivar; and they may be plants that have been altered in cultivation, including by genetic modification, but have not been formally denominated.[14] A cultigen or a component of a cultigen can be accepted as a cultivar if it is recognisable and has stable characters. Therefore, all cultivars are cultigens, because they are cultivated, but not all cultigens are cultivars, because some cultigens have not been formally distinguished and named as cultivars.

Formal definition edit

The Cultivated Plant Code notes that the word cultivar is used in two different senses: first, as a "classification category" the cultivar is defined in Article 2 of the International Code of Nomenclature for Cultivated Plants (2009, 8th edition) as follows: The basic category of cultivated plants whose nomenclature is governed by this Code is the cultivar.[15] There are two other classification categories for cultigens, the grex[16] and the group.[17] The Code then defines a cultivar as a "taxonomic unit within the classification category of cultivar". This is the sense of cultivar that is most generally understood and which is used as a general definition.

A cultivar is an assemblage of plants that (a) has been selected for a particular character or combination of characters, (b) is distinct, uniform and stable in those characters, and (c) when propagated by appropriate means, retains those characters.[18]

Different kinds edit

 
A cultivar of the orchid genus Oncidium

Which plants are chosen to be named as cultivars is simply a matter of convenience as the category was created to serve the practical needs of horticulture, agriculture, and forestry.[19]

Members of a particular cultivar are not necessarily genetically identical. The Cultivated Plant Code emphasizes that different cultivated plants may be accepted as different cultivars, even if they have the same genome, while cultivated plants with different genomes may be regarded as the same cultivar. The production of cultivars generally entails considerable human involvement although in a few cases it may be as little as simply selecting variation from plants growing in the wild (whether by collecting growing tissue to propagate from or by gathering seed).[20]

Cultivars generally occur as ornamentals and food crops: Malus 'Granny Smith' and Malus 'Red Delicious' are cultivars of apples propagated by cuttings or grafting, Lactuca 'Red Sails' and Lactuca 'Great Lakes' are lettuce cultivars propagated by seeds. Named cultivars of Hosta and Hemerocallis plants are cultivars produced by micropropagation or division.

Clones edit

 
Leucospermum 'Scarlet Ribbon'
A cross performed in Tasmania between L. glabrum and L. tottum

Cultivars that are produced asexually are genetically identical and known as clones; this includes plants propagated by division, layering, cuttings, grafts, and budding. The propagating material may be taken from a particular part of the plant, such as a lateral branch, or from a particular phase of the life cycle, such as a juvenile leaf, or from aberrant growth as occurs with witch's broom. Plants whose distinctive characters are derived from the presence of an intracellular organism may also form a cultivar provided the characters are reproduced reliably from generation to generation. Plants of the same chimera (which have mutant tissues close to normal tissue) or graft-chimeras (which have vegetative tissue from different kinds of plants and which originate by grafting) may also constitute a cultivar.[21]

Seed-produced edit

Some cultivars "come true from seed", retaining their distinguishing characteristics when grown from seed. Such plants are termed a "variety", "selection" or "strain" but these are ambiguous and confusing words that are best avoided. In general, asexually propagated cultivars grown from seeds produce highly variable seedling plants, and should not be labelled with, or sold under, the parent cultivar's name.[22]

Seed-raised cultivars may be produced by uncontrolled pollination when characteristics that are distinct, uniform and stable are passed from parents to progeny. Some are produced as "lines" that are produced by repeated self-fertilization or inbreeding or "multilines" that are made up of several closely related lines. Sometimes they are F1 hybrids which are the result of a deliberate repeatable single cross between two pure lines. A few F2 hybrid seed cultivars also exist, such as Achillea 'Summer Berries'.

Some cultivars are agamospermous plants, which retain their genetic composition and characteristics under reproduction.[23] Occasionally cultivars are raised from seed of a specially selected provenance – for example the seed may be taken from plants that are resistant to a particular disease.[24]

Genetically modified edit

Genetically modified plants with characteristics resulting from the deliberate implantation of genetic material from a different germplasm may form a cultivar. However, the International Code of Nomenclature for Cultivated Plants notes, "In practice such an assemblage is often marketed from one or more lines or multilines that have been genetically modified. These lines or multilines often remain in a constant state of development which makes the naming of such an assemblage as a cultivar a futile exercise."[25] However, retired transgenic varieties such as the fish tomato, which are no longer being developed, do not run into this obstacle and can be given a cultivar name.

Cultivars may be selected because of a change in the ploidy level of a plant which may produce more desirable characteristics.

Cultivar names edit

 
Viola 'Clear Crystals Apricot'
The specific epithet may be omitted from a cultivar name.

Every unique cultivar has a unique name within its denomination class (which is almost always the genus). Names of cultivars are regulated by the International Code of Nomenclature for Cultivated Plants, and may be registered with an International Cultivar Registration Authority (ICRA). There are sometimes separate registration authorities for different plant types such as roses and camellias. In addition, cultivars may be associated with commercial marketing names referred to in the Cultivated Plant Code as "trade designations" (see below).

Presenting in text edit

A cultivar name consists of a botanical name (of a genus, species, infraspecific taxon, interspecific hybrid or intergeneric hybrid) followed by a cultivar epithet. The cultivar epithet is enclosed by single quotes;[26] it should not be italicized if the botanical name is italicized;[27] and each of the words within the epithet is capitalized (with some permitted exceptions such as conjunctions).[28] It is permissible to place a cultivar epithet after a common name provided the common name is botanically unambiguous. Cultivar epithets published before 1 January 1959 were often given a Latin form and can be readily confused with the specific epithets in botanical names; after that date, newly coined cultivar epithets must be in a modern vernacular language to distinguish them from botanical epithets.[29]

For example, the full cultivar name of the King Edward potato is Solanum tuberosum 'King Edward'. 'King Edward' is the cultivar epithet, which, according to the Rules of the Cultivated Plant Code, is bounded by single quotation marks.[26] For patented or trademarked plant product lines developed from a given cultivar, the commercial product name is typically indicated by the symbols "TM" or "®", or is presented in capital letters with no quotation marks, following the cultivar name,[30] as in the following example, where "Bloomerang" is the commercial name and 'Penda' is the cultivar epithet: Syringa 'Penda' BLOOMERANG.[31]

Examples of correct text presentation:
Cryptomeria japonica 'Elegans'
Chamaecyparis lawsoniana 'Aureomarginata' (pre-1959 name, Latin in form)
Chamaecyparis lawsoniana 'Golden Wonder' (post-1959 name, English language)
Pinus densiflora 'Akebono' (post-1959 name, Japanese language)
Apple 'Sundown'
Some incorrect text presentation examples:
Cryptomeria japonica "Elegans" (double quotes are unacceptable)
Berberis thunbergii cv. 'Crimson Pygmy' (this once-common usage is now unacceptable, as it is no longer correct to use "cv." in this context; Berberis thunbergii 'Crimson Pygmy' is correct)
Rosa cv. 'Peace' (this is now incorrect for two reasons: firstly, the use of "cv."; secondly, "Peace" is a trade designation or "selling name" for the cultivar R. 'Madame A. Meilland' and should therefore be printed in a different typeface from the rest of the name, without quote marks, for example: Rosa Peace.)

Although "cv." has not been permitted by the International Code of Nomenclature for Cultivated Plants since the 1995 edition,[32][33] it is still widely used[34] and recommended by other authorities.[35][36]

Group names edit

Where several very similar cultivars exist they can be associated into a Group (formerly Cultivar-group). As Group names are used with cultivar names it is necessary to understand their way of presentation. Group names are presented in normal type and the first letter of each word capitalised as for cultivars, but they are not placed in single quotes. When used in a name, the first letter of the word "Group" is itself capitalized.[37]

Presenting in text edit

Brassica oleracea Capitata Group (the group of cultivars including all typical cabbages)
Brassica oleracea Botrytis Group (the group of cultivars including all typical cauliflowers)
Hydrangea macrophylla Groupe Hortensia (in French) = Hydrangea macrophylla Hortensia Group (in English)
Where cited with a cultivar name the group should be enclosed in parentheses, as follows:
Hydrangea macrophylla (Hortensia Group) 'Ayesha' [38]

Legal protection of cultivars and their names edit

Since the 1990s there has been an increasing use of legal protection for newly produced cultivars. Plant breeders expect legal protection for the cultivars they produce. According to proponents of such protections, if other growers can immediately propagate and sell these cultivars as soon as they come on the market, the breeder's benefit is largely lost.[39] Legal protection for cultivars is obtained through the use of Plant breeders' rights and plant Patents but the specific legislation and procedures needed to take advantage of this protection vary from country to country.[40]

Controversial use of legal protection for cultivars edit

The use of legal protection for cultivars can be controversial, particularly for food crops that are staples in developing countries,[41] or for plants selected from the wild and propagated for sale without any additional breeding work; some people consider this practice unethical.[42]

Trade designations and selling names edit

The formal scientific name of a cultivar, like Solanum tuberosum 'King Edward', is a way of uniquely designating a particular kind of plant. This scientific name is in the public domain and cannot be legally protected. Plant retailers wish to maximize their share of the market and one way of doing this is to replace the cumbersome Latin scientific names on plant labels in retail outlets with appealing marketing names that are easy to use, pronounce, and remember. Marketing names lie outside the scope of the Cultivated Plant Code which refers to them as "trade designations". If a retailer or wholesaler has the sole legal rights to a marketing name then that may offer a sales advantage. Plants protected by plant breeders' rights (PBR) may have a "true" cultivar name – the recognized scientific name in the public domain – and a "commercial synonym" – an additional marketing name that is legally protected. An example would be Rosa Fascination = 'Poulmax', in which Rosa is the genus, Fascination is the trade designation, and 'Poulmax' is scientific cultivar name.

Because a name that is attractive in one language may have less appeal in another country, a plant may be given different selling names from country to country. Quoting the original cultivar name allows the correct identification of cultivars around the world.[43]

The main body coordinating plant breeders' rights is the International Union for the Protection of New Varieties of Plants (Union internationale pour la protection des obtentions végétales, UPOV) and this organization maintains a database of new cultivars protected by PBR in all countries.[44]

International Cultivar Registration Authorities edit

 
Dahlia 'Akita'
A cultivar selected for flower form and colour

An International Cultivar Registration Authority (ICRA) is a voluntary, non-statutory organization appointed by the Commission for Nomenclature and Cultivar Registration of the International Society of Horticultural Science. ICRAs are generally formed by societies and institutions specializing in particular plant genera such as Dahlia or Rhododendron and are currently located in Europe, North America, China, India, Singapore, Australia, New Zealand, South Africa and Puerto Rico.[45]

Each ICRA produces an annual report and its reappointment is considered every four years. The main task is to maintain a register of the names within the group of interest and where possible this is published and placed in the public domain. One major aim is to prevent the duplication of cultivar and Group epithets within a genus, as well as ensuring that names are in accord with the latest edition of the Cultivated Plant Code. In this way, over the last 50 years or so, ICRAs have contributed to the stability of cultivated plant nomenclature. In recent times many ICRAs have also recorded trade designations and trademarks used in labelling plant material, to avoid confusion with established names.[46]

New names and other relevant data are collected by and submitted to the ICRA and in most cases there is no cost. The ICRA then checks each new epithet to ensure that it has not been used before and that it conforms with the Cultivated Plant Code. Each ICRA also ensures that new names are formally established (i.e. published in hard copy, with a description in a dated publication). They record details about the plant, such as parentage, the names of those concerned with its development and introduction, and a basic description highlighting its distinctive characters. ICRAs are not responsible for assessing the distinctiveness of the plant in question.[46] Most ICRAs can be contacted electronically and many maintain web sites for an up-to-date listing.[47]

See also edit

Notes edit

  1. ^ Cultivar (English: /ˈkʌltɪˌvɑːr, -ˌvɛər/) has two meanings, as explained in Formal definition: it is a classification category and a taxonomic unit within the category. When referring to a taxon, the word does not apply to an individual plant but to all plants that share the unique characteristics that define the cultivar.
  2. ^ This ignored its prior use as a transitive verb in Spanish denoting "to farm, to cultivate, to grow, or to practice" (Online Spanish dictionary 2010-01-11 at the Wayback Machine), and in Portuguese denoting to cultivate, to husband, to farm, to plant, to polish, to reclaim, to improve (Ectaco online Portuguese dictionary 2023-04-07 at the Wayback Machine).

References edit

  1. ^ Stanley J. Kays (3 October 2011). Cultivated vegetables of the world: a multilingual onomasticon. Springer. pp. 15–. ISBN 978-90-8686-720-2.
  2. ^ a b Bailey 1923, p. 113
  3. ^ Spencer & Cross 2007, p. 938
  4. ^ Lawrence 1953, pp. 19–20
  5. ^ . Department of Horticulture and Landscape Architecture - Purdue University. Archived from the original on 2012-01-13. Retrieved 2007-07-28. Also Archived 2 August 2012 at archive.today
  6. ^ Morton 1981, pp. 38–39
  7. ^ Lawrence 1955, p. 177
  8. ^ a b Lawrence 1955, p. 180
  9. ^ Lawrence 1955, p. 181
  10. ^ Lawrence 1955, pp. 179–180
  11. ^ Trehane 2004, p. 17
  12. ^ Lawrence 1960, p. 1
  13. ^ Cultivated Plant Code Art. 2.3 Brickell 2009, p. 1
  14. ^ Spencer, Cross & Lumley 2007, p. 47
  15. ^ Cultivated Plant Code. Art. 2.1 Brickell 2009, p. 6
  16. ^ Cultivated Plant Code. Art. 4 Brickell 2009, p. 12
  17. ^ Cultivated Plant Code. Art. 3 Brickell 2009, pp. 10–12
  18. ^ Cultivated Plant Code. Art. 2.2 Brickell 2009, p. 6
  19. ^ Cultivated Plant Code. Preamble & Principles Brickell 2009, p. 19
  20. ^ Cultivated Plant Code, Article 2.20 Brickell 2009, p. 9
  21. ^ Cultivated Plant Code, Articles 2.5–2.11 Brickell 2009, pp. 6–7
  22. ^ Courses / RHS Gardening 2005-12-26 at the Wayback Machine
  23. ^ Cultivated Plant Code, Articles 2.17–2.18 Brickell 2009, pp. 7–8
  24. ^ Cultivated Plant Code, Articles 2.12–2.16 Brickell 2009, pp. 7–8
  25. ^ Cultivated Plant Code, Articles 2.19 Brickell 2009, pp. 8–9
  26. ^ a b Cultivated Plant Code Article 14.1 Brickell 2009, p. 19
  27. ^ Cultivated Plant Code Recommendation 8A.1 Brickell 2009, p. 15
  28. ^ Cultivated Plant Code Article 21.3 Brickell 2009, p. 25
  29. ^ Cultivated Plant Code Art. 14 Brickell 2009, p. 19
  30. ^ Dirr, Michael; Warren, Keith (2019). The Tree Book: Superior Selections for Landscapes, Streetscapes, and Gardens. Timber Press, Inc. ISBN 978-1-60469-714-8.
  31. ^ "Syringa 'Penda' BLOOMERANG". Missouri Botanical Garden Plant Finder. from the original on 2021-11-30. Retrieved 2022-04-29.
  32. ^ "Naming a Cultivar ?". Australian National Botanic Gardens - Botanical Web Portal. 2000-07-28. from the original on 2021-03-12. Retrieved 2020-12-26.
  33. ^ (PDF). Food and Agriculture Organization. 2020-06-10. Archived from the original (PDF) on 2020-12-26. Retrieved 2020-12-26.
  34. ^ Chen, Guang; Liu, Chaolei; Gao, Zhenyu; Zhang, Yu; Zhang, Anpeng; Zhu, Li; Hu, Jiang; Ren, Deyong; Yu, Ling; Xu, Guohua; Qian, Qian (2018-01-05). "Variation in the Abundance of OsHAK1 Transcript Underlies the Differential Salinity Tolerance of an indica and a japonica Rice Cultivar". Frontiers in Plant Science. 8. Frontiers Media SA: 2216. doi:10.3389/fpls.2017.02216. ISSN 1664-462X. PMC 5760540. PMID 29354152.
  35. ^ "Horticulture 202 Laboratory 3". Texas A&M University General Horticulture. from the original on 2020-07-17. Retrieved 2020-12-26.
  36. ^ Finkle, Anita (2018-04-02). "What is the proper way to write a botanical name (Latin name)?". New York Botanical Garden. from the original on 2020-11-26. Retrieved 2020-12-26.
  37. ^ Cultivated Plant Code Art. 3 Brickell 2009, pp. 10–12
  38. ^ Cultivated Plant Code Art. 15 Brickell 2009, p. 19
  39. ^ P. Gepts (2004) Who Owns Biodiversity, and How Should the Owners Be Compensated? Plant Physiology 134, pp. 1295–1307
  40. ^ BSPB Plant breeding – The business and science of crop improvement 2011-09-11 at the Wayback Machine British Society of Plant Breeders booklet
  41. ^ "Adi, A.B.C., Intellectual Property Rights in Biotechnology and the Fate of Poor Farmers' Agriculture. Social Science Research Network". 13 January 2005. from the original on 2021-07-15. Retrieved 2020-07-23.
  42. ^ 'Who owns nature?' 2011-09-11 at the Wayback Machine (article by nurseryman and plant hunter Michael Wickenden, published in The Plantsman)
  43. ^ Spencer, Cross & Lumley 2007, pp. 76–81
  44. ^ Spencer, Cross & Lumley 2007, p. 78
  45. ^ Cultivated Plant Code Brickell 2009, pp. 62, 67–83
  46. ^ a b See International Cultivar Registration Authorities 2017-07-15 at the Wayback Machine
  47. ^ Staff (2010). . ishs.org. Archived from the original on 26 November 2014. Retrieved 5 March 2011.

Bibliography edit

  • Bailey, Liberty Hyde (1923). "Various cultigens, and transfers in nomenclature". Gentes Herbarum. 1 (Part 3): 113–136.
  • Brickell, Chris D. (2009). (PDF). Scripta Horticulturae. 10. International Society of Horticultural Science: 1–184. ISBN 978-90-6605-662-6. Archived from the original (PDF) on 2011-08-13. Retrieved 2011-02-07.
  • Lawrence, George H.M. (1953). "Cultivar, Distinguished from Variety". Baileya. 1: 19–20.
  • Lawrence, George H.M. (1955). "The Term and Category of Cultivar". Baileya. 3: 177–181.
  • Lawrence, George H.M. (1957). "The Designation of Cultivar-names". Baileya. 5: 162–165.
  • Lawrence, George H.M. (1960). "Notes on Cultivar Names". Baileya. 8: 1–4.
  • Morton, Alan G. (1981). History of Botanical Science: An Account of the Development of Botany from Ancient Times to the Present Day. London: Academic Press. ISBN 0-12-508382-3.
  • Spencer, Roger; Cross, Robert; Lumley, Peter (2007). Plant names: a guide to botanical nomenclature. (3rd ed.). Collingwood, Australia: CSIRO Publishing (also Earthscan, UK.). ISBN 978-0-643-09440-6.
  • Spencer, Roger D.; Cross, Robert G. (2007). "The International Code of Botanical Nomenclature (ICBN), the International Code of Nomenclature for Cultivated Plants (ICNCP), and the cultigen". Taxon. 56 (3): 938–940. doi:10.2307/25065875. JSTOR 25065875.
  • Trehane, Piers (2004). "50 years of the International Code of Nomenclature for Cultivated Plants". Acta Horticulturae. 634: 17–27. doi:10.17660/ActaHortic.2004.634.1.

External links edit

  • Sale point of the of The International Code of Nomenclature for Cultivated Plants
  • International Cultivar Registration Authorities
  • (from The Plantsman magazine)
  • Hortivar – The Food and Agriculture Organization of the United Nations Horticulture Cultivars Performance Database

cultivar, taxonomic, rank, just, below, subspecies, variety, botany, cultivar, kind, cultivated, plant, that, people, have, selected, desired, traits, which, retains, those, traits, when, propagated, methods, used, propagate, cultivars, include, division, root. For the taxonomic rank just below subspecies see Variety botany A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated Methods used to propagate cultivars include division root and stem cuttings offsets grafting tissue culture or carefully controlled seed production Most cultivars arise from deliberate human manipulation but some originate from wild plants that have distinctive characteristics Cultivar names are chosen according to rules of the International Code of Nomenclature for Cultivated Plants ICNCP and not all cultivated plants qualify as cultivars Horticulturists generally believe the word cultivar nb 1 was coined as a term meaning cultivated variety Osteospermum Pink Whirls A cultivar selected for its intriguing and colourful flowersPopular ornamental plants like roses camellias daffodils rhododendrons and azaleas are commonly cultivars produced by breeding and selection or as sports for floral colour or size plant form or other desirable characteristics 1 Similarly the world s agricultural food crops are almost exclusively cultivars that have been selected for characters such as improved yield flavour and resistance to disease and very few wild plants are now used as food sources Trees used in forestry are also special selections grown for their enhanced quality and yield of timber Cultivars form a major part of Liberty Hyde Bailey s broader group the cultigen 2 which is defined as a plant whose origin or selection is primarily due to intentional human activity 3 A cultivar is not the same as a botanical variety 4 which is a taxonomic rank below subspecies and there are differences in the rules for creating and using the names of botanical varieties and cultivars In recent times the naming of cultivars has been complicated by the use of statutory patents for plants and recognition of plant breeders rights 5 The International Union for the Protection of New Varieties of Plants UPOV French Union internationale pour la protection des obtentions vegetales offers legal protection of plant cultivars to persons or organisations that introduce new cultivars to commerce UPOV requires that a cultivar be distinct uniform and stable To be distinct it must have characters that easily distinguish it from any other known cultivar To be uniform and stable the cultivar must retain these characters in repeated propagation The naming of cultivars is an important aspect of cultivated plant taxonomy and the correct naming of a cultivar is prescribed by the Rules and Recommendations of the International Code of Nomenclature for Cultivated Plants ICNCP commonly denominated the Cultivated Plant Code A cultivar is given a cultivar name which consists of the scientific Latin botanical name followed by a cultivar epithet The cultivar epithet is usually in a vernacular language Contents 1 Etymology 1 1 Cultigens 2 Formal definition 3 Different kinds 3 1 Clones 3 2 Seed produced 3 3 Genetically modified 4 Cultivar names 4 1 Presenting in text 5 Group names 5 1 Presenting in text 6 Legal protection of cultivars and their names 6 1 Controversial use of legal protection for cultivars 6 2 Trade designations and selling names 7 International Cultivar Registration Authorities 8 See also 9 Notes 10 References 11 Bibliography 12 External linksEtymology edit nbsp Liberty Hyde Bailey 1858 1954 coined the words cultigen in 1918 and cultivar in 1923 Main article Cultivated plant taxonomy The word cultivar originated from the need to distinguish between wild plants and those with characteristics that arose in cultivation presently denominated cultigens This distinction dates to the Greek philosopher Theophrastus 370 285 BC the Father of Botany who was keenly aware of this difference Botanical historian Alan Morton noted that Theophrastus in his Historia Plantarum Enquiry into Plants had an inkling of the limits of culturally induced phenotypic changes and of the importance of genetic constitution Historia Plantarum Book 3 2 2 and Causa Plantarum Book 1 9 3 6 The International Code of Nomenclature for algae fungi and plants uses as its starting point for modern botanical nomenclature the Latin names in Linnaeus 1707 1778 Species Plantarum tenth edition and Genera Plantarum fifth edition In Species Plantarum Linnaeus enumerated all plants known to him either directly or from his extensive reading He recognised the rank of varietas botanical variety a rank below that of species and subspecies and he indicated these varieties with letters of the Greek alphabet such as a b and l before the varietal name rather than using the abbreviation var as is the present convention Most of the varieties that Linnaeus enumerated were of garden origin rather than being wild plants 7 In time the need to distinguish between wild plants and those with variations that had been cultivated increased In the nineteenth century many garden derived plants were given horticultural names sometimes in Latin and sometimes in a vernacular language From circa the 1900s cultivated plants in Europe were recognised in the Scandinavian Germanic and Slavic literature as stamm or sorte 8 but these words could not be used internationally because by international agreement any new denominations had to be in Latin 9 In the twentieth century an improved international nomenclature was proposed for cultivated plants 10 Liberty Hyde Bailey of Cornell University in New York United States created the word cultivar in 1923 when he wrote that The cultigen is a species or its equivalent that has appeared under domestication the plant is cultigenous I now propose another name cultivar for a botanical variety or for a race subordinate to species that has originated under cultivation it is not necessarily however referable to a recognized botanical species It is essentially the equivalent of the botanical variety except in respect to its origin 2 In that essay Bailey used only the rank of species for the cultigen but it was obvious to him that many domesticated plants were more like botanical varieties than species and that realization appears to have motivated the suggestion of the new category of cultivar Bailey created the word cultivar It is generally assumed to be a blend of cultivated and variety but Bailey never explicitly stated the etymology and it has been suggested that the word is actually a blend of cultigen and variety 11 The neologism cultivar was promoted as euphonious and free from ambiguity 8 nb 2 The first Cultivated Plant Code of 1953 subsequently commended its use and by 1960 it had achieved common international acceptance 12 Cultigens edit Main article Cultigen nbsp Bread wheat Triticum aestivum is considered a cultigen and is a distinct species from other wheats according to the biological species concept Many different cultivars have been created within this cultigen Many other cultigens are not considered to be distinct species and can be denominated otherwise The words cultigen and cultivar may be confused with each other A cultigen is any plant that is deliberately selected for or altered in cultivation as opposed to an indigen the Cultivated Plant Code states that cultigens are maintained as recognisable entities solely by continued propagation 13 Cultigens can have names at any of many taxonomic ranks including those of grex species cultivar group variety form and cultivar and they may be plants that have been altered in cultivation including by genetic modification but have not been formally denominated 14 A cultigen or a component of a cultigen can be accepted as a cultivar if it is recognisable and has stable characters Therefore all cultivars are cultigens because they are cultivated but not all cultigens are cultivars because some cultigens have not been formally distinguished and named as cultivars Formal definition editMain article International Code of Nomenclature for Cultivated Plants The Cultivated Plant Code notes that the word cultivar is used in two different senses first as a classification category the cultivar is defined in Article 2 of the International Code of Nomenclature for Cultivated Plants 2009 8th edition as follows The basic category of cultivated plants whose nomenclature is governed by this Code is the cultivar 15 There are two other classification categories for cultigens the grex 16 and the group 17 The Code then defines a cultivar as a taxonomic unit within the classification category of cultivar This is the sense of cultivar that is most generally understood and which is used as a general definition A cultivar is an assemblage of plants that a has been selected for a particular character or combination of characters b is distinct uniform and stable in those characters and c when propagated by appropriate means retains those characters 18 Different kinds edit nbsp A cultivar of the orchid genus OncidiumWhich plants are chosen to be named as cultivars is simply a matter of convenience as the category was created to serve the practical needs of horticulture agriculture and forestry 19 Members of a particular cultivar are not necessarily genetically identical The Cultivated Plant Code emphasizes that different cultivated plants may be accepted as different cultivars even if they have the same genome while cultivated plants with different genomes may be regarded as the same cultivar The production of cultivars generally entails considerable human involvement although in a few cases it may be as little as simply selecting variation from plants growing in the wild whether by collecting growing tissue to propagate from or by gathering seed 20 Cultivars generally occur as ornamentals and food crops Malus Granny Smith and Malus Red Delicious are cultivars of apples propagated by cuttings or grafting Lactuca Red Sails and Lactuca Great Lakes are lettuce cultivars propagated by seeds Named cultivars of Hosta and Hemerocallis plants are cultivars produced by micropropagation or division Clones edit Main article Clone botany nbsp Leucospermum Scarlet Ribbon A cross performed in Tasmania between L glabrum and L tottumCultivars that are produced asexually are genetically identical and known as clones this includes plants propagated by division layering cuttings grafts and budding The propagating material may be taken from a particular part of the plant such as a lateral branch or from a particular phase of the life cycle such as a juvenile leaf or from aberrant growth as occurs with witch s broom Plants whose distinctive characters are derived from the presence of an intracellular organism may also form a cultivar provided the characters are reproduced reliably from generation to generation Plants of the same chimera which have mutant tissues close to normal tissue or graft chimeras which have vegetative tissue from different kinds of plants and which originate by grafting may also constitute a cultivar 21 Seed produced edit Some cultivars come true from seed retaining their distinguishing characteristics when grown from seed Such plants are termed a variety selection or strain but these are ambiguous and confusing words that are best avoided In general asexually propagated cultivars grown from seeds produce highly variable seedling plants and should not be labelled with or sold under the parent cultivar s name 22 Seed raised cultivars may be produced by uncontrolled pollination when characteristics that are distinct uniform and stable are passed from parents to progeny Some are produced as lines that are produced by repeated self fertilization or inbreeding or multilines that are made up of several closely related lines Sometimes they are F1 hybrids which are the result of a deliberate repeatable single cross between two pure lines A few F2 hybrid seed cultivars also exist such as Achillea Summer Berries Some cultivars are agamospermous plants which retain their genetic composition and characteristics under reproduction 23 Occasionally cultivars are raised from seed of a specially selected provenance for example the seed may be taken from plants that are resistant to a particular disease 24 Genetically modified edit Main article Genetic engineering Genetically modified plants with characteristics resulting from the deliberate implantation of genetic material from a different germplasm may form a cultivar However the International Code of Nomenclature for Cultivated Plants notes In practice such an assemblage is often marketed from one or more lines or multilines that have been genetically modified These lines or multilines often remain in a constant state of development which makes the naming of such an assemblage as a cultivar a futile exercise 25 However retired transgenic varieties such as the fish tomato which are no longer being developed do not run into this obstacle and can be given a cultivar name Cultivars may be selected because of a change in the ploidy level of a plant which may produce more desirable characteristics Cultivar names edit nbsp Viola Clear Crystals Apricot The specific epithet may be omitted from a cultivar name Every unique cultivar has a unique name within its denomination class which is almost always the genus Names of cultivars are regulated by the International Code of Nomenclature for Cultivated Plants and may be registered with an International Cultivar Registration Authority ICRA There are sometimes separate registration authorities for different plant types such as roses and camellias In addition cultivars may be associated with commercial marketing names referred to in the Cultivated Plant Code as trade designations see below Presenting in text edit A cultivar name consists of a botanical name of a genus species infraspecific taxon interspecific hybrid or intergeneric hybrid followed by a cultivar epithet The cultivar epithet is enclosed by single quotes 26 it should not be italicized if the botanical name is italicized 27 and each of the words within the epithet is capitalized with some permitted exceptions such as conjunctions 28 It is permissible to place a cultivar epithet after a common name provided the common name is botanically unambiguous Cultivar epithets published before 1 January 1959 were often given a Latin form and can be readily confused with the specific epithets in botanical names after that date newly coined cultivar epithets must be in a modern vernacular language to distinguish them from botanical epithets 29 For example the full cultivar name of the King Edward potato is Solanum tuberosum King Edward King Edward is the cultivar epithet which according to the Rules of the Cultivated Plant Code is bounded by single quotation marks 26 For patented or trademarked plant product lines developed from a given cultivar the commercial product name is typically indicated by the symbols TM or or is presented in capital letters with no quotation marks following the cultivar name 30 as in the following example where Bloomerang is the commercial name and Penda is the cultivar epithet Syringa Penda BLOOMERANG 31 Examples of correct text presentation Cryptomeria japonica Elegans Chamaecyparis lawsoniana Aureomarginata pre 1959 name Latin in form Chamaecyparis lawsoniana Golden Wonder post 1959 name English language Pinus densiflora Akebono post 1959 name Japanese language Apple Sundown dd Some incorrect text presentation examples Cryptomeria japonica Elegans double quotes are unacceptable Berberis thunbergii cv Crimson Pygmy this once common usage is now unacceptable as it is no longer correct to use cv in this context Berberis thunbergii Crimson Pygmy is correct Rosa cv Peace this is now incorrect for two reasons firstly the use of cv secondly Peace is a trade designation or selling name for the cultivar R Madame A Meilland and should therefore be printed in a different typeface from the rest of the name without quote marks for example RosaPeace dd Although cv has not been permitted by the International Code of Nomenclature for Cultivated Plants since the 1995 edition 32 33 it is still widely used 34 and recommended by other authorities 35 36 Group names editMain article Cultivar group Where several very similar cultivars exist they can be associated into a Group formerly Cultivar group As Group names are used with cultivar names it is necessary to understand their way of presentation Group names are presented in normal type and the first letter of each word capitalised as for cultivars but they are not placed in single quotes When used in a name the first letter of the word Group is itself capitalized 37 Presenting in text edit Brassica oleracea Capitata Group the group of cultivars including all typical cabbages Brassica oleracea Botrytis Group the group of cultivars including all typical cauliflowers Hydrangea macrophylla Groupe Hortensia in French Hydrangea macrophylla Hortensia Group in English dd Where cited with a cultivar name the group should be enclosed in parentheses as follows Hydrangea macrophylla Hortensia Group Ayesha 38 dd Legal protection of cultivars and their names editFurther information Plant breeders rights and Trademark Since the 1990s there has been an increasing use of legal protection for newly produced cultivars Plant breeders expect legal protection for the cultivars they produce According to proponents of such protections if other growers can immediately propagate and sell these cultivars as soon as they come on the market the breeder s benefit is largely lost 39 Legal protection for cultivars is obtained through the use of Plant breeders rights and plant Patents but the specific legislation and procedures needed to take advantage of this protection vary from country to country 40 Controversial use of legal protection for cultivars edit The use of legal protection for cultivars can be controversial particularly for food crops that are staples in developing countries 41 or for plants selected from the wild and propagated for sale without any additional breeding work some people consider this practice unethical 42 Trade designations and selling names edit The formal scientific name of a cultivar like Solanum tuberosum King Edward is a way of uniquely designating a particular kind of plant This scientific name is in the public domain and cannot be legally protected Plant retailers wish to maximize their share of the market and one way of doing this is to replace the cumbersome Latin scientific names on plant labels in retail outlets with appealing marketing names that are easy to use pronounce and remember Marketing names lie outside the scope of the Cultivated Plant Code which refers to them as trade designations If a retailer or wholesaler has the sole legal rights to a marketing name then that may offer a sales advantage Plants protected by plant breeders rights PBR may have a true cultivar name the recognized scientific name in the public domain and a commercial synonym an additional marketing name that is legally protected An example would be Rosa Fascination Poulmax in which Rosa is the genus Fascination is the trade designation and Poulmax is scientific cultivar name Because a name that is attractive in one language may have less appeal in another country a plant may be given different selling names from country to country Quoting the original cultivar name allows the correct identification of cultivars around the world 43 The main body coordinating plant breeders rights is the International Union for the Protection of New Varieties of Plants Union internationale pour la protection des obtentions vegetales UPOV and this organization maintains a database of new cultivars protected by PBR in all countries 44 International Cultivar Registration Authorities editMain article International Cultivar Registration Authority nbsp Dahlia Akita A cultivar selected for flower form and colourAn International Cultivar Registration Authority ICRA is a voluntary non statutory organization appointed by the Commission for Nomenclature and Cultivar Registration of the International Society of Horticultural Science ICRAs are generally formed by societies and institutions specializing in particular plant genera such as Dahlia or Rhododendron and are currently located in Europe North America China India Singapore Australia New Zealand South Africa and Puerto Rico 45 Each ICRA produces an annual report and its reappointment is considered every four years The main task is to maintain a register of the names within the group of interest and where possible this is published and placed in the public domain One major aim is to prevent the duplication of cultivar and Group epithets within a genus as well as ensuring that names are in accord with the latest edition of the Cultivated Plant Code In this way over the last 50 years or so ICRAs have contributed to the stability of cultivated plant nomenclature In recent times many ICRAs have also recorded trade designations and trademarks used in labelling plant material to avoid confusion with established names 46 New names and other relevant data are collected by and submitted to the ICRA and in most cases there is no cost The ICRA then checks each new epithet to ensure that it has not been used before and that it conforms with the Cultivated Plant Code Each ICRA also ensures that new names are formally established i e published in hard copy with a description in a dated publication They record details about the plant such as parentage the names of those concerned with its development and introduction and a basic description highlighting its distinctive characters ICRAs are not responsible for assessing the distinctiveness of the plant in question 46 Most ICRAs can be contacted electronically and many maintain web sites for an up to date listing 47 See also editLists of cultivars Plant variety law Plant LandraceNotes edit Cultivar English ˈ k ʌ l t ɪ ˌ v ɑːr ˌ v ɛer has two meanings as explained in Formal definition it is a classification category and a taxonomic unit within the category When referring to a taxon the word does not apply to an individual plant but to all plants that share the unique characteristics that define the cultivar This ignored its prior use as a transitive verb in Spanish denoting to farm to cultivate to grow or to practice Online Spanish dictionary Archived 2010 01 11 at the Wayback Machine and in Portuguese denoting to cultivate to husband to farm to plant to polish to reclaim to improve Ectaco online Portuguese dictionary Archived 2023 04 07 at the Wayback Machine References edit Stanley J Kays 3 October 2011 Cultivated vegetables of the world a multilingual onomasticon Springer pp 15 ISBN 978 90 8686 720 2 a b Bailey 1923 p 113 Spencer amp Cross 2007 p 938 Lawrence 1953 pp 19 20 HORT217 Woody Landscape Plants Department of Horticulture and Landscape Architecture Purdue University Archived from the original on 2012 01 13 Retrieved 2007 07 28 Also Archived 2 August 2012 at archive today Morton 1981 pp 38 39 Lawrence 1955 p 177 a b Lawrence 1955 p 180 Lawrence 1955 p 181 Lawrence 1955 pp 179 180 Trehane 2004 p 17 Lawrence 1960 p 1 Cultivated Plant Code Art 2 3 Brickell 2009 p 1 Spencer Cross amp Lumley 2007 p 47 Cultivated Plant Code Art 2 1 Brickell 2009 p 6 Cultivated Plant Code Art 4 Brickell 2009 p 12 Cultivated Plant Code Art 3 Brickell 2009 pp 10 12 Cultivated Plant Code Art 2 2 Brickell 2009 p 6 Cultivated Plant Code Preamble amp Principles Brickell 2009 p 19 Cultivated Plant Code Article 2 20 Brickell 2009 p 9 Cultivated Plant Code Articles 2 5 2 11 Brickell 2009 pp 6 7 Courses RHS Gardening Archived 2005 12 26 at the Wayback Machine Cultivated Plant Code Articles 2 17 2 18 Brickell 2009 pp 7 8 Cultivated Plant Code Articles 2 12 2 16 Brickell 2009 pp 7 8 Cultivated Plant Code Articles 2 19 Brickell 2009 pp 8 9 a b Cultivated Plant Code Article 14 1 Brickell 2009 p 19 Cultivated Plant Code Recommendation 8A 1 Brickell 2009 p 15 Cultivated Plant Code Article 21 3 Brickell 2009 p 25 Cultivated Plant Code Art 14 Brickell 2009 p 19 Dirr Michael Warren Keith 2019 The Tree Book Superior Selections for Landscapes Streetscapes and Gardens Timber Press Inc ISBN 978 1 60469 714 8 Syringa Penda BLOOMERANG Missouri Botanical Garden Plant Finder Archived from the original on 2021 11 30 Retrieved 2022 04 29 Naming a Cultivar Australian National Botanic Gardens Botanical Web Portal 2000 07 28 Archived from the original on 2021 03 12 Retrieved 2020 12 26 How to name a new cultivar PDF Food and Agriculture Organization 2020 06 10 Archived from the original PDF on 2020 12 26 Retrieved 2020 12 26 Chen Guang Liu Chaolei Gao Zhenyu Zhang Yu Zhang Anpeng Zhu Li Hu Jiang Ren Deyong Yu Ling Xu Guohua Qian Qian 2018 01 05 Variation in the Abundance of OsHAK1 Transcript Underlies the Differential Salinity Tolerance of an indica and a japonica Rice Cultivar Frontiers in Plant Science 8 Frontiers Media SA 2216 doi 10 3389 fpls 2017 02216 ISSN 1664 462X PMC 5760540 PMID 29354152 Horticulture 202 Laboratory 3 Texas A amp M University General Horticulture Archived from the original on 2020 07 17 Retrieved 2020 12 26 Finkle Anita 2018 04 02 What is the proper way to write a botanical name Latin name New York Botanical Garden Archived from the original on 2020 11 26 Retrieved 2020 12 26 Cultivated Plant Code Art 3 Brickell 2009 pp 10 12 Cultivated Plant Code Art 15 Brickell 2009 p 19 P Gepts 2004 Who Owns Biodiversity and How Should the Owners Be Compensated Plant Physiology 134 pp 1295 1307 BSPB Plant breeding The business and science of crop improvement Archived 2011 09 11 at the Wayback Machine British Society of Plant Breeders booklet Adi A B C Intellectual Property Rights in Biotechnology and the Fate of Poor Farmers Agriculture Social Science Research Network 13 January 2005 Archived from the original on 2021 07 15 Retrieved 2020 07 23 Who owns nature Archived 2011 09 11 at the Wayback Machine article by nurseryman and plant hunter Michael Wickenden published in The Plantsman Spencer Cross amp Lumley 2007 pp 76 81 Spencer Cross amp Lumley 2007 p 78 Cultivated Plant Code Brickell 2009 pp 62 67 83 a b See International Cultivar Registration Authorities Archived 2017 07 15 at the Wayback Machine Staff 2010 ISHS Commission Nomenclature and Cultivar Registration International Cultivar Registration Authorities ICRAs ishs org Archived from the original on 26 November 2014 Retrieved 5 March 2011 Bibliography editBailey Liberty Hyde 1923 Various cultigens and transfers in nomenclature Gentes Herbarum 1 Part 3 113 136 Brickell Chris D 2009 International Code of Nomenclature for Cultivated Plants ICNCP or Cultivated Plant Code incorporating the Rules and Recommendations for naming plants in cultivation 8th ed adopted by the International Union of Biological Sciences International Commission for the Nomenclature of Cultivated Plants PDF Scripta Horticulturae 10 International Society of Horticultural Science 1 184 ISBN 978 90 6605 662 6 Archived from the original PDF on 2011 08 13 Retrieved 2011 02 07 Lawrence George H M 1953 Cultivar Distinguished from Variety Baileya 1 19 20 Lawrence George H M 1955 The Term and Category of Cultivar Baileya 3 177 181 Lawrence George H M 1957 The Designation of Cultivar names Baileya 5 162 165 Lawrence George H M 1960 Notes on Cultivar Names Baileya 8 1 4 Morton Alan G 1981 History of Botanical Science An Account of the Development of Botany from Ancient Times to the Present Day London Academic Press ISBN 0 12 508382 3 Spencer Roger Cross Robert Lumley Peter 2007 Plant names a guide to botanical nomenclature 3rd ed Collingwood Australia CSIRO Publishing also Earthscan UK ISBN 978 0 643 09440 6 Spencer Roger D Cross Robert G 2007 The International Code of Botanical Nomenclature ICBN the International Code of Nomenclature for Cultivated Plants ICNCP and the cultigen Taxon 56 3 938 940 doi 10 2307 25065875 JSTOR 25065875 Trehane Piers 2004 50 years of the International Code of Nomenclature for Cultivated Plants Acta Horticulturae 634 17 27 doi 10 17660 ActaHortic 2004 634 1 External links edit nbsp Look up cultivar in Wiktionary the free dictionary Sale point of the Latest Edition October 2009 of The International Code of Nomenclature for Cultivated Plants International Cultivar Registration Authorities The Language of Horticulture Opinion piece by Tony Lord from The Plantsman magazine Hortivar The Food and Agriculture Organization of the United Nations Horticulture Cultivars Performance Database Retrieved from https en wikipedia org w index php title Cultivar amp oldid 1198747054, wikipedia, wiki, book, books, library,

article

, read, download, free, free download, mp3, video, mp4, 3gp, jpg, jpeg, gif, png, picture, music, song, movie, book, game, games.