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Castanea sativa

Castanea sativa, the sweet chestnut, Spanish chestnut or just chestnut, is a species of tree in the family Fagaceae, native to Southern Europe and Asia Minor, and widely cultivated throughout the temperate world. A substantial, long-lived deciduous tree, it produces an edible seed, the chestnut, which has been used in cooking since ancient times.

Sweet chestnut
Sweet chestnut fruit
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Fagales
Family: Fagaceae
Genus: Castanea
Species:
C. sativa
Binomial name
Castanea sativa
Distribution map
  Probable native range and isolated population.
  Introduced and naturalized (synanthropic) area and isolated population since Neolithic.
Growth rings of Castanea sativa

Description edit

C. sativa attains a height of 20–35 metres (66–115 feet) with a trunk often 2 m (7 ft) in diameter. Around 20 trees are recorded with diameters over 4 m (13 ft) including one 7.5 m (25 ft) in diameter at breast height. A famous ancient tree known as the Hundred Horse Chestnut in Sicily was historically recorded at 18 m (59 ft) in diameter (although it has split into multiple trunks above ground).[3] The bark often has a net-shaped (retiform) pattern with deep furrows or fissures running spirally in both directions up the trunk. The trunk is mostly straight with branching starting at low heights. The oblong-lanceolate, boldly toothed leaves are 16–28 centimetres (6–11 inches) long and 5–9 cm (2–4 in) broad.

The flowers of both sexes are borne in 10–20 cm (4–8 in) long, upright catkins, the male flowers in the upper part and female flowers in the lower part. In the Northern Hemisphere, they appear in late June to July, and by autumn, the female flowers develop into spiny cupules containing 3–7 brownish nuts that are shed during October. The female flowers eventually form a spiky sheath that deters predators from the seed.[4] The sweet chestnut is naturally self incompatible, meaning that the plant cannot pollinate itself, making cross-pollination necessary.[5] Some cultivars only produce one large seed per cupule, while others produce up to three seeds.[5] The nut itself is composed of two skins: an external, shiny brown part, and an internal skin adhering to the fruit. Inside, there is an edible, creamy-white part developed from the cotyledons.[5]

Sweet chestnut trees live to an age of 500 to 600 years.[6] In cultivation they may even grow as old as 1,000 years or more.[5]

Taxonomy edit

The tree is to be distinguished from the horse chestnut Aesculus hippocastanum, to which it is only distantly related. The horse chestnut bears similar looking seeds (conkers) in a similar seed case, which are not palatable to humans. Other common names include "Spanish chestnut",[7] or "marron" (French for "chestnut"). The generic name Castanea is the old Latin name for the plant species,[8] while the specific epithet sativa means "cultivated by humans".[9] Some selected varieties are smaller and more compact in growth yielding earlier in life with different ripening time: the Marigoule, the Marisol and the Maraval.[10]

Distribution and habitat edit

 
Millennium-old specimen in Levie, Corsica, France

The species is native to Southern Europe and Asia Minor. It is found across the Mediterranean region, from the Caspian Sea to the Atlantic Ocean. It is thought to have survived the last ice age in several refuges in Southern Europe, on the southern coast of the Black Sea with a main centre on the southern slope of the Caucasus and in the region of north-western Syria, possibly extending into Lebanon.[11]

The species is widely distributed throughout Europe, where in 2004 C. sativa was grown on 2,250,000 hectares (5,600,000 acres) of forest, of which 1,780,000 ha (4,400,000 acres) were mainly cultivated for wood and 430,000 ha (1,100,000 acres) for fruit production. In some European countries, C. sativa has only been introduced recently, for example in Slovakia or the Netherlands.[12]

The tree requires a mild climate and adequate moisture for good growth and a good nut harvest. Its year-growth (but not the rest of the tree)[13] is sensitive to late spring and early autumn frosts; it is also intolerant of lime. Under forest conditions, it will tolerate moderate shade well. It can live to more than 2,000 years of age in natural conditions, such as the Hundred Horse Chestnut near Mount Etna in eastern Sicily.

Ecology edit

The leaves provide food for some animals, including Lepidoptera such as the case-bearer moth Coleophora anatipennella and North American rose chafer Macrodactylus subspinosus.

The two major fungal pathogens of the sweet chestnut are the chestnut blight (Cryphonectria parasitica) and the ink disease caused by Phytophthora cambivora and P. cinnamomi.[14][15] In North America as well as in Southern Europe C. parasitica destroyed most of the chestnut population in the 20th century. With biological control, the population of the sweet chestnut is not threatened anymore by the chestnut blight and is regenerating.[16][17] Ink disease is infesting trees mostly in humid soils, with the mycelium invading the root and resulting in wilting of the leaf. Absence of fruit formation leads to die back of the petal. The ink disease is named after the black exudates at the base of the trunk.[15] Nowadays there are cultivars that are resistant to the ink disease. Phytophthora cambivora caused serious damage in Asia and the US, and it still continues to destroy new plantations in Europe.[15]

Another serious pest which is difficult to control is the gall wasp (Dryocosmus kuriphylus) which was recently introduced in Southern Europe, originating in Asia.[16]

Cultivation edit

Cultivation forms edit

Three different cultivation systems for the sweet chestnut can be distinguished:[18]

  • Coppicing: Mainly for wood extraction. Standard conditions yield 15 m3 wood per ha and year.
  • Selve: Fruit production from grafted trees. The trees have a short tribe and a big crown. Trees have a high density and the ground between the trees is often used as pasture.
  • High forest: Wood and fruit production. This cultivation form is less intensive with a yield of 4–12 dt/ha and replacement of trees every 50–80 years. The trees grow from seeds and build a dense canopy.

The field management is dependent on the cultivation system. While cleaning the soil from the leaves and pruning is the norm, the use of fertilizer, irrigation and pesticides is less common and reserved for more intensive cultivation.[19]

Requirements edit

The sweet chestnut tree grows well on limestone-free, deeply weathered soil.[20] The optimal pH value of the soil is between 4.5 and 6, and the tree cannot tolerate soil compaction.[18] The tolerance to wet ground and to clay-rich soils is very low.[21] It is a heat-loving tree which needs a long vegetation period. The optimal average temperature is between 8 °C and 15 °C[18] (46-59°F) and in January the temperature should preferably not be below -1 °C[20] (30°F) but it may tolerate temperatures as low as -15 °C (5°F).[18] Low temperature in autumn can damage the fruit.[20] The maximal altitude is strongly dependent on the climate. In general, the climate should be similar to viticulture.[18] Optimal precipitation is between 400 and 1,600 millimetres (16 and 63 in).[21] Before planting, seeds must be stratified at 2-3 °C so germination can start 30–40 days later. After a year, the young trees are being transplanted.[18]

Harvest edit

A tree grown from seed may take 20 years or more before it bears fruits, but a grafted cultivar such as 'Marron de Lyon' or 'Paragon' may start production within five years of being planted. Both cultivars bear fruits with a single large kernel, rather than the usual two to four smaller kernels.[13]

The fruit yield per tree is usually between 30–100 kilograms (66–220 pounds), but can get as high as 300 kilograms (660 pounds).[18] Harvest time is between middle of September and middle of November. There are three harvesting techniques:

  • By hand: The sweet chestnuts are harvested by rake or broom, with a harvest speed of 5 to 30 kilograms (11 to 66 pounds) every hour depending on the soil relief. Also, the capsule makes the harvest more complicated and even painful for the worker.[21]
  • By hand with nets: This technique is less time-consuming and protects the fruits from injuries. However, setting up the nets is work intensive.
  • Mechanical: The fruits are collected with a machine that works similarly to a vacuum cleaner. Doing so is time-saving and economical, but it is possible that some fruits get injured, and an investment is needed. Furthermore, visual sorting is not possible.

Post-harvest treatment edit

The most widespread treatment before storage is water curing, a process in which the sweet chestnuts are immersed in water for nine days.[22] The aim of this practice is to limit the main storage problems threatening the sweet chestnut: fungi development and the presence of insect worms.[22] As an alternative to water curing, hot water treatment is also commercially used.

After water treatment, the sweet chestnuts are stored in a controlled environment with high carbon dioxide concentrations. In contrast to a cold storage system, where the fruits are stored at low temperatures in untreated air, the controlled environment method avoids flesh hardening which negatively impacts the processability of the product.[22]

Cultivars edit

The ornamental cultivar C. sativa 'Albomarginata'[23] has gained the Royal Horticultural Society's Award of Garden Merit.

French origin edit

American origin edit

  • Colossal
  • Labor Day

Uses edit

Sweet chestnut (Castanea sativa), raw, unpeeled
Nutritional value per 100 g (3.5 oz)
Energy891 kJ (213 kcal)
45.54 g
Dietary fiber8.1 g
2.26 g
Saturated0.425 g
Monounsaturated0.780 g
Polyunsaturated0.894 g
2.42 g
Tryptophan0.027 g
Threonine0.086 g
Isoleucine0.095 g
Leucine0.143 g
Lysine0.143 g
Methionine0.057 g
Cystine0.077 g
Phenylalanine0.102 g
Tyrosine0.067 g
Valine0.135 g
Arginine0.173 g
Histidine0.067 g
Alanine0.161 g
Aspartic acid0.417 g
Glutamic acid0.312 g
Glycine0.124 g
Proline0.127 g
Serine0.121 g
VitaminsQuantity
%DV
Vitamin A equiv.
0%
1 μg
Thiamine (B1)
20%
0.238 mg
Riboflavin (B2)
13%
0.168 mg
Niacin (B3)
7%
1.179 mg
Pantothenic acid (B5)
10%
0.509 mg
Vitamin B6
22%
0.376 mg
Folate (B9)
16%
62 μg
Vitamin B12
0%
0.00 μg
Vitamin C
48%
43.0 mg
MineralsQuantity
%DV
Calcium
2%
27 mg
Copper
50%
0.447 mg
Iron
6%
1.01 mg
Magnesium
8%
32 mg
Manganese
41%
0.952 mg
Phosphorus
7%
93 mg
Potassium
17%
518 mg
Sodium
0%
3 mg
Zinc
5%
0.52 mg
Other constituentsQuantity
Water48.65 g

Link to USDA Database entry
Percentages estimated using US recommendations for adults,[24] except for potassium, which is estimated based on expert recommendation from the National Academies.[25]

The species is widely cultivated for its edible seeds (also called nuts) and for its wood.

Sweet chestnut has been listed as one of the 38 substances used to prepare Bach flower remedies,[26] a kind of alternative medicine promoted for its supposed effect on health. However, according to Cancer Research UK, "there is no scientific evidence to prove that flower remedies can control, cure or prevent any type of disease, including cancer".[27]

Food edit

The species' large genetic diversity and different cultivars are exploited for uses such as flour, boiling, roasting, drying, and sweets.[16]

The raw nuts, though edible, have a skin which is astringent and unpleasant to eat when still moist; after drying for a time the thin skin loses its astringency but is still better removed to reach the white fruit underneath. Cooking dry in an oven or fire normally helps remove this skin. Chestnuts are traditionally roasted in their tough brown husks after removing the spiny cupules in which they grow on the tree, the husks being peeled off and discarded and the hot chestnuts dipped in salt before eating them. Roast chestnuts are traditionally sold in streets, markets and fairs by street vendors with mobile or static braziers.

The skin of raw peeled chestnuts can be relatively easily removed by quickly blanching the nuts after scoring them by a cross slit at the tufted end. Once cooked, chestnuts acquire a sweet flavor and a floury texture similar to the sweet potato. The cooked nuts can be used for stuffing poultry, as a vegetable or in nut roasts. They can also be used in confections, puddings, desserts and cakes. They are used for flour, bread making, a cereal substitute, coffee substitute, a thickener in soups and other cookery uses, as well as for fattening stock. A sugar can be extracted from them.[13] The Corsican variety of polenta (called pulenta) is made with sweet chestnut flour. A local variety of Corsican beer also uses chestnuts. The product is sold as a sweetened paste mixed with vanilla, crème de marrons [fr], sweetened or unsweetened as chestnut purée or purée de marron, and candied chestnuts as marrons glacés.[28] In Switzerland, it is often served as Vermicelles.

Roman soldiers were given chestnut porridge before going into battle.[4]

Leaf infusions are used in respiratory diseases and are a popular remedy for whooping cough.[13] A hair shampoo can be made from infusing leaves and fruit husks.[13]

Nutritional constituents edit

The fat content is low and dominated by unsaturated fatty acids.[29][30] Sweet chestnut is a good source of starch.[31][32] The energy value per 100 g (3.5 oz) of C. sativa amounts to 891 kJ (213 kcal) (table).[33] C. sativa is characterized by high moisture content which ranges from 41% to 59%.[34] The chestnut provides a good source of copper, phosphorus, manganese and potassium (nutrition table).[29] Its sugar content ranges from 14% to 20% dry weight, depending on the cultivar.[35] Fructose is mostly responsible for the sweet taste.[35]

Effect of processing edit

Sweet chestnut is suited for human nutrition. Most sweet chestnut is consumed in processed form, which has an impact on the nutrient composition. Its naturally high concentration of organic acids is a key factor influencing the organoleptic characteristics of fruits and vegetables, namely flavor.[31] Organic acids are thought to play an important role against diseases as an antioxidant.[36][37] Heat appears to be the most influencing factor when it comes to decreasing the organic acid content. However, even after heating sweet chestnuts, antioxidant activity remains relatively high.[38] On the other hand, the consumer must consider that roasting, boiling or frying has a big impact on the nutritional profile of chestnut.[36][39] Vitamin C significantly decreases between 25 and 54% when boiled and 2–77 % when roasted. Nevertheless, roasted or boiled chestnuts may still be a solid vitamin C source, since 100 gram still represent about 20% of the recommended daily dietary intake. The sugar content is also affected by the high temperatures. Four processes are decisive for the degrading process of sugar while cooking: hydrolysis of starch to oligosaccharide and monosaccharide, decomposition of sucrose to glucose and fructose, caramelization of sugars and degradation of sugars.[40] Organic acids are also affected by high temperatures: their content decreases about 50% after frying, and 15% after boiling.[38] Responsible for the aromatic characteristics of cooked chestnuts is the effect of degradation of saccharides, proteins and lipids, the caramelization of saccharides and the maillard reaction that is reducing sugar and amino acids.[41][42]

Wood edit

 
A sweet chestnut carved by English artist Steve Field to commemorate the English Civil War, and depicting Prince Rupert hiding from the Roundheads in the well at Wollescote Hall in Stourbridge, West Midlands

This tree responds very well to coppicing, which is still practised in Britain, and produces a good crop of tannin-rich wood every 12 to 30 years, depending on intended use and local growth rate. The tannin renders the young growing wood durable and weather resistant for outdoor use, thus suitable for posts, fencing or stakes.[43] The wood is of light colour, hard and strong. It is used to make furniture, barrels (sometimes used to age balsamic vinegar), and roof beams notably in southern Europe (for example in houses of the Alpujarra, Spain, in southern France and elsewhere). The timber has a density of 560 kg per cubic meter,[44] and due to its durability in ground contact is often used for external purposes such as fencing.[44] It is also a good fuel, though not favoured for open fires as it tends to spit.[13]

Tannin is found in the following proportions on a 10% moisture basis: bark (6.8%), wood (13.4%), seed husks (10 - 13%). The leaves also contain tannin.[13]

History edit

 
The 450-year-old Spanish chestnut tree at Balmerino in Fife, Scotland

Pollen data indicates that the first spreading of C. sativa due to human activity started around 2100–2050 B.C. in Anatolia, northeastern Greece and southeastern Bulgaria.[45] Compared to other crops, the sweet chestnut was probably of relatively minor importance and distributed very heterogeneously throughout these regions.[45] The first charcoal remains of sweet chestnut only date from around 850–950 B.C., making it very difficult to infer a precise origin history. A newer but more reliable source are the literary works of Ancient Greece, with the richest being Theophrastus's Historia plantarum, written in the third century B.C.[11] Theophrastus focuses mainly on the use of sweet chestnut wood as timber and charcoal, only mentioning the use of the fruit once when commenting on the digestive difficulties it causes, but praising its nourishing quality.[11] Several Greek authors wrote about medicinal properties of the sweet chestnut, specifically as a remedy against lacerations of the lips and of the oesophagus.[11]

Similar to the introduction of grape vine and olive cultivation to the Latin world, C. sativa is thought to have been introduced during the colonisation of the Italian peninsula by the Greeks.[46] Further clues pointing to this theory can be found in the work of Pliny the Elder, who mentions only Greek colonies in connection with sweet chestnut cultivation.[11] Today's phylogenetic map of the sweet chestnut, while not fully understood, shows greater genetic similarity between Italian and western Anatolian C. sativa trees compared to eastern Anatolian specimen, reinforcing these findings.[47] Nonetheless, until the end of the pre-Christian era, the spread and use of the chestnut in Italy remained limited.[11] Carbonised sweet chestnuts were found in a Roman villa at Torre Annunziata near Naples, destroyed by the eruption of Mount Vesuvius in A.D. 79.[48]

Clues in art and literature indicate a dislike of the sweet chestnut by the Roman aristocracy.[11] Like Theophrastus, Latin authors are sceptical of the sweet chestnut as a fruit, and Pliny the Elder even goes as far as admiring how well nature has hidden this fruit of apparently so little value.[11] In the beginning of the Christian era, people probably started to realize the value and versatility of sweet chestnut wood, leading to a slow spread of the cultivation of C. sativa trees, a theory that is supported by pollen data and literary sources, as well as the increased use of sweet chestnut wood as poles and in supporting structures, wood works and pier building between A.D. 100 and 600.[11]

Increasing sweet chestnut pollen appearances in Switzerland, France, Germany and the Iberian peninsula in the first century A.D. suggests the spreading of cultivated sweet chestnut trees by the Romans.[49][50] Contrary to that notion, other scientists found no indication of the Romans spreading C. sativa before the fifth century.[51] While the husks of sweet chestnuts, dated to the third or early fourth century, have been identified from the bottom of a Roman well at Great Holts Farm, in Boreham in Essex, England; this deposit includes remains of other exotic food plants and provides no evidence that any of them originated locally. No other evidence of sweet chestnut in Roman Britain has been confirmed.[52] Indeed, no centre of sweet chestnut cultivation outside the Italian peninsula in Roman times has been detected.[11] Widespread use of chestnut in western Europe started in the early Middle Ages and flourished in the late Middle Ages.[53] In the mid-seventh-century Lombard laws, a composition of one solidi is set for felling a chestnut tree (or, also, hazel, pear or apple) belonging to another person (Edictum Rothari, No. 301, 643 AD). Since the beginning of the 20th century, due to depopulation of the countryside and the abandonment of the sweet chestnut as a staple food as well as the spread of chestnut blight and ink disease, C. sativa cultivation has dramatically decreased.[53] Nowadays, sweet chestnut production is sometimes seen at a turning point again, because the development of high-value sweet chestnut products combined with changing needs of an urban society is leading to a revival in C. sativa cultivation.[12]

See also edit

References edit

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  39. ^ Ribeiro, Bárbara; Rangel, Joana; Valentão, Patrı́cia; Andrade, Paula B.; Pereira, José Alberto; Bölke, Hanna; Seabra, Rosa M. (January 2007). "Organic acids in two Portuguese chestnut (Castanea sativa Miller) varieties". Food Chemistry. 100 (2): 504–508. doi:10.1016/j.foodchem.2005.09.073. hdl:10198/735. ISSN 0308-8146.
  40. ^ Attanasio, Gerardina; Cinquanta, Luciano; Albanese, Donatella; Matteo, Marisa Di (December 2004). "Effects of drying temperatures on physico-chemical properties of dried and rehydrated chestnuts (Castanea sativa)". Food Chemistry. 88 (4): 583–590. doi:10.1016/j.foodchem.2004.01.071. ISSN 0308-8146.
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  42. ^ Li, Qian; Shi, Xianhe; Zhao, Qiaojiao; Cui, Yahui; Ouyang, Jie; Xu, Fang (June 2016). "Effect of cooking methods on nutritional quality and volatile compounds of Chinese chestnut (Castanea mollissima Blume)". Food Chemistry. 201: 80–86. doi:10.1016/j.foodchem.2016.01.068. ISSN 0308-8146. PMID 26868551.
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  47. ^ Villani, Fiorella (1990). "Genetic structure of Castanea sativa in Turkey: evidence of a hybrid zone". Journal of Evolutionary Biology. 12: 233–244. doi:10.1046/j.1420-9101.1999.00033.x. S2CID 84817286.
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  51. ^ Di Pasquale, Gaetano (December 2010). "Reworking the idea of chestnut (Castanea sativa Mill.) cultivation in Roman times: New data from ancient Campania". Plant Biosystems. 144 (4): 865–873. doi:10.1080/11263504.2010.491974. S2CID 86527704 – via ResearchGate.
  52. ^ Jarman, R.; Hazell, Z.; Campbell, G.; Webb, J.; Chambers, F.M. (2019). "Sweet Chestnut (Castanea sativa Mill.) in Britain: Re-assessment of its Status as a Roman Archaeophyte" (PDF). Britannia. 50: 49–74. doi:10.1017/S0068113X19000011. S2CID 165935525.
  53. ^ a b Livet, R. (1987). "J. R. Pitte - Terres de Castanide: hommes et paysages du châtaignier de l'Antiquité à nos jours [compte-rendu]". Méditerranée. 60: 52, review of: Pitte, Jean-Robert (1986). Terres de Castanide: Hommes et paysages du Châtaignier de l'Antiquité à nos jours. Fayard. ISBN 9782213017235.

External links edit

  •   Data related to Castanea sativa at Wikispecies
  •   Media related to Castanea sativa at Wikimedia Commons
  • Plants for a Future database: Castanea sativa
  • Castanea sativa - distribution map, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN)

castanea, sativa, sweet, chestnut, spanish, chestnut, just, chestnut, species, tree, family, fagaceae, native, southern, europe, asia, minor, widely, cultivated, throughout, temperate, world, substantial, long, lived, deciduous, tree, produces, edible, seed, c. Castanea sativa the sweet chestnut Spanish chestnut or just chestnut is a species of tree in the family Fagaceae native to Southern Europe and Asia Minor and widely cultivated throughout the temperate world A substantial long lived deciduous tree it produces an edible seed the chestnut which has been used in cooking since ancient times Sweet chestnutSweet chestnut fruitConservation statusLeast Concern IUCN 3 1 1 Scientific classificationKingdom PlantaeClade TracheophytesClade AngiospermsClade EudicotsClade RosidsOrder FagalesFamily FagaceaeGenus CastaneaSpecies C sativaBinomial nameCastanea sativaMill 2 Distribution map Probable native range and isolated population Introduced and naturalized synanthropic area and isolated population since Neolithic Growth rings of Castanea sativa Contents 1 Description 2 Taxonomy 3 Distribution and habitat 4 Ecology 5 Cultivation 5 1 Cultivation forms 5 2 Requirements 5 3 Harvest 5 4 Post harvest treatment 5 5 Cultivars 5 6 French origin 5 7 American origin 6 Uses 6 1 Food 6 1 1 Nutritional constituents 6 1 2 Effect of processing 6 2 Wood 7 History 8 See also 9 References 10 External linksDescription editC sativa attains a height of 20 35 metres 66 115 feet with a trunk often 2 m 7 ft in diameter Around 20 trees are recorded with diameters over 4 m 13 ft including one 7 5 m 25 ft in diameter at breast height A famous ancient tree known as the Hundred Horse Chestnut in Sicily was historically recorded at 18 m 59 ft in diameter although it has split into multiple trunks above ground 3 The bark often has a net shaped retiform pattern with deep furrows or fissures running spirally in both directions up the trunk The trunk is mostly straight with branching starting at low heights The oblong lanceolate boldly toothed leaves are 16 28 centimetres 6 11 inches long and 5 9 cm 2 4 in broad The flowers of both sexes are borne in 10 20 cm 4 8 in long upright catkins the male flowers in the upper part and female flowers in the lower part In the Northern Hemisphere they appear in late June to July and by autumn the female flowers develop into spiny cupules containing 3 7 brownish nuts that are shed during October The female flowers eventually form a spiky sheath that deters predators from the seed 4 The sweet chestnut is naturally self incompatible meaning that the plant cannot pollinate itself making cross pollination necessary 5 Some cultivars only produce one large seed per cupule while others produce up to three seeds 5 The nut itself is composed of two skins an external shiny brown part and an internal skin adhering to the fruit Inside there is an edible creamy white part developed from the cotyledons 5 Sweet chestnut trees live to an age of 500 to 600 years 6 In cultivation they may even grow as old as 1 000 years or more 5 Taxonomy editThe tree is to be distinguished from the horse chestnut Aesculus hippocastanum to which it is only distantly related The horse chestnut bears similar looking seeds conkers in a similar seed case which are not palatable to humans Other common names include Spanish chestnut 7 or marron French for chestnut The generic name Castanea is the old Latin name for the plant species 8 while the specific epithet sativa means cultivated by humans 9 Some selected varieties are smaller and more compact in growth yielding earlier in life with different ripening time the Marigoule the Marisol and the Maraval 10 Distribution and habitat edit nbsp Millennium old specimen in Levie Corsica FranceThe species is native to Southern Europe and Asia Minor It is found across the Mediterranean region from the Caspian Sea to the Atlantic Ocean It is thought to have survived the last ice age in several refuges in Southern Europe on the southern coast of the Black Sea with a main centre on the southern slope of the Caucasus and in the region of north western Syria possibly extending into Lebanon 11 The species is widely distributed throughout Europe where in 2004 C sativa was grown on 2 250 000 hectares 5 600 000 acres of forest of which 1 780 000 ha 4 400 000 acres were mainly cultivated for wood and 430 000 ha 1 100 000 acres for fruit production In some European countries C sativa has only been introduced recently for example in Slovakia or the Netherlands 12 The tree requires a mild climate and adequate moisture for good growth and a good nut harvest Its year growth but not the rest of the tree 13 is sensitive to late spring and early autumn frosts it is also intolerant of lime Under forest conditions it will tolerate moderate shade well It can live to more than 2 000 years of age in natural conditions such as the Hundred Horse Chestnut near Mount Etna in eastern Sicily Ecology editSee also List of Lepidoptera that feed on chestnut trees The leaves provide food for some animals including Lepidoptera such as the case bearer moth Coleophora anatipennella and North American rose chafer Macrodactylus subspinosus The two major fungal pathogens of the sweet chestnut are the chestnut blight Cryphonectria parasitica and the ink disease caused by Phytophthora cambivora and P cinnamomi 14 15 In North America as well as in Southern Europe C parasitica destroyed most of the chestnut population in the 20th century With biological control the population of the sweet chestnut is not threatened anymore by the chestnut blight and is regenerating 16 17 Ink disease is infesting trees mostly in humid soils with the mycelium invading the root and resulting in wilting of the leaf Absence of fruit formation leads to die back of the petal The ink disease is named after the black exudates at the base of the trunk 15 Nowadays there are cultivars that are resistant to the ink disease Phytophthora cambivora caused serious damage in Asia and the US and it still continues to destroy new plantations in Europe 15 Another serious pest which is difficult to control is the gall wasp Dryocosmus kuriphylus which was recently introduced in Southern Europe originating in Asia 16 Cultivation editCultivation forms edit Three different cultivation systems for the sweet chestnut can be distinguished 18 Coppicing Mainly for wood extraction Standard conditions yield 15 m3 wood per ha and year Selve Fruit production from grafted trees The trees have a short tribe and a big crown Trees have a high density and the ground between the trees is often used as pasture High forest Wood and fruit production This cultivation form is less intensive with a yield of 4 12 dt ha and replacement of trees every 50 80 years The trees grow from seeds and build a dense canopy The field management is dependent on the cultivation system While cleaning the soil from the leaves and pruning is the norm the use of fertilizer irrigation and pesticides is less common and reserved for more intensive cultivation 19 Requirements edit The sweet chestnut tree grows well on limestone free deeply weathered soil 20 The optimal pH value of the soil is between 4 5 and 6 and the tree cannot tolerate soil compaction 18 The tolerance to wet ground and to clay rich soils is very low 21 It is a heat loving tree which needs a long vegetation period The optimal average temperature is between 8 C and 15 C 18 46 59 F and in January the temperature should preferably not be below 1 C 20 30 F but it may tolerate temperatures as low as 15 C 5 F 18 Low temperature in autumn can damage the fruit 20 The maximal altitude is strongly dependent on the climate In general the climate should be similar to viticulture 18 Optimal precipitation is between 400 and 1 600 millimetres 16 and 63 in 21 Before planting seeds must be stratified at 2 3 C so germination can start 30 40 days later After a year the young trees are being transplanted 18 Harvest edit A tree grown from seed may take 20 years or more before it bears fruits but a grafted cultivar such as Marron de Lyon or Paragon may start production within five years of being planted Both cultivars bear fruits with a single large kernel rather than the usual two to four smaller kernels 13 The fruit yield per tree is usually between 30 100 kilograms 66 220 pounds but can get as high as 300 kilograms 660 pounds 18 Harvest time is between middle of September and middle of November There are three harvesting techniques By hand The sweet chestnuts are harvested by rake or broom with a harvest speed of 5 to 30 kilograms 11 to 66 pounds every hour depending on the soil relief Also the capsule makes the harvest more complicated and even painful for the worker 21 By hand with nets This technique is less time consuming and protects the fruits from injuries However setting up the nets is work intensive Mechanical The fruits are collected with a machine that works similarly to a vacuum cleaner Doing so is time saving and economical but it is possible that some fruits get injured and an investment is needed Furthermore visual sorting is not possible Post harvest treatment edit The most widespread treatment before storage is water curing a process in which the sweet chestnuts are immersed in water for nine days 22 The aim of this practice is to limit the main storage problems threatening the sweet chestnut fungi development and the presence of insect worms 22 As an alternative to water curing hot water treatment is also commercially used After water treatment the sweet chestnuts are stored in a controlled environment with high carbon dioxide concentrations In contrast to a cold storage system where the fruits are stored at low temperatures in untreated air the controlled environment method avoids flesh hardening which negatively impacts the processability of the product 22 Cultivars edit The ornamental cultivar C sativa Albomarginata 23 has gained the Royal Horticultural Society s Award of Garden Merit French origin edit Bouche de Betizac Maraval Marigoule Marsol Precoce MigouleAmerican origin edit Colossal Labor DayUses editSweet chestnut Castanea sativa raw unpeeledNutritional value per 100 g 3 5 oz Energy891 kJ 213 kcal Carbohydrates45 54 gDietary fiber8 1 gFat2 26 gSaturated0 425 gMonounsaturated0 780 gPolyunsaturated0 894 gProtein2 42 gTryptophan0 027 gThreonine0 086 gIsoleucine0 095 gLeucine0 143 gLysine0 143 gMethionine0 057 gCystine0 077 gPhenylalanine0 102 gTyrosine0 067 gValine0 135 gArginine0 173 gHistidine0 067 gAlanine0 161 gAspartic acid0 417 gGlutamic acid0 312 gGlycine0 124 gProline0 127 gSerine0 121 gVitaminsQuantity DV Vitamin A equiv 0 1 mgThiamine B1 20 0 238 mgRiboflavin B2 13 0 168 mgNiacin B3 7 1 179 mgPantothenic acid B5 10 0 509 mgVitamin B622 0 376 mgFolate B9 16 62 mgVitamin B120 0 00 mgVitamin C48 43 0 mgMineralsQuantity DV Calcium2 27 mgCopper50 0 447 mgIron6 1 01 mgMagnesium8 32 mgManganese41 0 952 mgPhosphorus7 93 mgPotassium17 518 mgSodium0 3 mgZinc5 0 52 mgOther constituentsQuantityWater48 65 gLink to USDA Database entry Percentages estimated using US recommendations for adults 24 except for potassium which is estimated based on expert recommendation from the National Academies 25 The species is widely cultivated for its edible seeds also called nuts and for its wood Sweet chestnut has been listed as one of the 38 substances used to prepare Bach flower remedies 26 a kind of alternative medicine promoted for its supposed effect on health However according to Cancer Research UK there is no scientific evidence to prove that flower remedies can control cure or prevent any type of disease including cancer 27 Food edit Main article Chestnut The species large genetic diversity and different cultivars are exploited for uses such as flour boiling roasting drying and sweets 16 The raw nuts though edible have a skin which is astringent and unpleasant to eat when still moist after drying for a time the thin skin loses its astringency but is still better removed to reach the white fruit underneath Cooking dry in an oven or fire normally helps remove this skin Chestnuts are traditionally roasted in their tough brown husks after removing the spiny cupules in which they grow on the tree the husks being peeled off and discarded and the hot chestnuts dipped in salt before eating them Roast chestnuts are traditionally sold in streets markets and fairs by street vendors with mobile or static braziers The skin of raw peeled chestnuts can be relatively easily removed by quickly blanching the nuts after scoring them by a cross slit at the tufted end Once cooked chestnuts acquire a sweet flavor and a floury texture similar to the sweet potato The cooked nuts can be used for stuffing poultry as a vegetable or in nut roasts They can also be used in confections puddings desserts and cakes They are used for flour bread making a cereal substitute coffee substitute a thickener in soups and other cookery uses as well as for fattening stock A sugar can be extracted from them 13 The Corsican variety of polenta called pulenta is made with sweet chestnut flour A local variety of Corsican beer also uses chestnuts The product is sold as a sweetened paste mixed with vanilla creme de marrons fr sweetened or unsweetened as chestnut puree or puree de marron and candied chestnuts as marrons glaces 28 In Switzerland it is often served as Vermicelles Roman soldiers were given chestnut porridge before going into battle 4 Leaf infusions are used in respiratory diseases and are a popular remedy for whooping cough 13 A hair shampoo can be made from infusing leaves and fruit husks 13 nbsp Roasted chestnuts nbsp Street vendors in Istanbul nbsp Marrons glaces nbsp Creme de marrons fr Nutritional constituents edit The fat content is low and dominated by unsaturated fatty acids 29 30 Sweet chestnut is a good source of starch 31 32 The energy value per 100 g 3 5 oz of C sativa amounts to 891 kJ 213 kcal table 33 C sativa is characterized by high moisture content which ranges from 41 to 59 34 The chestnut provides a good source of copper phosphorus manganese and potassium nutrition table 29 Its sugar content ranges from 14 to 20 dry weight depending on the cultivar 35 Fructose is mostly responsible for the sweet taste 35 Effect of processing edit Sweet chestnut is suited for human nutrition Most sweet chestnut is consumed in processed form which has an impact on the nutrient composition Its naturally high concentration of organic acids is a key factor influencing the organoleptic characteristics of fruits and vegetables namely flavor 31 Organic acids are thought to play an important role against diseases as an antioxidant 36 37 Heat appears to be the most influencing factor when it comes to decreasing the organic acid content However even after heating sweet chestnuts antioxidant activity remains relatively high 38 On the other hand the consumer must consider that roasting boiling or frying has a big impact on the nutritional profile of chestnut 36 39 Vitamin C significantly decreases between 25 and 54 when boiled and 2 77 when roasted Nevertheless roasted or boiled chestnuts may still be a solid vitamin C source since 100 gram still represent about 20 of the recommended daily dietary intake The sugar content is also affected by the high temperatures Four processes are decisive for the degrading process of sugar while cooking hydrolysis of starch to oligosaccharide and monosaccharide decomposition of sucrose to glucose and fructose caramelization of sugars and degradation of sugars 40 Organic acids are also affected by high temperatures their content decreases about 50 after frying and 15 after boiling 38 Responsible for the aromatic characteristics of cooked chestnuts is the effect of degradation of saccharides proteins and lipids the caramelization of saccharides and the maillard reaction that is reducing sugar and amino acids 41 42 Wood edit nbsp A sweet chestnut carved by English artist Steve Field to commemorate the English Civil War and depicting Prince Rupert hiding from the Roundheads in the well at Wollescote Hall in Stourbridge West MidlandsThis tree responds very well to coppicing which is still practised in Britain and produces a good crop of tannin rich wood every 12 to 30 years depending on intended use and local growth rate The tannin renders the young growing wood durable and weather resistant for outdoor use thus suitable for posts fencing or stakes 43 The wood is of light colour hard and strong It is used to make furniture barrels sometimes used to age balsamic vinegar and roof beams notably in southern Europe for example in houses of the Alpujarra Spain in southern France and elsewhere The timber has a density of 560 kg per cubic meter 44 and due to its durability in ground contact is often used for external purposes such as fencing 44 It is also a good fuel though not favoured for open fires as it tends to spit 13 Tannin is found in the following proportions on a 10 moisture basis bark 6 8 wood 13 4 seed husks 10 13 The leaves also contain tannin 13 History edit nbsp The 450 year old Spanish chestnut tree at Balmerino in Fife ScotlandPollen data indicates that the first spreading of C sativa due to human activity started around 2100 2050 B C in Anatolia northeastern Greece and southeastern Bulgaria 45 Compared to other crops the sweet chestnut was probably of relatively minor importance and distributed very heterogeneously throughout these regions 45 The first charcoal remains of sweet chestnut only date from around 850 950 B C making it very difficult to infer a precise origin history A newer but more reliable source are the literary works of Ancient Greece with the richest being Theophrastus s Historia plantarum written in the third century B C 11 Theophrastus focuses mainly on the use of sweet chestnut wood as timber and charcoal only mentioning the use of the fruit once when commenting on the digestive difficulties it causes but praising its nourishing quality 11 Several Greek authors wrote about medicinal properties of the sweet chestnut specifically as a remedy against lacerations of the lips and of the oesophagus 11 Similar to the introduction of grape vine and olive cultivation to the Latin world C sativa is thought to have been introduced during the colonisation of the Italian peninsula by the Greeks 46 Further clues pointing to this theory can be found in the work of Pliny the Elder who mentions only Greek colonies in connection with sweet chestnut cultivation 11 Today s phylogenetic map of the sweet chestnut while not fully understood shows greater genetic similarity between Italian and western Anatolian C sativa trees compared to eastern Anatolian specimen reinforcing these findings 47 Nonetheless until the end of the pre Christian era the spread and use of the chestnut in Italy remained limited 11 Carbonised sweet chestnuts were found in a Roman villa at Torre Annunziata near Naples destroyed by the eruption of Mount Vesuvius in A D 79 48 Clues in art and literature indicate a dislike of the sweet chestnut by the Roman aristocracy 11 Like Theophrastus Latin authors are sceptical of the sweet chestnut as a fruit and Pliny the Elder even goes as far as admiring how well nature has hidden this fruit of apparently so little value 11 In the beginning of the Christian era people probably started to realize the value and versatility of sweet chestnut wood leading to a slow spread of the cultivation of C sativa trees a theory that is supported by pollen data and literary sources as well as the increased use of sweet chestnut wood as poles and in supporting structures wood works and pier building between A D 100 and 600 11 Increasing sweet chestnut pollen appearances in Switzerland France Germany and the Iberian peninsula in the first century A D suggests the spreading of cultivated sweet chestnut trees by the Romans 49 50 Contrary to that notion other scientists found no indication of the Romans spreading C sativa before the fifth century 51 While the husks of sweet chestnuts dated to the third or early fourth century have been identified from the bottom of a Roman well at Great Holts Farm in Boreham in Essex England this deposit includes remains of other exotic food plants and provides no evidence that any of them originated locally No other evidence of sweet chestnut in Roman Britain has been confirmed 52 Indeed no centre of sweet chestnut cultivation outside the Italian peninsula in Roman times has been detected 11 Widespread use of chestnut in western Europe started in the early Middle Ages and flourished in the late Middle Ages 53 In the mid seventh century Lombard laws a composition of one solidi is set for felling a chestnut tree or also hazel pear or apple belonging to another person Edictum Rothari No 301 643 AD Since the beginning of the 20th century due to depopulation of the countryside and the abandonment of the sweet chestnut as a staple food as well as the spread of chestnut blight and ink disease C sativa cultivation has dramatically decreased 53 Nowadays sweet chestnut production is sometimes seen at a turning point again because the development of high value sweet chestnut products combined with changing needs of an urban society is leading to a revival in C sativa cultivation 12 See also editAmerican chestnut Chinese chestnut Japanese chestnutReferences edit Barstow M Khela S 2018 Castanea sativa IUCN Red List of Threatened Species 2018 e T202948A67740523 doi 10 2305 IUCN UK 2018 1 RLTS T202948A67740523 en Retrieved 17 July 2023 Miller Gardeners Dictionary ed 8 no 1 1768 Flora Europaea Castanea sativa Sweet Chestnuts Castanea sativa worldwide www monumentaltrees com Retrieved 2022 05 26 a b Kew Gardens Rhizotron amp Xstrata Treetop Walkway Castanea sativa a b c d San Miguel Ayanz Jesus Rigo Danielle de Caudullo Giovanni Durrant Tracy Houston Mauri Achille 2016 European atlas of forest tree species Luxembourg ISBN 9789279367403 OCLC 958294152 a href Template Cite book html title Template Cite book cite book a CS1 maint location missing publisher link Stephan Hahn 2004 Die Esskastanien Nahrungsquelle und bedrohte Naturressource ein Beitrag zur Kenntnis der Artenvielfalt Norderstedt Books on Demand ISBN 978 3833421921 OCLC 76668313 Royal Botanic Garden Sydney blooming calendar Archived 2009 10 05 at the Wayback Machine Gledhill D 1996 The Names of Plants Cambridge University Press ISBN 0521366755 Harrison Lorraine 2012 RHS Latin for gardeners United Kingdom Mitchell Beazley p 224 ISBN 9781845337315 Sweet Chestnut Trees The Walnut Tree Co www walnuttrees co uk a b c d e f g h i j Conedera Marco July 2004 The Cultivation of Castanea sativa Mill in Europe from its origin to its diffusion on a continental scale PDF Vegetation History and Archaeobotany 13 3 161 179 doi 10 1007 s00334 004 0038 7 S2CID 55465239 a b Conedera Marco 2004 Distribution and economic potential of the sweet chestnut Castanea sativa Mill in Europe Ecologia Mediterranea 30 2 179 193 doi 10 3406 ecmed 2004 1458 S2CID 126817906 via researchgate a b c d e f g Castanea sativa Plants for a Future Anagnostakis Sandra L 1987 Chestnut Blight The Classical Problem of an Introduced Pathogen Mycologia 79 1 23 37 doi 10 2307 3807741 JSTOR 3807741 a b c Vannini Andrea 2001 Ink disease in chestnuts impact on the European chestnut Forest Snow and Landscape Research 76 345 350 via researchgate a b c Bounous G October 2005 The Chestnut A Multipurpose Resource for the New Millennium Acta Horticulturae 693 33 40 doi 10 17660 actahortic 2005 693 1 ISSN 0567 7572 Archived from the original on 2022 02 01 Retrieved 2018 12 02 Anagnostakis Sandra L 1982 01 29 Biological Control of Chestnut Blight Science 215 4532 466 471 Bibcode 1982Sci 215 466A doi 10 1126 science 215 4532 466 ISSN 0036 8075 PMID 17771259 S2CID 36933270 a b c d e f g Ludders P February 2004 Esskastanie Erwerbs Obstbau 46 via ResearchGate Mariotti Barbara January 2009 Tradizione innovazione e sostenibilita una selvicoltura per il castagno da frutto Atti del Terzo Congresso Nazionale di Selvicoltura pp 851 857 doi 10 4129 CNS2008 113 ISBN 9788887553161 via ResearchGate a href Template Cite book html title Template Cite book cite book a journal ignored help a b c Koch Henrike 31 March 2012 Projektstudie Die Edelkastanie auf Obstwiesen Eine Alternative zum Kirschanbau Project study The sweet chestnut in orchards an alternative to cherry cultivation PDF State forests of Rhineland Palatinate in German Archived from the original PDF on 27 February 2019 a b c Conedera Marco 2016 European Atlas of Forest Tree Species Chapter Castanea sativa in Europe distribution habitat usage and threats Publication Office of the European Union ISBN 9789279367403 a b c Botondi 2009 Technological Parameters of Water Curing Affect Postharvest Physiology and Storage of Marrons Castanea Sativa Mill Marrone Fiorentino Postharvest Biology and Technology 51 97 103 doi 10 1016 j postharvbio 2008 06 010 via academia edu RHS Plant Selector Castanea sativa Albomarginata Retrieved 2020 04 17 United States Food and Drug Administration 2024 Daily Value on the Nutrition and Supplement Facts Labels Retrieved 2024 03 28 National Academies of Sciences Engineering and Medicine Health and Medicine Division Food and Nutrition Board Committee to Review the Dietary Reference Intakes for Sodium and Potassium 2019 Oria Maria Harrison Meghan Stallings Virginia A eds Dietary Reference Intakes for Sodium and Potassium The National Academies Collection Reports funded by National Institutes of Health Washington DC National Academies Press US ISBN 978 0 309 48834 1 PMID 30844154 a href Template Cite book html title Template Cite book cite book a CS1 maint multiple names authors list link D S Vohra 1 June 2004 Bach Flower Remedies A Comprehensive Study B Jain Publishers p 3 ISBN 978 81 7021 271 3 Retrieved 2 September 2013 Flower remedies Cancer Research UK 2017 08 30 Archived from the original on 2013 08 10 Retrieved September 2 2013 Lori Alden 2006 The Cook s Thesaurus nut pastes a b Borges Olga Goncalves Berta de Carvalho Jose L Soeiro Correia Paula Silva Ana Paula February 2008 Nutritional quality of chestnut Castanea sativa Mill cultivars from Portugal Food Chemistry 106 3 976 984 doi 10 1016 j foodchem 2007 07 011 hdl 10348 6419 ISSN 0308 8146 De Vasconcelos Maria Do Carmo Barbosa Mendes Bennett Richard N Rosa Eduardo A S Cardoso Jorge Ventura Ferreira May 2007 Primary and Secondary Metabolite Composition of Kernels from Three Cultivars of Portuguese Chestnut Castanea sativaMill at Different Stages of Industrial Transformation Journal of Agricultural and Food Chemistry 55 9 3508 3516 doi 10 1021 jf0629080 ISSN 0021 8561 PMID 17407304 a b Griffith Vaughan J G John 2009 The new Oxford book of food plants Geissler Catherine Nicholson Barbara Dowle Elisabeth Rice Elizabeth Oxford UK Oxford University Press ISBN 9780191567742 OCLC 500808884 a href Template Cite book html title Template Cite book cite book a CS1 maint multiple names authors list link Kunsch U Scharer H Patrian B Hurter J Conedera M Sassella A Jermini M Jelmini G July 1999 Quality Assessment of Chestnut Fruits Acta Horticulturae 494 119 128 doi 10 17660 actahortic 1999 494 17 ISSN 0567 7572 Archived from the original on 2022 03 03 Retrieved 2018 12 02 Fernandes Angela Barreira Joao C M Antonio Amilcar L Bento Albino Luisa Botelho M Ferreira Isabel C F R September 2011 Assessing the effects of gamma irradiation and storage time in energetic value and in major individual nutrients of chestnuts Food and Chemical Toxicology 49 9 2429 2432 doi 10 1016 j fct 2011 06 062 ISSN 0278 6915 PMID 21740949 De La Montana Mi guelez J Mi guez Bernardez M Garci a Queijeiro J M February 2004 Composition of varieties of chestnuts from Galicia Spain Food Chemistry 84 3 401 404 doi 10 1016 s0308 8146 03 00249 8 ISSN 0308 8146 a b Neri L Dimitri G Sacchetti G February 2010 Chemical composition and antioxidant activity of cured chestnuts from three sweet chestnut Castanea sativa Mill ecotypes from Italy Journal of Food Composition and Analysis 23 1 23 29 doi 10 1016 j jfca 2009 03 002 ISSN 0889 1575 a b Silva Branca M Andrade Paula B Valentao Patricia Ferreres Federico Seabra Rosa M Ferreira Margarida A July 2004 Quince Cydonia oblonga Miller Fruit Pulp Peel and Seed and Jam Antioxidant Activity Journal of Agricultural and Food Chemistry 52 15 4705 4712 doi 10 1021 jf040057v ISSN 0021 8561 PMID 15264903 via Academia edu Blomhoff Rune Carlsen Monica H Andersen Lene Frost Jacobs David R September 2006 Health benefits of nuts potential role of antioxidants British Journal of Nutrition 96 S2 S52 S60 doi 10 1017 BJN20061864 ISSN 1475 2662 PMID 17125534 a b Barros Ana I R N A Nunes Fernando M Goncalves Berta Bennett Richard N Silva Ana Paula September 2011 Effect of cooking on total vitamin C contents and antioxidant activity of sweet chestnuts Castanea sativa Mill Food Chemistry 128 1 165 172 doi 10 1016 j foodchem 2011 03 013 hdl 10348 6571 ISSN 0308 8146 PMID 25214344 Ribeiro Barbara Rangel Joana Valentao Patri cia Andrade Paula B Pereira Jose Alberto Bolke Hanna Seabra Rosa M January 2007 Organic acids in two Portuguese chestnut Castanea sativa Miller varieties Food Chemistry 100 2 504 508 doi 10 1016 j foodchem 2005 09 073 hdl 10198 735 ISSN 0308 8146 Attanasio Gerardina Cinquanta Luciano Albanese Donatella Matteo Marisa Di December 2004 Effects of drying temperatures on physico chemical properties of dried and rehydrated chestnuts Castanea sativa Food Chemistry 88 4 583 590 doi 10 1016 j foodchem 2004 01 071 ISSN 0308 8146 Morini G Maga J A May 1995 Volatile compounds in roasted and boiled Chinese chestnuts Castanea molissima LWT Food Science and Technology 28 6 638 640 doi 10 1016 0023 6438 95 90014 4 ISSN 0023 6438 Li Qian Shi Xianhe Zhao Qiaojiao Cui Yahui Ouyang Jie Xu Fang June 2016 Effect of cooking methods on nutritional quality and volatile compounds of Chinese chestnut Castanea mollissima Blume Food Chemistry 201 80 86 doi 10 1016 j foodchem 2016 01 068 ISSN 0308 8146 PMID 26868551 Oleg Polunin Trees and Bushes of Britain and Europe Ed Paladin 1973 pp 51 188 and 195 a b Chestnut Archived 2010 02 05 at the Wayback Machine Niche Timbers Accessed 19 08 2009 a b van Zeist Willem 1991 Late Quaternary Vegetation of the Near East Wiesbaden L Reichert ISBN 978 3882265309 Forni Gaetano 1990 Gli albori dell agricoltura origine ed evoluzion fino agli etruschi ed italici Rome REDA Villani Fiorella 1990 Genetic structure of Castanea sativa in Turkey evidence of a hybrid zone Journal of Evolutionary Biology 12 233 244 doi 10 1046 j 1420 9101 1999 00033 x S2CID 84817286 Meyer Frederick 1980 Carbonized food plants of Pompeii Herculaneum and the Villa at Torre Annunziata Economic Botany 34 4 401 437 doi 10 1007 BF02858317 S2CID 22625450 van Mourik J M 1986 Pollen profiles of slope deposits in the Galician area N W Spain Amsterdam ISBN 978 9068090185 a href Template Cite book html title Template Cite book cite book a CS1 maint location missing publisher link Santos Luisa September 2000 History of vegetation during the Holocene in the Courel and Queixa Sierras Galicia northwest Iberian Peninsula Journal of Quaternary Science 15 6 621 632 Bibcode 2000JQS 15 621S doi 10 1002 1099 1417 200009 15 6 lt 621 AID JQS524 gt 3 0 CO 2 L Di Pasquale Gaetano December 2010 Reworking the idea of chestnut Castanea sativa Mill cultivation in Roman times New data from ancient Campania Plant Biosystems 144 4 865 873 doi 10 1080 11263504 2010 491974 S2CID 86527704 via ResearchGate Jarman R Hazell Z Campbell G Webb J Chambers F M 2019 Sweet Chestnut Castanea sativa Mill in Britain Re assessment of its Status as a Roman Archaeophyte PDF Britannia 50 49 74 doi 10 1017 S0068113X19000011 S2CID 165935525 a b Livet R 1987 J R Pitte Terres de Castanide hommes et paysages du chataignier de l Antiquite a nos jours compte rendu Mediterranee 60 52 review of Pitte Jean Robert 1986 Terres de Castanide Hommes et paysages du Chataignier de l Antiquite a nos jours Fayard ISBN 9782213017235 External links edit nbsp Data related to Castanea sativa at Wikispecies nbsp Media related to Castanea sativa at Wikimedia Commons U C Davis California Castanea sativa in horticulture Plants for a Future database Castanea sativa Castanea sativa distribution map genetic conservation units and related resources European Forest Genetic Resources Programme EUFORGEN Retrieved from https en wikipedia org w index php title Castanea sativa amp oldid 1212076474, wikipedia, wiki, book, books, library,

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