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Mallard

The mallard (/ˈmælɑːrd, ˈmælərd/) or wild duck (Anas platyrhynchos) is a dabbling duck that breeds throughout the temperate and subtropical Americas, Eurasia, and North Africa, and has been introduced to New Zealand, Australia, Peru, Brazil, Uruguay, Argentina, Chile, Colombia, the Falkland Islands, and South Africa. This duck belongs to the subfamily Anatinae of the waterfowl family Anatidae. Males have purple patches on their wings, while the females (hens or ducks) have mainly brown-speckled plumage. Both sexes have an area of white-bordered black or iridescent blue feathers called a speculum on their wings; males especially tend to have blue speculum feathers. The mallard is 50–65 cm (20–26 in) long, of which the body makes up around two-thirds the length. The wingspan is 81–98 cm (32–39 in) and the bill is 4.4 to 6.1 cm (1.7 to 2.4 in) long. It is often slightly heavier than most other dabbling ducks, weighing 0.7–1.6 kg (1.5–3.5 lb). Mallards live in wetlands, eat water plants and small animals, and are social animals preferring to congregate in groups or flocks of varying sizes.

Mallard
Temporal range: Late Pleistocene–present
Female (left) and male (right)
Female call
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Anseriformes
Family: Anatidae
Genus: Anas
Species:
A. platyrhynchos
Binomial name
Anas platyrhynchos
Linnaeus, 1758
Subspecies

A. p. platyrhynchos Linnaeus, 1758
A. p. domesticus Linnaeus, 1758
A. p. conboschas C. L. Brehm, 1831 (disputed)

Range of A. platyrhynchos
  Breeding
  Resident
  Passage
  Non-breeding
  Vagrant (seasonality uncertain)
  Possibly extant and introduced
  Extant and introduced (seasonality uncertain)
  Possibly extant and introduced (seasonality uncertain)
Synonyms
  • Anas boschas Linnaeus, 1758
  • Anas adunca Linnaeus, 1758

The female lays 8 to 13 creamy white to greenish-buff spotless eggs, on alternate days. Incubation takes 27 to 28 days and fledging takes 50 to 60 days. The ducklings are precocial and fully capable of swimming as soon as they hatch.

The mallard is considered to be a species of least concern by the International Union for Conservation of Nature (IUCN). Unlike many waterfowl, mallards are considered an invasive species in some regions. It is a very adaptable species, being able to live and even thrive in urban areas which may have supported more localised, sensitive species of waterfowl before development. The non-migratory mallard interbreeds with indigenous wild ducks of closely related species through genetic pollution by producing fertile offspring. Complete hybridisation of various species of wild duck gene pools could result in the extinction of many indigenous waterfowl. This species is the main ancestor of most breeds of domestic duck, and its naturally evolved wild gene pool has been genetically polluted by the domestic and feral mallard populations.

Taxonomy and evolutionary history

 
An American black duck (upper left) and a male mallard (lower right) in eclipse plumage

The mallard was one of the many bird species originally described in the 1758 10th edition of Systema Naturae by Carl Linnaeus.[2] He gave it two binomial names: Anas platyrhynchos and Anas boschas.[3] The latter was generally preferred until 1906 when Einar Lönnberg established that A. platyrhynchos had priority, as it appeared on an earlier page in the text.[4] The scientific name comes from Latin Anas, "duck" and Ancient Greek πλατυρυγχος, platyrhynchus, "broad-billed" (from πλατύς, platys, "broad" and ρυγχός, rhunkhos, "bill").[5] The genome of Anas platyrhynchos was sequenced in 2013.[6]

The name mallard originally referred to any wild drake, and it is sometimes still used this way.[7] It was derived from the Old French malart or mallart for "wild drake" although its true derivation is unclear.[8] It may be related to, or at least influenced by, an Old High German masculine proper name Madelhart, clues lying in the alternative English forms "maudelard" and "mawdelard".[9] Masle (male) has also been proposed as an influence.[10]

Mallards frequently interbreed with their closest relatives in the genus Anas, such as the American black duck, and also with species more distantly related, such as the northern pintail, leading to various hybrids that may be fully fertile.[11] This is quite unusual among such different species, and is apparently because the mallard evolved very rapidly and recently, during the Late Pleistocene.[12][failed verification] The distinct lineages of this radiation are usually kept separate due to non-overlapping ranges and behavioural cues, but have not yet reached the point where they are fully genetically incompatible.[12] Mallards and their domestic conspecifics are also fully interfertile.[13]

Genetic analysis has shown that certain mallards appear to be closer to their Indo-Pacific relatives, while others are related to their American relatives.[14] Mitochondrial DNA data for the D-loop sequence suggest that mallards may have evolved in the general area of Siberia. Mallard bones rather abruptly appear in food remains of ancient humans and other deposits of fossil bones in Europe, without a good candidate for a local predecessor species.[15] The large Ice Age palaeosubspecies that made up at least the European and West Asian populations during the Pleistocene has been named Anas platyrhynchos palaeoboschas.[16]

Mallards are differentiated in their mitochondrial DNA between North American and Eurasian populations,[17] but the nuclear genome displays a notable lack of genetic structure.[18] Haplotypes typical of American mallard relatives and eastern spot-billed ducks can be found in mallards around the Bering Sea.[19] The Aleutian Islands hold a population of mallards that appear to be evolving towards becoming a subspecies, as gene flow with other populations is very limited.[15]

Also, the paucity of morphological differences between the Old World mallards and the New World mallard demonstrates the extent to which the genome is shared among them such that birds like the Chinese spot-billed duck are highly similar to the Old World mallard, and birds such as the Hawaiian duck are highly similar to the New World mallard.[20]

The size of the mallard varies clinally; for example, birds from Greenland, though larger, have smaller bills, paler plumage, and stockier bodies than birds further south and are sometimes classified as a separate subspecies, the Greenland mallard (A. p. conboschas).[21]

Description

 
Juvenile male and female
 
Duckling

The mallard is a medium-sized waterfowl species that is often slightly heavier than most other dabbling ducks. It is 50–65 cm (20–26 in) long – of which the body makes up around two-thirds – has a wingspan of 81–98 cm (32–39 in),[22]: 505  and weighs 0.7–1.6 kg (1.5–3.5 lb).[23] Among standard measurements, the wing chord is 25.7 to 30.6 cm (10.1 to 12.0 in), the bill is 4.4 to 6.1 cm (1.7 to 2.4 in), and the tarsus is 4.1 to 4.8 cm (1.6 to 1.9 in).[24] The breeding male mallard is unmistakable, with a glossy bottle-green head and a white collar that demarcates the head from the purple-tinged brown breast, grey-brown wings, and a pale grey belly.[25] The rear of the male is black, with white-bordered dark tail feathers.[22]: 506  The bill of the male is a yellowish-orange tipped with black, with that of the female generally darker and ranging from black to mottled orange and brown.[26] The female mallard is predominantly mottled, with each individual feather showing sharp contrast from buff to very dark brown, a coloration shared by most female dabbling ducks, and has buff cheeks, eyebrow, throat, and neck, with a darker crown and eye-stripe.[22]: 506 

Both male and female mallards have distinct iridescent purple-blue speculum feathers edged with white, which are prominent in flight or at rest but temporarily shed during the annual summer moult.[27] Upon hatching, the plumage of the duckling is yellow on the underside and face (with streaks by the eyes) and black on the back (with some yellow spots) all the way to the top and back of the head.[28] Its legs and bill are also black.[28] As it nears a month in age, the duckling's plumage starts becoming drab, looking more like the female, though more streaked, and its legs lose their dark grey colouring.[22]: 506  Two months after hatching, the fledgling period has ended, and the duckling is now a juvenile.[29] The duckling is able to fly 50–60 days after hatching. Its bill soon loses its dark grey colouring, and its sex can finally be distinguished visually by three factors: 1) the bill is yellow in males, but black and orange in females;[30] 2) the breast feathers are reddish-brown in males, but brown in females;[30] and 3) in males, the centre tail feather (drake feather) is curled, but in females, the centre tail feather is straight.[30] During the final period of maturity leading up to adulthood (6–10 months of age), the plumage of female juveniles remains the same while the plumage of male juveniles gradually changes to its characteristic colours.[31] This change in plumage also applies to adult mallard males when they transition in and out of their non-breeding eclipse plumage at the beginning and the end of the summer moulting period.[31] The adulthood age for mallards is fourteen months, and the average life expectancy is three years, but they can live to twenty.[32]

Several species of duck have brown-plumaged females that can be confused with the female mallard.[33] The female gadwall (Mareca strepera) has an orange-lined bill, white belly, black and white speculum that is seen as a white square on the wings in flight, and is a smaller bird.[22]: 506  More similar to the female mallard in North America are the American black duck (A. rubripes), which is notably darker-hued in both sexes than the mallard,[34] and the mottled duck (A. fulvigula), which is somewhat darker than the female mallard, and with slightly different bare-part colouration and no white edge on the speculum.[34]

 
Owing to their highly 'malleable' genetic code, mallards can display a large amount of variation,[35] as seen here with this female, who displays faded or 'apricot' plumage.

In captivity, domestic ducks come in wild-type plumages, white, and other colours.[36] Most of these colour variants are also known in domestic mallards not bred as livestock, but kept as pets, aviary birds, etc., where they are rare but increasing in availability.[36]

A noisy species, the female has the deep quack stereotypically associated with ducks.[22]: 507  Male mallards make a sound phonetically similar to that of the female, a typical quack, but it is deeper and quieter compared to that of the female. Research conducted by Middlesex University found that the vocalisations of mallards vary depending on their environment, with urban mallards being much louder and more vociferous compared to populations of the species found in suburban and rural areas, with this being an adaptation to persistent levels of anthropogenic noise.[37][38]

When incubating a nest, or when offspring are present, females vocalise differently, making a call that sounds like a truncated version of the usual quack. This maternal vocalisation is highly attractive to their young. The repetition and frequency modulation of these quacks form the auditory basis for species identification in offspring, a process known as acoustic conspecific identification.[39] In addition, females hiss if the nest or offspring are threatened or interfered with. When taking off, the wings of a mallard produce a characteristic faint whistling noise.[40]

The mallard is a rare example of both Allen's Rule and Bergmann's Rule in birds.[41] Bergmann's Rule, which states that polar forms tend to be larger than related ones from warmer climates, has numerous examples in birds,[42] as in case of the Greenland mallard which is larger than the mallards further south.[21] Allen's Rule says that appendages like ears tend to be smaller in polar forms to minimise heat loss, and larger in tropical and desert equivalents to facilitate heat diffusion, and that the polar taxa are stockier overall.[43] Examples of this rule in birds are rare as they lack external ears, but the bill of ducks is supplied with a few blood vessels to prevent heat loss,[44] and, as in the Greenland mallard, the bill is smaller than that of birds farther south, illustrating the rule.[21]

Due to the variability of the mallard's genetic code, which gives it its vast interbreeding capability, mutations in the genes that decide plumage colour are very common and have resulted in a wide variety of hybrids, such as Brewer's duck (mallard × gadwall, Mareca strepera).[45]

Distribution and habitat

The mallard is widely distributed across the Northern and Southern Hemispheres; in North America its range extends from southern and central Alaska to Mexico, the Hawaiian Islands,[46] across the Palearctic,[47] from Iceland[48] and southern Greenland[46] and parts of Morocco (North Africa)[48] in the west, Scandinavia[48] and Britain[48] to the north, and to Siberia,[49] Japan,[50] and South Korea.[50] Also in the east, it ranges to south-eastern and south-western Australia[51] and New Zealand[52] in the Southern hemisphere.[22]: 505 [1] It is strongly migratory in the northern parts of its breeding range, and winters farther south.[53][54] For example, in North America, it winters south to the southern United States and northern Mexico,[55][56] but also regularly strays into Central America and the Caribbean between September and May.[57] A drake later named "Trevor" attracted media attention in 2018 when it turned up on the island of Niue, an atypical location for mallards.[58][59]

The mallard inhabits a wide range of habitats and climates, from the Arctic tundra to subtropical regions.[60] It is found in both fresh- and salt-water wetlands, including parks, small ponds, rivers, lakes and estuaries, as well as shallow inlets and open sea within sight of the coastline.[61] Water depths of less than 0.9 metres (3.0 ft) are preferred, with birds avoiding areas more than a few metres deep.[62] They are attracted to bodies of water with aquatic vegetation.[22]: 507 

Behaviour

 
Drake mallard performing the grunt-whistle[63]

Feeding

The mallard is omnivorous and very flexible in its choice of food.[64] Its diet may vary based on several factors, including the stage of the breeding cycle, short-term variations in available food, nutrient availability, and interspecific and intraspecific competition.[65] The majority of the mallard's diet seems to be made up of gastropods,[66] insects (including beetles, flies, lepidopterans, dragonflies, and caddisflies),[67] crustaceans,[68] other arthropods,[69] worms,[66] many varieties of seeds and plant matter,[66] and roots and tubers.[68] During the breeding season, male birds were recorded to have eaten 37.6% animal matter and 62.4% plant matter, most notably the grass Echinochloa crus-galli, and nonlaying females ate 37.0% animal matter and 63.0% plant matter, while laying females ate 71.9% animal matter and only 28.1% plant matter.[70] Plants generally make up the larger part of a bird's diet, especially during autumn migration and in the winter.[71][72]

The mallard usually feeds by dabbling for plant food or grazing; there are reports of it eating frogs.[73] However, in 2017 a flock of mallards in Romania were observed hunting small migratory birds, including grey wagtails and black redstarts, the first documented occasion they had been seen attacking and consuming large vertebrates.[74] It usually nests on a river bank, but not always near water. It is highly gregarious outside of the breeding season and forms large flocks, which are known as "sordes".[75]

Breeding

Female mallard with five ducklings

Mallards usually form pairs (in October and November in the Northern Hemisphere) until the female lays eggs at the start of the nesting season, which is around the beginning of spring.[76] At this time she is left by the male who joins up with other males to await the moulting period, which begins in June (in the Northern Hemisphere).[77][78] During the brief time before this, however, the males are still sexually potent and some of them either remain on standby to sire replacement clutches (for female mallards that have lost or abandoned their previous clutch)[79] or forcibly mate with females that appear to be isolated or unattached regardless of their species and whether or not they have a brood of ducklings.[79][80]

Nesting sites are typically on the ground, hidden in vegetation where the female's speckled plumage serves as effective camouflage,[81] but female mallards have also been known to nest in hollows in trees, boathouses, roof gardens and on balconies, sometimes resulting in hatched offspring having difficulty following their parent to water.[82]

 
Egg, Collection Museum Wiesbaden

Egg clutches number 8–13 creamy white to greenish-buff eggs free of speckles.[83][84] They measure about 58 mm (2.3 in) in length and 32 mm (1.3 in) in width.[84] The eggs are laid on alternate days, and incubation begins when the clutch is almost complete.[84] Incubation takes 27–28 days and fledging takes 50–60 days.[83][85] The ducklings are precocial and fully capable of swimming as soon as they hatch.[86] However, filial imprinting compels them to instinctively stay near the mother, not only for warmth and protection but also to learn about and remember their habitat as well as how and where to forage for food.[87] Though adoptions are known to occur, female mallards typically do not tolerate stray ducklings near their broods, and will violently attack and drive away any unfamiliar young, sometimes going as far as to kill them.[88]

When ducklings mature into flight-capable juveniles, they learn about and remember their traditional migratory routes (unless they are born and raised in captivity). In New Zealand, where mallards are naturalised, the nesting season has been found to be longer, eggs and clutches are larger and nest survival is generally greater compared with mallards in their native range.[89]

In cases where a nest or brood fails, some mallards may mate for a second time in an attempt to raise a second clutch, typically around early-to-mid summer. In addition, mallards may occasionally breed during the autumn in cases of unseasonably warm weather; one such instance of a ‘late’ clutch occurred in November 2011, in which a female successfully hatched and raised a clutch of eleven ducklings at the London Wetland Centre.[90]

During the breeding season, both male and female mallards can become aggressive, driving off competitors to themselves or their mate by charging at them.[91] Males tend to fight more than females, and attack each other by repeatedly pecking at their rival's chest, ripping out feathers and even skin on rare occasions. Female mallards are also known to carry out 'inciting displays', which encourage other ducks in the flock to begin fighting.[92] It is possible that this behaviour allows the female to evaluate the strength of potential partners.[93]

The drakes that end up being left out after the others have paired off with mating partners sometimes target an isolated female duck, even one of a different species, and proceed to chase and peck at her until she weakens, at which point the males take turns copulating with the female.[94] Lebret (1961) calls this behaviour "Attempted Rape Flight", and Stanley Cramp and K.E.L. Simmons (1977) speak of "rape-intent flights".[94] Male mallards also occasionally chase other male ducks of a different species, and even each other, in the same way.[94] In one documented case of "homosexual necrophilia", a male mallard copulated with another male he was chasing after the chased male died upon flying into a glass window.[94] This paper was awarded an Ig Nobel Prize in 2003.[95]

Mallards are opportunistically targeted by brood parasites, occasionally having eggs laid in their nests by redheads, ruddy ducks, lesser scaup, gadwalls, northern shovelers, northern pintails, cinnamon teal, common goldeneyes, and other mallards.[96] These eggs are generally accepted when they resemble the eggs of the host mallard, but the hen may attempt to eject them or even abandon the nest if parasitism occurs during egg laying.[97]

Predators and threats

 
A male mute swan (Cygnus olor) driving off a female mallard

In addition to human hunting, mallards of all ages (but especially young ones) and in all locations must contend with a wide diversity of predators including raptors and owls, mustelids, corvids, snakes, raccoons, opossums, skunks, turtles, large fish, felids, and canids, the last two including domestic ones.[98] The most prolific natural predators of adult mallards are red foxes (which most often pick off brooding females) and the faster or larger birds of prey, e.g. peregrine falcons, Aquila or Haliaeetus eagles.[99] In North America, adult mallards face no fewer than 15 species of birds of prey, from northern harriers (Circus hudsonius) and short-eared owls (Asio flammeus) (both smaller than a mallard) to huge bald (Haliaeetus leucocephalus) and golden eagles (Aquila chrysaetos), and about a dozen species of mammalian predators, not counting several more avian and mammalian predators who threaten eggs and nestlings.[97]

Mallards are also preyed upon by other waterside apex predators, such as grey herons (Ardea cinerea),[100] great blue herons (Ardea herodias) and black-crowned night herons (Nycticorax nycticorax), the European herring gull (Larus argentatus), the wels catfish (Silurus glanis), and the northern pike (Esox lucius).[101] Crows (Corvus spp.) are also known to kill ducklings and adults on occasion.[102] Also, mallards may be attacked by larger anseriformes such as swans (Cygnus spp.) and geese during the breeding season, and are frequently driven off by these birds over territorial disputes. Mute swans (Cygnus olor) have been known to attack or even kill mallards if they feel that the ducks pose a threat to their offspring.[103] Common loons (Gavia inmer) are similarly territorial and aggressive towards other birds in such disputes, and will frequently drive mallards away from their territory.[104] However, in 2019, a pair of common loons in Wisconsin were observed raising a mallard duckling for several weeks, having seemingly adopted the bird after it had been abandoned by its parents.[105]

The predation-avoidance behaviour of sleeping with one eye open, allowing one brain hemisphere to remain aware while the other half sleeps, was first demonstrated in mallards, although it is believed to be widespread among birds in general.[106]

Status and conservation

Several drakes swim in a pond

Since 1998, the mallard has been rated as a species of least concern on the IUCN Red List of Endangered Species. This is because it has a large range–more than 20,000,000 km2 (7,700,000 mi2)[107] and because its population is increasing, rather than declining by 30% over ten years or three generations and thus is not warranted a vulnerable rating. Also, the population size of the mallard is very large.[108]

Unlike many waterfowl, mallards have benefited from human alterations to the world – so much so that they are now considered an invasive species in some regions.[109] They are a common sight in urban parks, lakes, ponds, and other human-made water features in the regions they inhabit, and are often tolerated or encouraged in human habitat due to their placid nature towards humans and their beautiful and iridescent colours.[27] While most are not domesticated, mallards are so successful at coexisting in human regions that the main conservation risk they pose comes from the loss of genetic diversity among a region's traditional ducks once humans and mallards colonise an area. Mallards are very adaptable, being able to live and even thrive in urban areas which may have supported more localised, sensitive species of waterfowl before development.[110] The release of feral mallards in areas where they are not native sometimes creates problems through interbreeding with indigenous waterfowl.[109][111] These non-migratory mallards interbreed with indigenous wild ducks from local populations of closely related species through genetic pollution by producing fertile offspring.[111] Complete hybridisation of various species of wild duck gene pools could result in the extinction of many indigenous waterfowl.[111] The mallard itself is the ancestor of most domestic ducks, and its naturally evolved wild gene pool gets genetically polluted in turn by the domestic and feral populations.[112]

Over time, a continuum of hybrids ranging between almost typical examples of either species develop; the speciation process is beginning to reverse itself.[113] This has created conservation concerns for relatives of the mallard, such as the Hawaiian duck,[114][115] the New Zealand grey duck (A. s. superciliosa) subspecies of the Pacific black duck,[114][116] the American black duck,[117][118] the mottled duck,[119] Meller's duck,[120] the yellow-billed duck,[113] and the Mexican duck,[114][119] in the latter case even leading to a dispute as to whether these birds should be considered a species[121] (and thus entitled to more conservation research and funding) or included in the mallard species. Ecological changes and hunting have also led to a decline of local species; for example, the New Zealand grey duck population declined drastically due to overhunting in the mid-20th century.[116] Hybrid offspring of Hawaiian ducks seem to be less well adapted to native habitat, and using them in re-introduction projects apparently reduces success.[114][122] In summary, the problems of mallards "hybridising away" relatives is more a consequence of local ducks declining than of mallards spreading; allopatric speciation and isolating behaviour have produced today's diversity of mallard-like ducks despite the fact that, in most, if not all, of these populations, hybridisation must have occurred to some extent.[123]

Invasiveness

 
The last male Mariana mallard

Mallards are causing severe "genetic pollution" to South Africa's biodiversity by breeding with endemic ducks[124] even though the Agreement on the Conservation of African-Eurasian Migratory Waterbirdsan agreement to protect the local waterfowl populations – applies to the mallard as well as other ducks.[125] The hybrids of mallards and the yellow-billed duck are fertile, capable of producing hybrid offspring.[126] If this continues, only hybrids occur and in the long term result in the extinction of various indigenous waterfowl.[126] The mallard can crossbreed with 63 other species, posing a severe threat to indigenous waterfowl's genetic integrity.[127] Mallards and their hybrids compete with indigenous birds for resources, including nest sites, roosting sites, and food.[124]

 
Mallard x Pacific black duck hybrid, Tasmania

Availability of mallards, mallard ducklings, and fertilised mallard eggs for public sale and private ownership, either as poultry or as pets, is currently legal in the United States, except for the state of Florida, which has currently banned domestic ownership of mallards. This is to prevent hybridisation with the native mottled duck.[128]

The mallard is considered an invasive species in Australia and New Zealand,[22]: 505  where it competes with the Pacific black duck (known as the grey duck locally in New Zealand) which was over-hunted in the past. There, and elsewhere, mallards are spreading with increasing urbanisation and hybridising with local relatives.[114]

The eastern or Chinese spot-billed duck is currently introgressing into the mallard populations of the Primorsky Krai, possibly due to habitat changes from global warming.[19] The Mariana mallard was a resident allopatric population – in most respects a good species – apparently initially derived from mallard-Pacific black duck hybrids;[129] unfortunately, it became extinct in the late 20th century.[130]

The Laysan duck is an insular relative of the mallard, with a very small and fluctuating population.[131][1] Mallards sometimes arrive on its island home during migration, and can be expected to occasionally have remained and hybridised with Laysan ducks as long as these species have existed.[132] However, these hybrids are less well adapted to the peculiar ecological conditions of Laysan Island than the local ducks, and thus have lower fitness. Laysan ducks were found throughout the Hawaiian archipelago before 400 AD, after which they suffered a rapid decline during the Polynesian colonisation.[133] Now, their range includes only Laysan Island.[133] It is one of the successfully translocated birds, after having become nearly extinct in the early 20th century.[134]

Relationship with humans

 
Mallard (wild duck) resting on a poolside in San Francisco

Domestication

Mallards have often been ubiquitous in their regions among the ponds, rivers, and streams of human parks, farms, and other human-made waterways – even to the point of visiting water features in human courtyards.[135]

 
George Hetzel, mallard still life painting, 1883–1884

Mallards have had a long relationship with humans. Almost all domestic duck breeds derive from the mallard, with the exception of a few Muscovy breeds,[136] and are listed under the trinomial name A. p. domesticus. Mallards are generally monogamous while domestic ducks are mostly polygamous. Domestic ducks have no territorial behaviour and are less aggressive than mallards.[137] Domestic ducks are mostly kept for meat; their eggs are also eaten, and have a strong flavour.[137] They were first domesticated in Southeast Asia at least 4,000 years ago, during the Neolithic Age, and were also farmed by the Romans in Europe, and the Malays in Asia.[138] As the domestic duck and the mallard are the same species as each other, it is common for mallards to mate with domestic ducks and produce hybrid offspring that are fully fertile.[139] Because of this, mallards have been found to be contaminated with the genes of the domestic duck.[139]

While the keeping of domestic breeds is more popular, pure-bred mallards are sometimes kept for eggs and meat,[140] although they may require wing clipping to restrict flying.

Hunting

Mallards are one of the most common varieties of ducks hunted as a sport due to the large population size. The ideal location for hunting mallards is considered to be where the water level is somewhat shallow where the birds can be found foraging for food.[141] Hunting mallards might cause the population to decline in some places, at some times, and with some populations.[142] In certain countries, the mallard may be legally shot but is protected under national acts and policies. For example, in the United Kingdom, the mallard is protected under the Wildlife and Countryside Act 1981, which restricts certain hunting methods or taking or killing mallards.[143]

As food

Since ancient times, the mallard has been eaten as food. The wild mallard was eaten in Neolithic Greece.[144] Usually, only the breast and thigh meat is eaten.[145] It does not need to be hung before preparation, and is often braised or roasted, sometimes flavoured with bitter orange or with port.[146]

References

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For other uses see Mallard disambiguation The mallard ˈ m ae l ɑːr d ˈ m ae l er d or wild duck Anas platyrhynchos is a dabbling duck that breeds throughout the temperate and subtropical Americas Eurasia and North Africa and has been introduced to New Zealand Australia Peru Brazil Uruguay Argentina Chile Colombia the Falkland Islands and South Africa This duck belongs to the subfamily Anatinae of the waterfowl family Anatidae Males have purple patches on their wings while the females hens or ducks have mainly brown speckled plumage Both sexes have an area of white bordered black or iridescent blue feathers called a speculum on their wings males especially tend to have blue speculum feathers The mallard is 50 65 cm 20 26 in long of which the body makes up around two thirds the length The wingspan is 81 98 cm 32 39 in and the bill is 4 4 to 6 1 cm 1 7 to 2 4 in long It is often slightly heavier than most other dabbling ducks weighing 0 7 1 6 kg 1 5 3 5 lb Mallards live in wetlands eat water plants and small animals and are social animals preferring to congregate in groups or flocks of varying sizes MallardTemporal range Late Pleistocene present PreꞒ Ꞓ O S D C P T J K Pg N Female left and male right source source track track track track track track track Female callConservation statusLeast Concern IUCN 3 1 1 Scientific classificationKingdom AnimaliaPhylum ChordataClass AvesOrder AnseriformesFamily AnatidaeGenus AnasSpecies A platyrhynchosBinomial nameAnas platyrhynchosLinnaeus 1758SubspeciesA p platyrhynchos Linnaeus 1758A p domesticus Linnaeus 1758A p conboschas C L Brehm 1831 disputed Range of A platyrhynchos Breeding Resident Passage Non breeding Vagrant seasonality uncertain Possibly extant and introduced Extant and introduced seasonality uncertain Possibly extant and introduced seasonality uncertain SynonymsAnas boschas Linnaeus 1758 Anas adunca Linnaeus 1758The female lays 8 to 13 creamy white to greenish buff spotless eggs on alternate days Incubation takes 27 to 28 days and fledging takes 50 to 60 days The ducklings are precocial and fully capable of swimming as soon as they hatch The mallard is considered to be a species of least concern by the International Union for Conservation of Nature IUCN Unlike many waterfowl mallards are considered an invasive species in some regions It is a very adaptable species being able to live and even thrive in urban areas which may have supported more localised sensitive species of waterfowl before development The non migratory mallard interbreeds with indigenous wild ducks of closely related species through genetic pollution by producing fertile offspring Complete hybridisation of various species of wild duck gene pools could result in the extinction of many indigenous waterfowl This species is the main ancestor of most breeds of domestic duck and its naturally evolved wild gene pool has been genetically polluted by the domestic and feral mallard populations Contents 1 Taxonomy and evolutionary history 2 Description 3 Distribution and habitat 4 Behaviour 4 1 Feeding 4 2 Breeding 5 Predators and threats 6 Status and conservation 6 1 Invasiveness 7 Relationship with humans 7 1 Domestication 7 2 Hunting 7 3 As food 8 References 9 External linksTaxonomy and evolutionary history Edit An American black duck upper left and a male mallard lower right in eclipse plumage The mallard was one of the many bird species originally described in the 1758 10th edition of Systema Naturae by Carl Linnaeus 2 He gave it two binomial names Anas platyrhynchos and Anas boschas 3 The latter was generally preferred until 1906 when Einar Lonnberg established that A platyrhynchos had priority as it appeared on an earlier page in the text 4 The scientific name comes from Latin Anas duck and Ancient Greek platyrygxos platyrhynchus broad billed from platys platys broad and rygxos rhunkhos bill 5 The genome of Anas platyrhynchos was sequenced in 2013 6 The name mallard originally referred to any wild drake and it is sometimes still used this way 7 It was derived from the Old French malart or mallart for wild drake although its true derivation is unclear 8 It may be related to or at least influenced by an Old High German masculine proper name Madelhart clues lying in the alternative English forms maudelard and mawdelard 9 Masle male has also been proposed as an influence 10 Mallards frequently interbreed with their closest relatives in the genus Anas such as the American black duck and also with species more distantly related such as the northern pintail leading to various hybrids that may be fully fertile 11 This is quite unusual among such different species and is apparently because the mallard evolved very rapidly and recently during the Late Pleistocene 12 failed verification The distinct lineages of this radiation are usually kept separate due to non overlapping ranges and behavioural cues but have not yet reached the point where they are fully genetically incompatible 12 Mallards and their domestic conspecifics are also fully interfertile 13 Genetic analysis has shown that certain mallards appear to be closer to their Indo Pacific relatives while others are related to their American relatives 14 Mitochondrial DNA data for the D loop sequence suggest that mallards may have evolved in the general area of Siberia Mallard bones rather abruptly appear in food remains of ancient humans and other deposits of fossil bones in Europe without a good candidate for a local predecessor species 15 The large Ice Age palaeosubspecies that made up at least the European and West Asian populations during the Pleistocene has been named Anas platyrhynchos palaeoboschas 16 Mallards are differentiated in their mitochondrial DNA between North American and Eurasian populations 17 but the nuclear genome displays a notable lack of genetic structure 18 Haplotypes typical of American mallard relatives and eastern spot billed ducks can be found in mallards around the Bering Sea 19 The Aleutian Islands hold a population of mallards that appear to be evolving towards becoming a subspecies as gene flow with other populations is very limited 15 Also the paucity of morphological differences between the Old World mallards and the New World mallard demonstrates the extent to which the genome is shared among them such that birds like the Chinese spot billed duck are highly similar to the Old World mallard and birds such as the Hawaiian duck are highly similar to the New World mallard 20 The size of the mallard varies clinally for example birds from Greenland though larger have smaller bills paler plumage and stockier bodies than birds further south and are sometimes classified as a separate subspecies the Greenland mallard A p conboschas 21 Description Edit Juvenile male and female Call source source A group of mallards quacking Problems playing this file See media help Duckling The mallard is a medium sized waterfowl species that is often slightly heavier than most other dabbling ducks It is 50 65 cm 20 26 in long of which the body makes up around two thirds has a wingspan of 81 98 cm 32 39 in 22 505 and weighs 0 7 1 6 kg 1 5 3 5 lb 23 Among standard measurements the wing chord is 25 7 to 30 6 cm 10 1 to 12 0 in the bill is 4 4 to 6 1 cm 1 7 to 2 4 in and the tarsus is 4 1 to 4 8 cm 1 6 to 1 9 in 24 The breeding male mallard is unmistakable with a glossy bottle green head and a white collar that demarcates the head from the purple tinged brown breast grey brown wings and a pale grey belly 25 The rear of the male is black with white bordered dark tail feathers 22 506 The bill of the male is a yellowish orange tipped with black with that of the female generally darker and ranging from black to mottled orange and brown 26 The female mallard is predominantly mottled with each individual feather showing sharp contrast from buff to very dark brown a coloration shared by most female dabbling ducks and has buff cheeks eyebrow throat and neck with a darker crown and eye stripe 22 506 Both male and female mallards have distinct iridescent purple blue speculum feathers edged with white which are prominent in flight or at rest but temporarily shed during the annual summer moult 27 Upon hatching the plumage of the duckling is yellow on the underside and face with streaks by the eyes and black on the back with some yellow spots all the way to the top and back of the head 28 Its legs and bill are also black 28 As it nears a month in age the duckling s plumage starts becoming drab looking more like the female though more streaked and its legs lose their dark grey colouring 22 506 Two months after hatching the fledgling period has ended and the duckling is now a juvenile 29 The duckling is able to fly 50 60 days after hatching Its bill soon loses its dark grey colouring and its sex can finally be distinguished visually by three factors 1 the bill is yellow in males but black and orange in females 30 2 the breast feathers are reddish brown in males but brown in females 30 and 3 in males the centre tail feather drake feather is curled but in females the centre tail feather is straight 30 During the final period of maturity leading up to adulthood 6 10 months of age the plumage of female juveniles remains the same while the plumage of male juveniles gradually changes to its characteristic colours 31 This change in plumage also applies to adult mallard males when they transition in and out of their non breeding eclipse plumage at the beginning and the end of the summer moulting period 31 The adulthood age for mallards is fourteen months and the average life expectancy is three years but they can live to twenty 32 Several species of duck have brown plumaged females that can be confused with the female mallard 33 The female gadwall Mareca strepera has an orange lined bill white belly black and white speculum that is seen as a white square on the wings in flight and is a smaller bird 22 506 More similar to the female mallard in North America are the American black duck A rubripes which is notably darker hued in both sexes than the mallard 34 and the mottled duck A fulvigula which is somewhat darker than the female mallard and with slightly different bare part colouration and no white edge on the speculum 34 Owing to their highly malleable genetic code mallards can display a large amount of variation 35 as seen here with this female who displays faded or apricot plumage In captivity domestic ducks come in wild type plumages white and other colours 36 Most of these colour variants are also known in domestic mallards not bred as livestock but kept as pets aviary birds etc where they are rare but increasing in availability 36 A noisy species the female has the deep quack stereotypically associated with ducks 22 507 Male mallards make a sound phonetically similar to that of the female a typical quack but it is deeper and quieter compared to that of the female Research conducted by Middlesex University found that the vocalisations of mallards vary depending on their environment with urban mallards being much louder and more vociferous compared to populations of the species found in suburban and rural areas with this being an adaptation to persistent levels of anthropogenic noise 37 38 When incubating a nest or when offspring are present females vocalise differently making a call that sounds like a truncated version of the usual quack This maternal vocalisation is highly attractive to their young The repetition and frequency modulation of these quacks form the auditory basis for species identification in offspring a process known as acoustic conspecific identification 39 In addition females hiss if the nest or offspring are threatened or interfered with When taking off the wings of a mallard produce a characteristic faint whistling noise 40 The mallard is a rare example of both Allen s Rule and Bergmann s Rule in birds 41 Bergmann s Rule which states that polar forms tend to be larger than related ones from warmer climates has numerous examples in birds 42 as in case of the Greenland mallard which is larger than the mallards further south 21 Allen s Rule says that appendages like ears tend to be smaller in polar forms to minimise heat loss and larger in tropical and desert equivalents to facilitate heat diffusion and that the polar taxa are stockier overall 43 Examples of this rule in birds are rare as they lack external ears but the bill of ducks is supplied with a few blood vessels to prevent heat loss 44 and as in the Greenland mallard the bill is smaller than that of birds farther south illustrating the rule 21 Due to the variability of the mallard s genetic code which gives it its vast interbreeding capability mutations in the genes that decide plumage colour are very common and have resulted in a wide variety of hybrids such as Brewer s duck mallard gadwall Mareca strepera 45 Iridescent speculum feathers of the male Female showing pattern of the back and the coloured wing patchesDistribution and habitat EditThe mallard is widely distributed across the Northern and Southern Hemispheres in North America its range extends from southern and central Alaska to Mexico the Hawaiian Islands 46 across the Palearctic 47 from Iceland 48 and southern Greenland 46 and parts of Morocco North Africa 48 in the west Scandinavia 48 and Britain 48 to the north and to Siberia 49 Japan 50 and South Korea 50 Also in the east it ranges to south eastern and south western Australia 51 and New Zealand 52 in the Southern hemisphere 22 505 1 It is strongly migratory in the northern parts of its breeding range and winters farther south 53 54 For example in North America it winters south to the southern United States and northern Mexico 55 56 but also regularly strays into Central America and the Caribbean between September and May 57 A drake later named Trevor attracted media attention in 2018 when it turned up on the island of Niue an atypical location for mallards 58 59 The mallard inhabits a wide range of habitats and climates from the Arctic tundra to subtropical regions 60 It is found in both fresh and salt water wetlands including parks small ponds rivers lakes and estuaries as well as shallow inlets and open sea within sight of the coastline 61 Water depths of less than 0 9 metres 3 0 ft are preferred with birds avoiding areas more than a few metres deep 62 They are attracted to bodies of water with aquatic vegetation 22 507 Behaviour Edit Drake mallard performing the grunt whistle 63 Feeding Edit The mallard is omnivorous and very flexible in its choice of food 64 Its diet may vary based on several factors including the stage of the breeding cycle short term variations in available food nutrient availability and interspecific and intraspecific competition 65 The majority of the mallard s diet seems to be made up of gastropods 66 insects including beetles flies lepidopterans dragonflies and caddisflies 67 crustaceans 68 other arthropods 69 worms 66 many varieties of seeds and plant matter 66 and roots and tubers 68 During the breeding season male birds were recorded to have eaten 37 6 animal matter and 62 4 plant matter most notably the grass Echinochloa crus galli and nonlaying females ate 37 0 animal matter and 63 0 plant matter while laying females ate 71 9 animal matter and only 28 1 plant matter 70 Plants generally make up the larger part of a bird s diet especially during autumn migration and in the winter 71 72 The mallard usually feeds by dabbling for plant food or grazing there are reports of it eating frogs 73 However in 2017 a flock of mallards in Romania were observed hunting small migratory birds including grey wagtails and black redstarts the first documented occasion they had been seen attacking and consuming large vertebrates 74 It usually nests on a river bank but not always near water It is highly gregarious outside of the breeding season and forms large flocks which are known as sordes 75 Breeding Edit source source source source source source source source source source source source source source Female mallard with five ducklings Mallards usually form pairs in October and November in the Northern Hemisphere until the female lays eggs at the start of the nesting season which is around the beginning of spring 76 At this time she is left by the male who joins up with other males to await the moulting period which begins in June in the Northern Hemisphere 77 78 During the brief time before this however the males are still sexually potent and some of them either remain on standby to sire replacement clutches for female mallards that have lost or abandoned their previous clutch 79 or forcibly mate with females that appear to be isolated or unattached regardless of their species and whether or not they have a brood of ducklings 79 80 Nesting sites are typically on the ground hidden in vegetation where the female s speckled plumage serves as effective camouflage 81 but female mallards have also been known to nest in hollows in trees boathouses roof gardens and on balconies sometimes resulting in hatched offspring having difficulty following their parent to water 82 Egg Collection Museum Wiesbaden Egg clutches number 8 13 creamy white to greenish buff eggs free of speckles 83 84 They measure about 58 mm 2 3 in in length and 32 mm 1 3 in in width 84 The eggs are laid on alternate days and incubation begins when the clutch is almost complete 84 Incubation takes 27 28 days and fledging takes 50 60 days 83 85 The ducklings are precocial and fully capable of swimming as soon as they hatch 86 However filial imprinting compels them to instinctively stay near the mother not only for warmth and protection but also to learn about and remember their habitat as well as how and where to forage for food 87 Though adoptions are known to occur female mallards typically do not tolerate stray ducklings near their broods and will violently attack and drive away any unfamiliar young sometimes going as far as to kill them 88 When ducklings mature into flight capable juveniles they learn about and remember their traditional migratory routes unless they are born and raised in captivity In New Zealand where mallards are naturalised the nesting season has been found to be longer eggs and clutches are larger and nest survival is generally greater compared with mallards in their native range 89 In cases where a nest or brood fails some mallards may mate for a second time in an attempt to raise a second clutch typically around early to mid summer In addition mallards may occasionally breed during the autumn in cases of unseasonably warm weather one such instance of a late clutch occurred in November 2011 in which a female successfully hatched and raised a clutch of eleven ducklings at the London Wetland Centre 90 During the breeding season both male and female mallards can become aggressive driving off competitors to themselves or their mate by charging at them 91 Males tend to fight more than females and attack each other by repeatedly pecking at their rival s chest ripping out feathers and even skin on rare occasions Female mallards are also known to carry out inciting displays which encourage other ducks in the flock to begin fighting 92 It is possible that this behaviour allows the female to evaluate the strength of potential partners 93 The drakes that end up being left out after the others have paired off with mating partners sometimes target an isolated female duck even one of a different species and proceed to chase and peck at her until she weakens at which point the males take turns copulating with the female 94 Lebret 1961 calls this behaviour Attempted Rape Flight and Stanley Cramp and K E L Simmons 1977 speak of rape intent flights 94 Male mallards also occasionally chase other male ducks of a different species and even each other in the same way 94 In one documented case of homosexual necrophilia a male mallard copulated with another male he was chasing after the chased male died upon flying into a glass window 94 This paper was awarded an Ig Nobel Prize in 2003 95 Mallards are opportunistically targeted by brood parasites occasionally having eggs laid in their nests by redheads ruddy ducks lesser scaup gadwalls northern shovelers northern pintails cinnamon teal common goldeneyes and other mallards 96 These eggs are generally accepted when they resemble the eggs of the host mallard but the hen may attempt to eject them or even abandon the nest if parasitism occurs during egg laying 97 Predators and threats Edit A male mute swan Cygnus olor driving off a female mallard In addition to human hunting mallards of all ages but especially young ones and in all locations must contend with a wide diversity of predators including raptors and owls mustelids corvids snakes raccoons opossums skunks turtles large fish felids and canids the last two including domestic ones 98 The most prolific natural predators of adult mallards are red foxes which most often pick off brooding females and the faster or larger birds of prey e g peregrine falcons Aquila or Haliaeetus eagles 99 In North America adult mallards face no fewer than 15 species of birds of prey from northern harriers Circus hudsonius and short eared owls Asio flammeus both smaller than a mallard to huge bald Haliaeetus leucocephalus and golden eagles Aquila chrysaetos and about a dozen species of mammalian predators not counting several more avian and mammalian predators who threaten eggs and nestlings 97 Mallards are also preyed upon by other waterside apex predators such as grey herons Ardea cinerea 100 great blue herons Ardea herodias and black crowned night herons Nycticorax nycticorax the European herring gull Larus argentatus the wels catfish Silurus glanis and the northern pike Esox lucius 101 Crows Corvus spp are also known to kill ducklings and adults on occasion 102 Also mallards may be attacked by larger anseriformes such as swans Cygnus spp and geese during the breeding season and are frequently driven off by these birds over territorial disputes Mute swans Cygnus olor have been known to attack or even kill mallards if they feel that the ducks pose a threat to their offspring 103 Common loons Gavia inmer are similarly territorial and aggressive towards other birds in such disputes and will frequently drive mallards away from their territory 104 However in 2019 a pair of common loons in Wisconsin were observed raising a mallard duckling for several weeks having seemingly adopted the bird after it had been abandoned by its parents 105 The predation avoidance behaviour of sleeping with one eye open allowing one brain hemisphere to remain aware while the other half sleeps was first demonstrated in mallards although it is believed to be widespread among birds in general 106 Status and conservation Edit source source source source source source source source source source source source Several drakes swim in a pond Since 1998 the mallard has been rated as a species of least concern on the IUCN Red List of Endangered Species This is because it has a large range more than 20 000 000 km2 7 700 000 mi2 107 and because its population is increasing rather than declining by 30 over ten years or three generations and thus is not warranted a vulnerable rating Also the population size of the mallard is very large 108 Unlike many waterfowl mallards have benefited from human alterations to the world so much so that they are now considered an invasive species in some regions 109 They are a common sight in urban parks lakes ponds and other human made water features in the regions they inhabit and are often tolerated or encouraged in human habitat due to their placid nature towards humans and their beautiful and iridescent colours 27 While most are not domesticated mallards are so successful at coexisting in human regions that the main conservation risk they pose comes from the loss of genetic diversity among a region s traditional ducks once humans and mallards colonise an area Mallards are very adaptable being able to live and even thrive in urban areas which may have supported more localised sensitive species of waterfowl before development 110 The release of feral mallards in areas where they are not native sometimes creates problems through interbreeding with indigenous waterfowl 109 111 These non migratory mallards interbreed with indigenous wild ducks from local populations of closely related species through genetic pollution by producing fertile offspring 111 Complete hybridisation of various species of wild duck gene pools could result in the extinction of many indigenous waterfowl 111 The mallard itself is the ancestor of most domestic ducks and its naturally evolved wild gene pool gets genetically polluted in turn by the domestic and feral populations 112 Over time a continuum of hybrids ranging between almost typical examples of either species develop the speciation process is beginning to reverse itself 113 This has created conservation concerns for relatives of the mallard such as the Hawaiian duck 114 115 the New Zealand grey duck A s superciliosa subspecies of the Pacific black duck 114 116 the American black duck 117 118 the mottled duck 119 Meller s duck 120 the yellow billed duck 113 and the Mexican duck 114 119 in the latter case even leading to a dispute as to whether these birds should be considered a species 121 and thus entitled to more conservation research and funding or included in the mallard species Ecological changes and hunting have also led to a decline of local species for example the New Zealand grey duck population declined drastically due to overhunting in the mid 20th century 116 Hybrid offspring of Hawaiian ducks seem to be less well adapted to native habitat and using them in re introduction projects apparently reduces success 114 122 In summary the problems of mallards hybridising away relatives is more a consequence of local ducks declining than of mallards spreading allopatric speciation and isolating behaviour have produced today s diversity of mallard like ducks despite the fact that in most if not all of these populations hybridisation must have occurred to some extent 123 Invasiveness Edit The last male Mariana mallardMallards are causing severe genetic pollution to South Africa s biodiversity by breeding with endemic ducks 124 even though the Agreement on the Conservation of African Eurasian Migratory Waterbirds an agreement to protect the local waterfowl populations applies to the mallard as well as other ducks 125 The hybrids of mallards and the yellow billed duck are fertile capable of producing hybrid offspring 126 If this continues only hybrids occur and in the long term result in the extinction of various indigenous waterfowl 126 The mallard can crossbreed with 63 other species posing a severe threat to indigenous waterfowl s genetic integrity 127 Mallards and their hybrids compete with indigenous birds for resources including nest sites roosting sites and food 124 Mallard x Pacific black duck hybrid Tasmania Availability of mallards mallard ducklings and fertilised mallard eggs for public sale and private ownership either as poultry or as pets is currently legal in the United States except for the state of Florida which has currently banned domestic ownership of mallards This is to prevent hybridisation with the native mottled duck 128 The mallard is considered an invasive species in Australia and New Zealand 22 505 where it competes with the Pacific black duck known as the grey duck locally in New Zealand which was over hunted in the past There and elsewhere mallards are spreading with increasing urbanisation and hybridising with local relatives 114 The eastern or Chinese spot billed duck is currently introgressing into the mallard populations of the Primorsky Krai possibly due to habitat changes from global warming 19 The Mariana mallard was a resident allopatric population in most respects a good species apparently initially derived from mallard Pacific black duck hybrids 129 unfortunately it became extinct in the late 20th century 130 The Laysan duck is an insular relative of the mallard with a very small and fluctuating population 131 1 Mallards sometimes arrive on its island home during migration and can be expected to occasionally have remained and hybridised with Laysan ducks as long as these species have existed 132 However these hybrids are less well adapted to the peculiar ecological conditions of Laysan Island than the local ducks and thus have lower fitness Laysan ducks were found throughout the Hawaiian archipelago before 400 AD after which they suffered a rapid decline during the Polynesian colonisation 133 Now their range includes only Laysan Island 133 It is one of the successfully translocated birds after having become nearly extinct in the early 20th century 134 Relationship with humans EditFurther information Domestic duck Mallard wild duck resting on a poolside in San Francisco Domestication Edit Mallards have often been ubiquitous in their regions among the ponds rivers and streams of human parks farms and other human made waterways even to the point of visiting water features in human courtyards 135 George Hetzel mallard still life painting 1883 1884 Mallards have had a long relationship with humans Almost all domestic duck breeds derive from the mallard with the exception of a few Muscovy breeds 136 and are listed under the trinomial name A p domesticus Mallards are generally monogamous while domestic ducks are mostly polygamous Domestic ducks have no territorial behaviour and are less aggressive than mallards 137 Domestic ducks are mostly kept for meat their eggs are also eaten and have a strong flavour 137 They were first domesticated in Southeast Asia at least 4 000 years ago during the Neolithic Age and were also farmed by the Romans in Europe and the Malays in Asia 138 As the domestic duck and the mallard are the same species as each other it is common for mallards to mate with domestic ducks and produce hybrid offspring that are fully fertile 139 Because of this mallards have been found to be contaminated with the genes of the domestic duck 139 While the keeping of domestic breeds is more popular pure bred mallards are sometimes kept for eggs and meat 140 although they may require wing clipping to restrict flying Hunting Edit Mallards are one of the most common varieties of ducks hunted as a sport due to the large population size The ideal location for hunting mallards is considered to be where the water level is somewhat shallow where the birds can be found foraging for food 141 Hunting mallards might cause the population to decline in some places at some times and with some populations 142 In certain countries the mallard may be legally shot but is protected under national acts and policies For example in the United Kingdom the mallard is protected under the Wildlife and Countryside Act 1981 which restricts certain hunting methods or taking or killing mallards 143 As food Edit Since ancient times the mallard has been eaten as food The wild mallard was eaten in Neolithic Greece 144 Usually only the breast and thigh meat is eaten 145 It does not need to be hung before preparation and is often braised or roasted sometimes flavoured with bitter orange or with port 146 References Edit a b c BirdLife International 2019 amended version of 2017 assessment Anas platyrhynchos IUCN Red List of Threatened Species 2019 e T22680186A155457360 doi 10 2305 IUCN UK 2019 3 RLTS T22680186A155457360 en Retrieved 23 February 2022 Linnaeus Carl 1758 Systema naturae per regna tria naturae secundum classes ordines genera species cum characteribus differentiis synonymis locis Tomus I Editio decima reformata in Latin Laurentius Salvius p 125 Jobling James A 2010 Helm Dictionary of Scientific Bird Names Bloomsbury Publishing ISBN 9781408133262 Johnsgard Paul A 1961 Anas boschas platyrhynchos Linnaeus Evolutionary relationships among the North American mallards The Auk 78 1 3 43 11 12 doi 10 2307 4082232 JSTOR 4082232 S2CID 41605830 Jobling James A 2010 The Helm Dictionary of Scientific Bird Names Christopher Helm pp 46 309 ISBN 978 1 4081 2501 4 Huang Y Li Y Burt D W Chen H Zhang Y Qian Wubin Kim Heebal Gan Shangquan Zhao Yiqiang Li Jianwen Yi Kang Feng Huapeng Zhu Pengyang Li Bo Liu Qiuyue Fairley Suan Magor Katharine E Du Zhenlin Hu Xiaoxiang Goodman Laurie Tafer Hakim Vignal Alain Lee Taeheon Kim Kyu Won Sheng Zheya An Yang Searle Steve Herrero Javier Groenen Martien A M et al 2013 The duck genome and transcriptome provide insight into an avian influenza virus reservoir species Nature Genetics 45 7 776 783 doi 10 1038 ng 2657 PMC 4003391 PMID 23749191 Magnus PD 2012 Scientific Enquiry and Natural Kinds from Planets to Mallards Palgrave Macmillan ISBN 9781137271259 Buckingham James Silk Sterling John Maurice Frederick Denison Stebbing Henry Dilke Charles Wentworth Hervey Thomas Kibble Dixon William Hepworth Maccoll Norman Rendall Vernon Horace 1904 The Athenaeum A Journal of Literature Science the Fine Arts Music and the Drama J Francis mallard Oxford English Dictionary 2nd ed Oxford University Press 1989 Wedgwood Hensleigh 1862 Dictionary of English Etymology Trubner and Company Phillips John C 1915 Experimental studies of hybridization among ducks and pheasants Journal of Experimental Zoology 18 1 69 112 doi 10 1002 jez 1400180103 a b Steadman David W 2005 Late Pleistocene Birds from Kingston Saltpeter Cave Southern Appalachian Mountains Georgia The Bulletin of the Florida Museum of Natural History 231 248 Anonymous 1937 Proceedings of the Royal Society of Edinburgh Oliver amp Boyd p 210 Johnson Kevin P Sorenson M D 1999 Phylogeny and biogeography of dabbling ducks genus Anas a comparison of molecular and morphological evidence PDF The Auk 116 3 792 805 doi 10 2307 4089339 JSTOR 4089339 a b Kulikova Irina V Drovetski S V Gibson D D Harrigan R J Rohwer S Sorenson Michael D Winker K Zhuravlev Yury N McCracken Kevin G 2005 Phylogeography of the Mallard Anas platyrhynchos hybridization dispersal and lineage sorting contribute to complex geographic structure The Auk 122 3 949 965 doi 10 1642 0004 8038 2005 122 0949 POTMAP 2 0 CO 2 S2CID 85668932 Erratum The Auk 122 4 1309 doi 10 1642 0004 8038 2005 122 1309 POTMAP2 0 CO 2 Delacour Jean 1964 The Waterfowl of the World Country Life Kraus R H S Zeddeman A van Hooft P Sartakov D Soloviev S A Ydenberg Ronald C Prins Herbert H T 2011 Evolution and connectivity in the world wide migration system of the mallard Inferences from mitochondrial DNA BMC Genetics 12 99 99 doi 10 1186 1471 2156 12 99 PMC 3258206 PMID 22093799 Kraus R H S van Hooft P Megens H J Tsvey A Fokin S Y Ydenberg Ronald C Prins Herbert H T 2013 Global lack of flyway structure in a cosmopolitan bird revealed by a genome wide survey of single nucleotide polymorphisms Molecular Ecology published January 2013 22 1 41 55 doi 10 1111 mec 12098 PMID 23110616 S2CID 11190535 a b Kulikova Irina V Zhuravlev Yury N McCracken Kevin G 2004 Asymmetric hybridization and sex biased gene flow between Eastern Spot billed Ducks Anas zonorhyncha and Mallards A platyrhynchos in the Russian Far East The Auk 121 3 930 949 doi 10 1642 0004 8038 2004 121 0930 AHASGF 2 0 CO 2 S2CID 17470882 Lavretsky Philip McCracken Kevin G Peters Jeffrey L January 2014 Phylogenetics of a recent radiation in the mallards and allies Aves Anas inferences from a genomic transect and the multispecies coalescent Molecular Phylogenetics and Evolution 70 402 411 doi 10 1016 j ympev 2013 08 008 ISSN 1095 9513 PMID 23994490 a b c Ogilvie M A Young Steve 2002 Wildfowl of the World New Holland Publishers ISBN 9781843303282 a b c d e f g h i Cramp Stanley ed 1977 Handbook of the Birds of Europe the Middle East and North Africa the Birds of the Western Palearctic Vol 1 Ostrich to Ducks Oxford University Press ISBN 9780198573586 Dunning John B Jr ed 1992 CRC 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links Edit Wikimedia Commons has media related to Anas platyrhynchos Wikispecies has information related to Anas platyrhynchos Mallard media Internet Bird Collection Mallard photo gallery at VIREO Drexel University Retrieved from https en wikipedia org w index php title Mallard amp oldid 1136421512, wikipedia, wiki, book, books, library,

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