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Wikipedia

Pig

January 01, 1970

For other uses, see Pig (disambiguation).
"Swine" redirects here. For other uses, see Swine (disambiguation).

The pig (Sus domesticus), also called swine (pl.: swine) or hog, is an omnivorous, domesticated, even-toed, hoofed mammal. It is named the domestic pig when distinguishing it from other members of the genus Sus. It is considered a subspecies of Sus scrofa (the wild boar or Eurasian boar) by some authorities, but as a distinct species by others. Pigs were domesticated in the Neolithic, both in East Asia and in the Near East. When domesticated pigs arrived in Europe, they extensively interbred with wild boar but retained their domesticated features.

Pig
Domestic pigs
Domesticated
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Suidae
Genus: Sus
Species:
S. domesticus
Binomial name
Sus domesticus
Erxleben, 1777
Synonyms[1]

Pigs are farmed primarily for meat, called pork. The animal's skin or hide is used for leather. China is the world's largest pork producer, followed by the European Union and then the United States. Around 1.5 billion pigs are raised each year, producing some 120 million tonnes of meat, often cured as bacon. Some are kept as pets.

Pigs have featured in human culture since Neolithic times, appearing in art and literature for children and adults, and celebrated in cities such as Bologna for their meat products.

Description

The pig has a large head, with a long snout strengthened by a special prenasal bone and a disk of cartilage at the tip.[2] The snout is used to dig into the soil to find food and is an acute sense organ. The dental formula of adult pigs is 3.1.4.33.1.4.3, giving a total of 44 teeth. The rear teeth are adapted for crushing. In males, the canine teeth can form tusks, which grow continuously and are sharpened by grinding against each other.[2] There are four hoofed toes on each foot; the two larger central toes bear most of the weight, while the outer two are also used in soft ground.[3] Most pigs have rather sparsely bristled hair on their skin, though there are some woolly-coated breeds such as the Mangalitsa.[4] Adult pigs generally weigh between 140 and 300 kg (310 and 660 lb), though some breeds can exceed this range. Exceptionally, a pig called Big Bill weighed 1,157 kg (2,551 lb) and had a shoulder height of 1.5 m (4.9 ft).[5]

Pigs possess both apocrine and eccrine sweat glands, although the latter are limited to the snout.[6] Pigs, like other "hairless" mammals such as elephants, do not use thermal sweat glands in cooling.[7] Pigs are less able than many other mammals to dissipate heat from wet mucous membranes in the mouth by panting. Their thermoneutral zone is 16–22 °C (61–72 °F).[8] At higher temperatures, pigs lose heat by wallowing in mud or water via evaporative cooling, although it has been suggested that wallowing may serve other functions, such as protection from sunburn, ecto-parasite control, and scent-marking.[9] Pigs are among four mammalian species with mutations in the nicotinic acetylcholine receptor that protect against snake venom. Mongooses, honey badgers, hedgehogs, and pigs all have different modifications to the receptor pocket which prevents α-neurotoxin from binding.[10] Pigs have small lungs for their body size, and are thus more susceptible than other domesticated animals to fatal bronchitis and pneumonia.[11] The genome of the pig has been sequenced; it contains about 22,342 protein-coding genes.[12][13][14]

  •  
    Skeleton
  •  
    Skull
  •  
    Bones of the foot

Evolution

Phylogeny

Domestic pigs are related to other pig species as shown in the cladogram, based on phylogenetic analysis using mitochondrial DNA.[15]

Suidae

Babirusa  

Indonesia
Africa

Visayan warty pig  

Philippine warty pig  

Wild boar

 

Domestic pig

East Asian pigs  

European pigs  

Eurasia
Southeast Asia

Taxonomy

The pig is most often considered to be a subspecies of the wild boar, which was given the name Sus scrofa by Carl Linnaeus in 1758; following from this, the formal name of the pig is Sus scrofa domesticus.[16][17] However, in 1777, Johann Christian Polycarp Erxleben classified the pig as a separate species from the wild boar. He gave it the name Sus domesticus, still used by some taxonomists.[18] The American Society of Mammalogists considers it a separate species.[19]

Domestication in the Neolithic

Further information: Domestication
 
The initial emergence of wild pigs, followed by the genetic divergence between boars and pigs and the domestication of pigs [20]

Archaeological evidence shows that pigs were domesticated from wild boar in the Near East in or around the Tigris Basin,[21] being managed in a semi-wild state much as they are managed by some modern New Guineans.[22] There were pigs in Cyprus more than 11,400 years ago, introduced from the mainland, implying domestication in the adjacent mainland by then.[23] Pigs were separately domesticated in China, starting some 8,000 years ago.[24][25][26] In the Near East, pig husbandry spread for the next few millennia. It reduced gradually during the Bronze Age, as rural populations instead focused on commodity-producing livestock, but it was sustained in cities.[27]

Domestication did not involve reproductive isolation with population bottlenecks. Western Asian pigs were introduced into Europe, where they crossed with wild boar. There appears to have been interbreeding with a now extinct ghost population of wild pigs during the Pleistocene. The genomes of domestic pigs show strong selection for genes affecting behavior and morphology. Human selection for domestic traits likely counteracted the homogenizing effect of gene flow from wild boars and created domestication islands in the genome.[28][29] Pigs arrived in Europe from the Near East at least 8,500 years ago. Over the next 3,000 years they interbred with European wild boar until their genome showed less than 5% Near Eastern ancestry, yet retained their domesticated features.[30]

DNA evidence from subfossil remains of teeth and jawbones of Neolithic pigs shows that the first domestic pigs in Europe were brought from the Near East. This stimulated the domestication of local European wild boar, resulting in a third domestication event with the Near Eastern genes dying out in European pig stock. More recently there have been complex exchanges, with European domesticated lines being exported, in turn, to the ancient Near East.[31][32] Historical records indicate that Asian pigs were again introduced into Europe during the 18th and early 19th centuries.[25]

History

Columbian Exchange

Among the animals that the Spanish introduced to the Chiloé Archipelago in the 16th century Columbian Exchange, pigs were the most successful in adapting to local conditions. The pigs benefited from abundant shellfish and algae exposed by the large tides of the archipelago.[33] Pigs were brought to southeastern North America from Europe by de Soto and other early Spanish explorers. Escaped pigs became feral.[34]

Feral pigs

Main article: Feral pig
 
A family of feral pigs

Pigs have escaped from farms and gone feral in many parts of the world. Feral pigs in the southeastern United States have migrated north to the Midwest, where many state agencies have programs to remove them.[35][36][37] Feral pigs in New Zealand and northern Queensland have caused substantial environmental damage.[38][39] Feral hybrids of the European wild boar with the domestic pig are disruptive to both environment and agriculture, as they destroy crops, spread animal diseases including Foot-and-mouth disease, and consume wildlife such as juvenile seabirds and young tortoises.[40] Feral pig damage is especially an issue in southeastern South America.[41][42]

Reproduction

Physiology

Female pigs reach sexual maturity at 3–12 months of age and come into estrus every 18–24 days if they are not successfully bred. The variation in ovulation rate can be attributed to intrinsic factors such as age and genotype, as well as extrinsic factors like nutrition, environment, and the supplementation of exogenous hormones. The gestation period averages 112–120 days.[43]

 
Piglets keeping warm together

Estrus lasts two to three days, and the female's displayed receptiveness to mate is known as standing heat. Standing heat is a reflexive response that is stimulated when the female is in contact with the saliva of a sexually mature boar. Androstenol is one of the pheromones produced in the submaxillary salivary glands of boars that trigger the female's response.[44] The female cervix contains a series of five interdigitating pads, or folds, that hold the boar's corkscrew-shaped penis during copulation.[45] Females have bicornuate uteruses and two conceptuses must be present in both uterine horns to enable pregnancy to proceed.[46] The mother's body recognises that it is pregnant on days 11 to 12 of pregnancy, and is marked by the corpus luteum's producing the sex hormone progesterone.[47] To sustain the pregnancy, the embryo signals to the corpus luteum with the hormones estradiol and prostaglandin E2.[48] This signaling acts on both the endometrium and luteal tissue to prevent the regression of the corpus luteum by activation of genes that are responsible for corpus luteum maintenance.[49] During mid to late pregnancy, the corpus luteum relies primarily on luteinizing hormone for maintenance until birth.[48]

Archeological evidence indicates that medieval European pigs farrowed, or bore a litter of piglets, once per year.[50] By the nineteenth century, European piglets routinely double-farrowed, or bore two litters of piglets per year. It is unclear when this shift occurred.[51] Pigs have a maximum life span of about 27 years.[52]

Nest-building

A characteristic of pigs which they share with carnivores is nest-building. Sows root in the ground to create depressions the size of their body, and then build nest mounds, using twigs and leaves, softer in the middle, in which to give birth. When the mound reaches the desired height, she places large branches, up to 2 metres in length, on the surface. She enters the mound and roots around to create a depression within the gathered material. She then gives birth in a lying position, unlike other artiodactyls which usually stand while birthing.[53]

Nest-building occurs during the last 24 hours before the onset of farrowing, and becomes most intense 12 to 6 hours before farrowing.[54] The sow separates from the group and seeks a suitable nest site with well-drained soil and shelter from rain and wind. This provides the offspring with shelter, comfort, and thermoregulation. The nest provides protection against weather and predators, while keeping the piglets close to the sow and away from the rest of the herd. This ensures they do not get trampled on, and prevents other piglets from stealing milk from the sow.[55] The onset of nest-building is triggered by a rise in prolactin level, caused by a decrease in progesterone and an increase in prostaglandin; the gathering of nest material seems to be regulated more by external stimuli such as temperature.[54]

Nursing and suckling

Pigs have complex nursing and suckling behaviour.[56] Nursing occurs every 50–60 minutes, and the sow requires stimulation from piglets before milk let-down. Sensory inputs (vocalisation, odours from mammary and birth fluids, and hair patterns of the sow) are particularly important immediately post-birth to facilitate teat location by the piglets.[57] Initially, the piglets compete for position at the udder; then the piglets massage around their respective teats with their snouts, during which time the sow grunts at slow, regular intervals. Each series of grunts varies in frequency, tone and magnitude, indicating the stages of nursing to the piglets.[58]

The phase of competition for teats and of nosing the udder lasts for about a minute, ending when milk begins to flow. The piglets then hold the teats in their mouths and suck with slow mouth movements (one per second), and the rate of the sow's grunting increases for approximately 20 seconds. The grunt peak in the third phase of suckling does not coincide with milk ejection, but rather the release of oxytocin from the pituitary into the bloodstream.[59] Phase four coincides with the period of main milk flow (10–20 seconds) when the piglets suddenly withdraw slightly from the udder and start sucking with rapid mouth movements of about three per second. The sow grunts rapidly, lower in tone and often in quick runs of three or four, during this phase. Finally, the flow stops and so does the grunting of the sow. The piglets may dart from teat to teat and recommence suckling with slow movements, or nosing the udder. Piglets massage and suckle the sow's teats after milk flow ceases as a way of letting the sow know their nutritional status. This helps her to regulate the amount of milk released from that teat in future sucklings. The more intense the post-feed massaging of a teat, the more milk that teat later releases.[60]

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    Sows typically have 12–14 nipples.
  •  
    A sow with suckling piglets

Teat order

In pigs, dominance hierarchies are formed at an early age. Piglets are precocious, and attempt to suckle soon after being born. The piglets are born with sharp teeth and fight for the anterior teats, as these produce more milk. Once established, this teat order remains stable; each piglet tends to feed on a particular teat or group of teats.[53] Stimulation of the anterior teats appears to be important in causing milk letdown,[61] so it might be advantageous to the entire litter to have these teats occupied by healthy piglets. Piglets locate teats by sight and then by olfaction.[62]

Behaviour

Social

 
American Yorkshire pigs in a wallow

Pig behaviour is intermediate between that of other artiodactyls and of carnivores.[53] Pigs seek out the company of other pigs and often huddle to maintain physical contact, but they do not naturally form large herds. They live in groups of about 8–10 adult sows, some young individuals, and some single males.[54] Pigs confined in a simplified, crowded, or uncomfortable environment may resort to tail-biting; farmers sometimes dock the tails of pigs to prevent the problem, or may enrich the environment with toys or other objects to reduce the risk.[63][64]

Temperature control

Because of their relative lack of sweat glands, pigs often control their body temperature using behavioural thermoregulation. Wallowing, coating the body with mud, is a common behaviour.[9] They do not submerge completely under the mud, but vary the depth and duration of wallowing depending on environmental conditions.[9] Adult pigs start wallowing once the ambient temperature is around 17–21 °C (63–70 °F). They cover themselves in mud from head to tail.[9] They may use mud as a sunscreen, or to keep parasites away.[9] Most bristled pigs "blow their coat", meaning that they shed most of the longer, coarser stiff hair once a year, usually in spring or early summer, to prepare for the warmer months ahead.[65]

Eating, feeding, sleeping

 
Pigs around a rotary feeder

Where pigs are allowed to roam freely, they walk roughly 4 km daily, scavenging within a home range of around a hectare. Farmers in Africa often choose such a low-input, free-range production system.[66]

If conditions permit, pigs feed continuously for many hours and then sleep for many hours, in contrast to ruminants, which tend to feed for a short time and then sleep for a short time. Pigs are omnivorous and versatile in their feeding behaviour. They primarily eat leaves, stems, roots, fruits, and flowers.[67]

Rooting is an instinctual comforting behaviour in pigs characterized by nudging the snout into something. It first happens when piglets are born to obtain their mother's milk, and can become a habitual, obsessive behaviour, most prominent in animals weaned too early. Pigs root and dig into the ground to forage for food. Rooting is also a means of communication.[68]

Intelligence

 
A pig using a specifically designed joystick[69]

Pigs are relatively intelligent animals, roughly on par with dogs. They distinguish each other as individuals, spend time in play, and form structured communities. They have good long-term memory and they experience emotions, changing their behaviour in response to the emotional states of other pigs. In terms of experimental tasks, pigs can perform tasks that require them to identify the locations of objects; they can solve mazes; and they can work with a simple language of symbols. They display self-recognition in a mirror. Pigs have been trained to associate different sorts of music (Bach and a military march) with food and social isolation respectively, and could communicate the resulting positive or negative emotion to untrained pigs.[70][71] Pigs can be trained to use a joystick with their snout to select a target on screen.[69]

Senses

 
A trained pig using its sensitive nose to assist the search for wild truffles in France

Pigs have panoramic vision of approximately 310° and binocular vision of 35° to 50°. It is thought they have no eye accommodation.[72] Other animals that have no accommodation, e.g. sheep, lift their heads to see distant objects.[73] The extent to which pigs have colour vision is still a source of some debate; however, the presence of cone cells in the retina with two distinct wavelength sensitivities (blue and green) suggests that at least some colour vision is present.[74]

Pigs have a well-developed sense of smell; this is exploited in Europe where trained pigs find underground truffles.[75] Pigs have 1,113 genes for smell receptors, compared to 1,094 in dogs; this may indicate an acute sense of smell, but against this, insects have only around 50 to 100 such genes but make extensive use of olfaction.[76] Olfactory rather than visual stimuli are used in the identification of other pigs.[77] Hearing is well developed; sounds are localised by moving the head. Pigs use auditory stimuli extensively for communication in all social activities.[78] Alarm or aversive stimuli are transmitted to other pigs not only by auditory cues but also by pheromones.[79] Similarly, recognition between the sow and her piglets is by olfactory and vocal cues.[80]

Pests and diseases

 
Trichinella spiralis larvae in uncooked pig meat

Pigs are subject to many pests and diseases which can seriously affect productivity and cause death. These include parasites such as Ascaris roundworms, virus diseases such as the tick-borne African Swine Fever, bacterial infections such as Clostridium, arthritis caused by Mycoplasma, and stillbirths caused by Parvovirus.[81]

Some parasites of pigs are a public health risk as they can be transmitted to humans in undercooked pork. These are the pork tapeworm Taenia solium; a protozoan, Toxoplasma gondii; and a nematode, Trichinella spiralis. Transmission can be prevented by thorough sanitation on the farm; by meat inspection and careful commercial processing; and by thorough cooking, or alternatively by sufficient freezing and curing.[82]

In agriculture

Further information: Pig farming § Terminology

Production

Main article: Pig farming

Pigs have been raised outdoors, and sometimes allowed to forage in woods or pastures. In industrialized nations, pig production has largely switched to large-scale intensive pig farming. This has lowered production costs but has caused concern about possible cruelty. As consumers have become concerned with the humane treatment of livestock, demand for pasture-raised pork in these nations has increased.[83] Most pigs in the US receive ractopamine, a beta-agonist drug, which promotes muscle instead of fat and quicker weight gain, requiring less feed to reach finishing weight, and producing less manure. China has requested that pork exports be ractopamine-free.[84] With a population of around 1 billion individuals, the domesticated pig is one of the most numerous large mammals on the planet.[85][86]

Like all animals, pigs are susceptible to adverse impacts from climate change, such as heat stress from increased annual temperatures and more intense heatwaves. Heat stress has increased rapidly between 1981 and 2017 on pig farms in Europe. Installing a ground-coupled heat exchanger is an effective intervention.[87]

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    Indoor pig farm, Sweden, 1911
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    Sow in stall with separate piglet balcony to prevent crushing, Germany, 1959
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    Free range pigs with field shelters, England, 2006
  • FAO data for 2021
  •  
    Pork is tied with chicken as the most commonly consumed meat worldwide.
  •  
    Pork production has grown substantially over the recent 60 years.
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    Production of pork worldwide, by country in 2021.

Breeds

Main article: List of pig breeds

Around 600 breeds of pig have been created by farmers around the world, mainly in Europe and Asia, differing in coloration, shape, and size.[88] According to The Livestock Conservancy, as of 2016, three breeds of pig are critically rare (having a global population of fewer than 2000). They are the Choctaw hog, the Mulefoot, and the Ossabaw Island hog.[89] The smallest known pig breed in the world is the Göttingen minipig, typically weighing about 26 kilograms (57 lb) as a healthy, full-grown adult.[90]

As pets

 
A mini pet pig

Vietnamese Pot-bellied pigs, a miniature breed of pig, have been kept as pets in the United States, beginning in the latter half of the 20th century.

Pigs are intelligent, social creatures. They are considered hypoallergenic and are known to do quite well with people who have the usual animal allergies. Since these animals are known to have a life expectancy of 15 to 20 years, they require a long-term commitment.

Given pigs are bred primarily as livestock and have not been bred as companion animals for very long, selective breeding for a placid or biddable temperament is not well established. Pigs have radically different psychology to dogs and exhibit fight-or-flight instincts, independent nature, and natural assertiveness.[91] Male and female swine that have not been de-sexed may express unwanted aggressive behavior, and are prone to developing serious health issues.[92] As rooting is found to be comforting, pigs kept in the house may root household objects, furniture or surfaces. Pet pigs should be let outside to allow them to fulfill their natural desire of rooting around.

Economy

Global pig stock
in 2019
Number in millions
1. China (Mainland)310.4 (36.5%)
2. European Union143.1 (16.83%)
3. United States78.7 (9.26%)
4. Brazil40.6 (4.77%)
5. Russia23.7 (2.79%)
6. Myanmar21.6 (2.54%)
7. Vietnam19.6 (2.31%)
8. Mexico18.4 (2.16%)
9. Canada14.1 (1.66%)
10. Philippines12.7 (1.49%)
World total850.3
Source: UN Food and Agriculture Organization

Approximately 1.5 billion pigs are slaughtered each year for meat.[93]

The pork belly futures contract became an icon of commodities trading. It appears in depictions of the arena in popular entertainment, such as the 1983 film Trading Places.[94] Trade in pork bellies declined, and they were delisted from the Chicago Mercantile Exchange in 2011.[94][95]

In 2023, China produced more pork than any other country, 55 million tonnes, followed by the European Union with 22.8 million tonnes and the United States with 12.5 million tonnes. Global production in 2023 was 120 million tonnes.[96] India, despite its large population, consumed under 0.3 million tonnes of pork in 2023.[97] International trade in pork (meat not consumed in the producing country) reached 13 million tonnes in 2020.[98]

Uses

Products

Pigs are farmed primarily for meat, called pork. Pork is eaten in the form of pork chops, loin or rib roasts, shoulder joints, steaks, and loin (also called fillet). The many meat products made from pork include ham, bacon (mainly from the back and belly), and sausages.[99] Pork is further made into charcuterie products such as terrines, galantines, pâtés and confits.[100] Some sausages such as salami are fermented and air-dried, to be eaten raw. There are many types, the original Italian varieties including Genovese, Milanese, and Cacciatorino, with spicier kinds from the South of Italy including Calabrese, Napoletano, and Peperone.[101]

The hide is made into pigskin leather, which is soft and durable; it can be brushed to form suede leather. These are used for products such as gloves, wallets, suede shoes, and leather jackets.[102] In the 16th century, pig skin was the most popular book-binding material in Germany, though calf skin was more common elsewhere.[103]


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    Pork chops
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    Streaky or side bacon
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    Salami, a fermented and air-dried sausage, originally made in Italy
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    A 16th century book bound in pig skin
  •  
    A woman's suede gloves, England, c. 1820

In medicine

 
The growth in publication of medical research papers using pigs and miniature pigs, and the research done on miniature pigs by organ system[104]

Pigs, both as live animals and as a source of post-mortem tissues, are valuable animal models because of their biological, physiological, and anatomical similarities to human beings. For instance, human skin is very similar to the pigskin, therefore pigskin has been used in many preclinical studies.[105][106]

Pigs are good non-human candidates for organ donation to humans, and in 2021 became the first animal to successfully donate an organ to a human body.[107][108] The procedure used a donor pig genetically engineered not to have a specific carbohydrate that the human body considers a threat–Galactose-alpha-1,3-galactose.[109] Pigs are good for human donation as the risk of cross-species disease transmission is reduced by the considerable phylogenetic distance from humans.[110] They are readily available, and the danger of creating new human diseases is low as domesticated pigs have been in close contact with humans for thousands of years.[111]

Impact of pig husbandry

On public health

Further information: Zoonotic disease

Pig farms can serve as reservoirs of viral diseases that are dangerous to humans and so contribute to their outbreaks in human populations.[112] The 2009 swine flu pandemic was caused by an influenza A variant which had first emerged in pigs.[113][114] Pigs were also essential to the first outbreak of the Nipah virus in 1999, with 93% of the infected humans having had contact with pigs.[112] While Japanese encephalitis is primarily spread by mosquitoes, pigs are a known intermediary host.[115] There is also a potential for porcine coronaviruses such as porcine epidemic diarrhea virus or swine acute diarrhea syndrome coronavirus to spill over into human populations.[112]

On the environment

Main article: Environmental impact of pig farming
 
A typical waste lagoon in North Carolina

As with the other forms of meat, producing pork is more energy-intensive than plant-based foods, and it is associated with more greenhouse gas emissions per calorie. However, emissions from pork are many times smaller than those of beef, veal and mutton, though larger than of chicken meat.[116]

Intensive pig production is also associated with water pollution concerns, as the swine waste is often stored above ground in so-called lagoons. These lagoons typically have high levels of nitrogen and phosphorus, and can contain toxic heavy metals like zinc and copper, microbial pathogens, or hold elevated concentrations of pharmaceuticals from subtherapeutic antibiotic use in swine.[117] This wastewater from lagoons is liable to reach groundwater on farms, though there is little evidence for it reaching deeper into local drinking water supplies.[118] However, lagoon spills, such as from heavy rains in the wake of a hurricane, can lead to fish kills and algal blooms in local rivers.[117] In the United States, 35,000 mi (56,000 km) of river across over 20 states were estimated to have been contaminated by manure leakage as of 2015.[119] There is also evidence that evaporation from lagoons can cause nitrogen and phosphorus to spread through the air as dry particles then reach other water basins when they fall out through dry deposition. This process then also contributes to water eutrophication.[117]

On animal welfare

Further information: Cruelty to animals § Welfare concerns of farm animals, and Intensive pig farming
 
Sows in gestation crates, United States, 2010

Intensive pig production involves practices such as castration, earmarking, tattooing for litter identification, tail docking, which are often done without the use of anesthetic.[120] [121] Teeth clipping of piglets is also done to curtail cannibalism, behavioural instability and aggression, and tail biting, which are induced by the cramped environment.[122][123] In indoor farming, pigs are allowed to be kept with less than one square meter of space per pig.[124]

Pigs often begin life in a farrowing or gestation crate, which is a small pen with a central cage, designed to allow the piglets to feed from their mother, the sow, while preventing her from moving around, crushing her children, and reducing aggression.[125] The crates are so small that the pigs cannot turn around.[126][127] While wild piglets remain with their mothers for around 12 to 14 weeks, farmed piglets are weaned and removed from their mothers at between two and five weeks old.[128][129] Of the piglets born alive, 10% to 18% will not reach weaning age, instead succumbing to disease, starvation, dehydration, or accidental crushing by their mothers.[122][130] Unusually small runt piglets are typically killed immediately by staff through blunt trauma to the head.[131][132] Further, intensive farming involves sows giving birth to large litter sizes at an unnatural frequency, which increases the rate of stillborn piglets, and causes as many as 25%-50% of sows to die of prolapse[133][134]

In culture

Main article: Pigs in culture

Pigs, widespread in societies around the world since Neolithic times, have been used for many purposes in art, literature, and other expressions of human culture. In classical times, the Romans considered pork the finest of meats, enjoying sausages, and depicting them in their art.[135] Across Europe, pigs have been celebrated in carnivals since the Middle Ages,[136] becoming specially important in Medieval Germany in cities such as Nuremberg,[137] and in Early Modern Italy in cities such as Bologna.[138][139] Pigs, especially miniature breeds, are occasionally kept as pets.[140][141]

In literature, both for children[142] and adults, pig characters appear in allegories, comic stories, and serious novels.[136][143][144] In art, pigs have been represented in a wide range of media and styles from the earliest times in many cultures.[145] Pig names are used in idioms and animal epithets, often derogatory, since pigs have long been linked with dirtiness and greed,[146][147] while places such as Swindon are named for their association with swine.[148] The eating of pork is forbidden in Islam and Judaism,[149][150] but pigs are sacred in some other religions.[151][152]

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For other uses see Pig disambiguation Swine redirects here For other uses see Swine disambiguation The pig Sus domesticus also called swine pl swine or hog is an omnivorous domesticated even toed hoofed mammal It is named the domestic pig when distinguishing it from other members of the genus Sus It is considered a subspecies of Sus scrofa the wild boar or Eurasian boar by some authorities but as a distinct species by others Pigs were domesticated in the Neolithic both in East Asia and in the Near East When domesticated pigs arrived in Europe they extensively interbred with wild boar but retained their domesticated features Pig Domestic pigs Conservation status Domesticated Scientific classification Domain Eukaryota Kingdom Animalia Phylum Chordata Class Mammalia Order Artiodactyla Family Suidae Genus Sus Species S domesticus Binomial name Sus domesticusErxleben 1777 Synonyms 1 Sus domestica Sus scrofa domesticus Linnaeus 1758 Pigs are farmed primarily for meat called pork The animal s skin or hide is used for leather China is the world s largest pork producer followed by the European Union and then the United States Around 1 5 billion pigs are raised each year producing some 120 million tonnes of meat often cured as bacon Some are kept as pets Pigs have featured in human culture since Neolithic times appearing in art and literature for children and adults and celebrated in cities such as Bologna for their meat products Contents 1 Description 2 Evolution 2 1 Phylogeny 2 2 Taxonomy 2 3 Domestication in the Neolithic 3 History 3 1 Columbian Exchange 3 2 Feral pigs 4 Reproduction 4 1 Physiology 4 2 Nest building 4 3 Nursing and suckling 4 4 Teat order 5 Behaviour 5 1 Social 5 2 Temperature control 5 3 Eating feeding sleeping 5 4 Intelligence 5 5 Senses 6 Pests and diseases 7 In agriculture 7 1 Production 7 2 Breeds 8 As pets 9 Economy 10 Uses 10 1 Products 10 2 In medicine 11 Impact of pig husbandry 11 1 On public health 11 2 On the environment 11 3 On animal welfare 12 In culture 13 References 14 External linksDescriptionThe pig has a large head with a long snout strengthened by a special prenasal bone and a disk of cartilage at the tip 2 The snout is used to dig into the soil to find food and is an acute sense organ The dental formula of adult pigs is 3 1 4 3 3 1 4 3 giving a total of 44 teeth The rear teeth are adapted for crushing In males the canine teeth can form tusks which grow continuously and are sharpened by grinding against each other 2 There are four hoofed toes on each foot the two larger central toes bear most of the weight while the outer two are also used in soft ground 3 Most pigs have rather sparsely bristled hair on their skin though there are some woolly coated breeds such as the Mangalitsa 4 Adult pigs generally weigh between 140 and 300 kg 310 and 660 lb though some breeds can exceed this range Exceptionally a pig called Big Bill weighed 1 157 kg 2 551 lb and had a shoulder height of 1 5 m 4 9 ft 5 Pigs possess both apocrine and eccrine sweat glands although the latter are limited to the snout 6 Pigs like other hairless mammals such as elephants do not use thermal sweat glands in cooling 7 Pigs are less able than many other mammals to dissipate heat from wet mucous membranes in the mouth by panting Their thermoneutral zone is 16 22 C 61 72 F 8 At higher temperatures pigs lose heat by wallowing in mud or water via evaporative cooling although it has been suggested that wallowing may serve other functions such as protection from sunburn ecto parasite control and scent marking 9 Pigs are among four mammalian species with mutations in the nicotinic acetylcholine receptor that protect against snake venom Mongooses honey badgers hedgehogs and pigs all have different modifications to the receptor pocket which prevents a neurotoxin from binding 10 Pigs have small lungs for their body size and are thus more susceptible than other domesticated animals to fatal bronchitis and pneumonia 11 The genome of the pig has been sequenced it contains about 22 342 protein coding genes 12 13 14 nbsp Skeleton nbsp Skull nbsp Bones of the footEvolutionPhylogeny Domestic pigs are related to other pig species as shown in the cladogram based on phylogenetic analysis using mitochondrial DNA 15 Suidae Babirusa nbsp Indonesia Red river hog nbsp Warthogs nbsp Africa Visayan warty pig nbsp Philippine warty pig nbsp Bornean bearded pig nbsp Javan warty pig nbsp Wild boar nbsp Domestic pig East Asian pigs nbsp European pigs nbsp Eurasia Southeast Asia Taxonomy The pig is most often considered to be a subspecies of the wild boar which was given the name Sus scrofa by Carl Linnaeus in 1758 following from this the formal name of the pig is Sus scrofa domesticus 16 17 However in 1777 Johann Christian Polycarp Erxleben classified the pig as a separate species from the wild boar He gave it the name Sus domesticus still used by some taxonomists 18 The American Society of Mammalogists considers it a separate species 19 Domestication in the Neolithic Further information Domestication nbsp The initial emergence of wild pigs followed by the genetic divergence between boars and pigs and the domestication of pigs 20 Archaeological evidence shows that pigs were domesticated from wild boar in the Near East in or around the Tigris Basin 21 being managed in a semi wild state much as they are managed by some modern New Guineans 22 There were pigs in Cyprus more than 11 400 years ago introduced from the mainland implying domestication in the adjacent mainland by then 23 Pigs were separately domesticated in China starting some 8 000 years ago 24 25 26 In the Near East pig husbandry spread for the next few millennia It reduced gradually during the Bronze Age as rural populations instead focused on commodity producing livestock but it was sustained in cities 27 Domestication did not involve reproductive isolation with population bottlenecks Western Asian pigs were introduced into Europe where they crossed with wild boar There appears to have been interbreeding with a now extinct ghost population of wild pigs during the Pleistocene The genomes of domestic pigs show strong selection for genes affecting behavior and morphology Human selection for domestic traits likely counteracted the homogenizing effect of gene flow from wild boars and created domestication islands in the genome 28 29 Pigs arrived in Europe from the Near East at least 8 500 years ago Over the next 3 000 years they interbred with European wild boar until their genome showed less than 5 Near Eastern ancestry yet retained their domesticated features 30 DNA evidence from subfossil remains of teeth and jawbones of Neolithic pigs shows that the first domestic pigs in Europe were brought from the Near East This stimulated the domestication of local European wild boar resulting in a third domestication event with the Near Eastern genes dying out in European pig stock More recently there have been complex exchanges with European domesticated lines being exported in turn to the ancient Near East 31 32 Historical records indicate that Asian pigs were again introduced into Europe during the 18th and early 19th centuries 25 HistoryColumbian Exchange Among the animals that the Spanish introduced to the Chiloe Archipelago in the 16th century Columbian Exchange pigs were the most successful in adapting to local conditions The pigs benefited from abundant shellfish and algae exposed by the large tides of the archipelago 33 Pigs were brought to southeastern North America from Europe by de Soto and other early Spanish explorers Escaped pigs became feral 34 Feral pigs Main article Feral pig nbsp A family of feral pigs Pigs have escaped from farms and gone feral in many parts of the world Feral pigs in the southeastern United States have migrated north to the Midwest where many state agencies have programs to remove them 35 36 37 Feral pigs in New Zealand and northern Queensland have caused substantial environmental damage 38 39 Feral hybrids of the European wild boar with the domestic pig are disruptive to both environment and agriculture as they destroy crops spread animal diseases including Foot and mouth disease and consume wildlife such as juvenile seabirds and young tortoises 40 Feral pig damage is especially an issue in southeastern South America 41 42 ReproductionPhysiology Female pigs reach sexual maturity at 3 12 months of age and come into estrus every 18 24 days if they are not successfully bred The variation in ovulation rate can be attributed to intrinsic factors such as age and genotype as well as extrinsic factors like nutrition environment and the supplementation of exogenous hormones The gestation period averages 112 120 days 43 nbsp Piglets keeping warm together Estrus lasts two to three days and the female s displayed receptiveness to mate is known as standing heat Standing heat is a reflexive response that is stimulated when the female is in contact with the saliva of a sexually mature boar Androstenol is one of the pheromones produced in the submaxillary salivary glands of boars that trigger the female s response 44 The female cervix contains a series of five interdigitating pads or folds that hold the boar s corkscrew shaped penis during copulation 45 Females have bicornuate uteruses and two conceptuses must be present in both uterine horns to enable pregnancy to proceed 46 The mother s body recognises that it is pregnant on days 11 to 12 of pregnancy and is marked by the corpus luteum s producing the sex hormone progesterone 47 To sustain the pregnancy the embryo signals to the corpus luteum with the hormones estradiol and prostaglandin E2 48 This signaling acts on both the endometrium and luteal tissue to prevent the regression of the corpus luteum by activation of genes that are responsible for corpus luteum maintenance 49 During mid to late pregnancy the corpus luteum relies primarily on luteinizing hormone for maintenance until birth 48 Archeological evidence indicates that medieval European pigs farrowed or bore a litter of piglets once per year 50 By the nineteenth century European piglets routinely double farrowed or bore two litters of piglets per year It is unclear when this shift occurred 51 Pigs have a maximum life span of about 27 years 52 Nest building A characteristic of pigs which they share with carnivores is nest building Sows root in the ground to create depressions the size of their body and then build nest mounds using twigs and leaves softer in the middle in which to give birth When the mound reaches the desired height she places large branches up to 2 metres in length on the surface She enters the mound and roots around to create a depression within the gathered material She then gives birth in a lying position unlike other artiodactyls which usually stand while birthing 53 Nest building occurs during the last 24 hours before the onset of farrowing and becomes most intense 12 to 6 hours before farrowing 54 The sow separates from the group and seeks a suitable nest site with well drained soil and shelter from rain and wind This provides the offspring with shelter comfort and thermoregulation The nest provides protection against weather and predators while keeping the piglets close to the sow and away from the rest of the herd This ensures they do not get trampled on and prevents other piglets from stealing milk from the sow 55 The onset of nest building is triggered by a rise in prolactin level caused by a decrease in progesterone and an increase in prostaglandin the gathering of nest material seems to be regulated more by external stimuli such as temperature 54 Nursing and suckling Pigs have complex nursing and suckling behaviour 56 Nursing occurs every 50 60 minutes and the sow requires stimulation from piglets before milk let down Sensory inputs vocalisation odours from mammary and birth fluids and hair patterns of the sow are particularly important immediately post birth to facilitate teat location by the piglets 57 Initially the piglets compete for position at the udder then the piglets massage around their respective teats with their snouts during which time the sow grunts at slow regular intervals Each series of grunts varies in frequency tone and magnitude indicating the stages of nursing to the piglets 58 The phase of competition for teats and of nosing the udder lasts for about a minute ending when milk begins to flow The piglets then hold the teats in their mouths and suck with slow mouth movements one per second and the rate of the sow s grunting increases for approximately 20 seconds The grunt peak in the third phase of suckling does not coincide with milk ejection but rather the release of oxytocin from the pituitary into the bloodstream 59 Phase four coincides with the period of main milk flow 10 20 seconds when the piglets suddenly withdraw slightly from the udder and start sucking with rapid mouth movements of about three per second The sow grunts rapidly lower in tone and often in quick runs of three or four during this phase Finally the flow stops and so does the grunting of the sow The piglets may dart from teat to teat and recommence suckling with slow movements or nosing the udder Piglets massage and suckle the sow s teats after milk flow ceases as a way of letting the sow know their nutritional status This helps her to regulate the amount of milk released from that teat in future sucklings The more intense the post feed massaging of a teat the more milk that teat later releases 60 nbsp Sows typically have 12 14 nipples nbsp A sow with suckling piglets Teat order In pigs dominance hierarchies are formed at an early age Piglets are precocious and attempt to suckle soon after being born The piglets are born with sharp teeth and fight for the anterior teats as these produce more milk Once established this teat order remains stable each piglet tends to feed on a particular teat or group of teats 53 Stimulation of the anterior teats appears to be important in causing milk letdown 61 so it might be advantageous to the entire litter to have these teats occupied by healthy piglets Piglets locate teats by sight and then by olfaction 62 BehaviourSocial nbsp American Yorkshire pigs in a wallow Pig behaviour is intermediate between that of other artiodactyls and of carnivores 53 Pigs seek out the company of other pigs and often huddle to maintain physical contact but they do not naturally form large herds They live in groups of about 8 10 adult sows some young individuals and some single males 54 Pigs confined in a simplified crowded or uncomfortable environment may resort to tail biting farmers sometimes dock the tails of pigs to prevent the problem or may enrich the environment with toys or other objects to reduce the risk 63 64 Temperature control Because of their relative lack of sweat glands pigs often control their body temperature using behavioural thermoregulation Wallowing coating the body with mud is a common behaviour 9 They do not submerge completely under the mud but vary the depth and duration of wallowing depending on environmental conditions 9 Adult pigs start wallowing once the ambient temperature is around 17 21 C 63 70 F They cover themselves in mud from head to tail 9 They may use mud as a sunscreen or to keep parasites away 9 Most bristled pigs blow their coat meaning that they shed most of the longer coarser stiff hair once a year usually in spring or early summer to prepare for the warmer months ahead 65 Eating feeding sleeping nbsp Pigs around a rotary feeder Where pigs are allowed to roam freely they walk roughly 4 km daily scavenging within a home range of around a hectare Farmers in Africa often choose such a low input free range production system 66 If conditions permit pigs feed continuously for many hours and then sleep for many hours in contrast to ruminants which tend to feed for a short time and then sleep for a short time Pigs are omnivorous and versatile in their feeding behaviour They primarily eat leaves stems roots fruits and flowers 67 Rooting is an instinctual comforting behaviour in pigs characterized by nudging the snout into something It first happens when piglets are born to obtain their mother s milk and can become a habitual obsessive behaviour most prominent in animals weaned too early Pigs root and dig into the ground to forage for food Rooting is also a means of communication 68 Intelligence nbsp A pig using a specifically designed joystick 69 Pigs are relatively intelligent animals roughly on par with dogs They distinguish each other as individuals spend time in play and form structured communities They have good long term memory and they experience emotions changing their behaviour in response to the emotional states of other pigs In terms of experimental tasks pigs can perform tasks that require them to identify the locations of objects they can solve mazes and they can work with a simple language of symbols They display self recognition in a mirror Pigs have been trained to associate different sorts of music Bach and a military march with food and social isolation respectively and could communicate the resulting positive or negative emotion to untrained pigs 70 71 Pigs can be trained to use a joystick with their snout to select a target on screen 69 Senses nbsp A trained pig using its sensitive nose to assist the search for wild truffles in France Pigs have panoramic vision of approximately 310 and binocular vision of 35 to 50 It is thought they have no eye accommodation 72 Other animals that have no accommodation e g sheep lift their heads to see distant objects 73 The extent to which pigs have colour vision is still a source of some debate however the presence of cone cells in the retina with two distinct wavelength sensitivities blue and green suggests that at least some colour vision is present 74 Pigs have a well developed sense of smell this is exploited in Europe where trained pigs find underground truffles 75 Pigs have 1 113 genes for smell receptors compared to 1 094 in dogs this may indicate an acute sense of smell but against this insects have only around 50 to 100 such genes but make extensive use of olfaction 76 Olfactory rather than visual stimuli are used in the identification of other pigs 77 Hearing is well developed sounds are localised by moving the head Pigs use auditory stimuli extensively for communication in all social activities 78 Alarm or aversive stimuli are transmitted to other pigs not only by auditory cues but also by pheromones 79 Similarly recognition between the sow and her piglets is by olfactory and vocal cues 80 Pests and diseases nbsp Trichinella spiralis larvae in uncooked pig meat Pigs are subject to many pests and diseases which can seriously affect productivity and cause death These include parasites such as Ascaris roundworms virus diseases such as the tick borne African Swine Fever bacterial infections such as Clostridium arthritis caused by Mycoplasma and stillbirths caused by Parvovirus 81 Some parasites of pigs are a public health risk as they can be transmitted to humans in undercooked pork These are the pork tapeworm Taenia solium a protozoan Toxoplasma gondii and a nematode Trichinella spiralis Transmission can be prevented by thorough sanitation on the farm by meat inspection and careful commercial processing and by thorough cooking or alternatively by sufficient freezing and curing 82 In agricultureFurther information Pig farming Terminology Production Main article Pig farming Pigs have been raised outdoors and sometimes allowed to forage in woods or pastures In industrialized nations pig production has largely switched to large scale intensive pig farming This has lowered production costs but has caused concern about possible cruelty As consumers have become concerned with the humane treatment of livestock demand for pasture raised pork in these nations has increased 83 Most pigs in the US receive ractopamine a beta agonist drug which promotes muscle instead of fat and quicker weight gain requiring less feed to reach finishing weight and producing less manure China has requested that pork exports be ractopamine free 84 With a population of around 1 billion individuals the domesticated pig is one of the most numerous large mammals on the planet 85 86 Like all animals pigs are susceptible to adverse impacts from climate change such as heat stress from increased annual temperatures and more intense heatwaves Heat stress has increased rapidly between 1981 and 2017 on pig farms in Europe Installing a ground coupled heat exchanger is an effective intervention 87 nbsp Indoor pig farm Sweden 1911 nbsp Sow in stall with separate piglet balcony to prevent crushing Germany 1959 nbsp Free range pigs with field shelters England 2006 FAO data for 2021 nbsp Pork is tied with chicken as the most commonly consumed meat worldwide nbsp Pork production has grown substantially over the recent 60 years nbsp Production of pork worldwide by country in 2021 Breeds Main article List of pig breeds Around 600 breeds of pig have been created by farmers around the world mainly in Europe and Asia differing in coloration shape and size 88 According to The Livestock Conservancy as of 2016 three breeds of pig are critically rare having a global population of fewer than 2000 They are the Choctaw hog the Mulefoot and the Ossabaw Island hog 89 The smallest known pig breed in the world is the Gottingen minipig typically weighing about 26 kilograms 57 lb as a healthy full grown adult 90 As pets nbsp A mini pet pig Vietnamese Pot bellied pigs a miniature breed of pig have been kept as pets in the United States beginning in the latter half of the 20th century Pigs are intelligent social creatures They are considered hypoallergenic and are known to do quite well with people who have the usual animal allergies Since these animals are known to have a life expectancy of 15 to 20 years they require a long term commitment Given pigs are bred primarily as livestock and have not been bred as companion animals for very long selective breeding for a placid or biddable temperament is not well established Pigs have radically different psychology to dogs and exhibit fight or flight instincts independent nature and natural assertiveness 91 Male and female swine that have not been de sexed may express unwanted aggressive behavior and are prone to developing serious health issues 92 As rooting is found to be comforting pigs kept in the house may root household objects furniture or surfaces Pet pigs should be let outside to allow them to fulfill their natural desire of rooting around EconomyGlobal pig stockin 2019Number in millions1 China Mainland 310 4 36 5 2 European Union143 1 16 83 3 United States78 7 9 26 4 Brazil40 6 4 77 5 Russia23 7 2 79 6 Myanmar21 6 2 54 7 Vietnam19 6 2 31 8 Mexico18 4 2 16 9 Canada14 1 1 66 10 Philippines12 7 1 49 World total850 3Source UN Food and Agriculture Organization Approximately 1 5 billion pigs are slaughtered each year for meat 93 The pork belly futures contract became an icon of commodities trading It appears in depictions of the arena in popular entertainment such as the 1983 film Trading Places 94 Trade in pork bellies declined and they were delisted from the Chicago Mercantile Exchange in 2011 94 95 In 2023 China produced more pork than any other country 55 million tonnes followed by the European Union with 22 8 million tonnes and the United States with 12 5 million tonnes Global production in 2023 was 120 million tonnes 96 India despite its large population consumed under 0 3 million tonnes of pork in 2023 97 International trade in pork meat not consumed in the producing country reached 13 million tonnes in 2020 98 UsesProducts Pigs are farmed primarily for meat called pork Pork is eaten in the form of pork chops loin or rib roasts shoulder joints steaks and loin also called fillet The many meat products made from pork include ham bacon mainly from the back and belly and sausages 99 Pork is further made into charcuterie products such as terrines galantines pates and confits 100 Some sausages such as salami are fermented and air dried to be eaten raw There are many types the original Italian varieties including Genovese Milanese and Cacciatorino with spicier kinds from the South of Italy including Calabrese Napoletano and Peperone 101 The hide is made into pigskin leather which is soft and durable it can be brushed to form suede leather These are used for products such as gloves wallets suede shoes and leather jackets 102 In the 16th century pig skin was the most popular book binding material in Germany though calf skin was more common elsewhere 103 nbsp Pork chops nbsp Streaky or side bacon nbsp Salami a fermented and air dried sausage originally made in Italy nbsp A 16th century book bound in pig skin nbsp A woman s suede gloves England c 1820 In medicine nbsp The growth in publication of medical research papers using pigs and miniature pigs and the research done on miniature pigs by organ system 104 Pigs both as live animals and as a source of post mortem tissues are valuable animal models because of their biological physiological and anatomical similarities to human beings For instance human skin is very similar to the pigskin therefore pigskin has been used in many preclinical studies 105 106 Pigs are good non human candidates for organ donation to humans and in 2021 became the first animal to successfully donate an organ to a human body 107 108 The procedure used a donor pig genetically engineered not to have a specific carbohydrate that the human body considers a threat Galactose alpha 1 3 galactose 109 Pigs are good for human donation as the risk of cross species disease transmission is reduced by the considerable phylogenetic distance from humans 110 They are readily available and the danger of creating new human diseases is low as domesticated pigs have been in close contact with humans for thousands of years 111 Impact of pig husbandryOn public health Further information Zoonotic disease Pig farms can serve as reservoirs of viral diseases that are dangerous to humans and so contribute to their outbreaks in human populations 112 The 2009 swine flu pandemic was caused by an influenza A variant which had first emerged in pigs 113 114 Pigs were also essential to the first outbreak of the Nipah virus in 1999 with 93 of the infected humans having had contact with pigs 112 While Japanese encephalitis is primarily spread by mosquitoes pigs are a known intermediary host 115 There is also a potential for porcine coronaviruses such as porcine epidemic diarrhea virus or swine acute diarrhea syndrome coronavirus to spill over into human populations 112 On the environment Main article Environmental impact of pig farming nbsp A typical waste lagoon in North Carolina As with the other forms of meat producing pork is more energy intensive than plant based foods and it is associated with more greenhouse gas emissions per calorie However emissions from pork are many times smaller than those of beef veal and mutton though larger than of chicken meat 116 Intensive pig production is also associated with water pollution concerns as the swine waste is often stored above ground in so called lagoons These lagoons typically have high levels of nitrogen and phosphorus and can contain toxic heavy metals like zinc and copper microbial pathogens or hold elevated concentrations of pharmaceuticals from subtherapeutic antibiotic use in swine 117 This wastewater from lagoons is liable to reach groundwater on farms though there is little evidence for it reaching deeper into local drinking water supplies 118 However lagoon spills such as from heavy rains in the wake of a hurricane can lead to fish kills and algal blooms in local rivers 117 In the United States 35 000 mi 56 000 km of river across over 20 states were estimated to have been contaminated by manure leakage as of 2015 119 There is also evidence that evaporation from lagoons can cause nitrogen and phosphorus to spread through the air as dry particles then reach other water basins when they fall out through dry deposition This process then also contributes to water eutrophication 117 On animal welfare Further information Cruelty to animals Welfare concerns of farm animals and Intensive pig farming nbsp Sows in gestation crates United States 2010 Intensive pig production involves practices such as castration earmarking tattooing for litter identification tail docking which are often done without the use of anesthetic 120 121 Teeth clipping of piglets is also done to curtail cannibalism behavioural instability and aggression and tail biting which are induced by the cramped environment 122 123 In indoor farming pigs are allowed to be kept with less than one square meter of space per pig 124 Pigs often begin life in a farrowing or gestation crate which is a small pen with a central cage designed to allow the piglets to feed from their mother the sow while preventing her from moving around crushing her children and reducing aggression 125 The crates are so small that the pigs cannot turn around 126 127 While wild piglets remain with their mothers for around 12 to 14 weeks farmed piglets are weaned and removed from their mothers at between two and five weeks old 128 129 Of the piglets born alive 10 to 18 will not reach weaning age instead succumbing to disease starvation dehydration or accidental crushing by their mothers 122 130 Unusually small runt piglets are typically killed immediately by staff through blunt trauma to the head 131 132 Further intensive farming involves sows giving birth to large litter sizes at an unnatural frequency which increases the rate of stillborn piglets and causes as many as 25 50 of sows to die of prolapse 133 134 In cultureMain article Pigs in culture Pigs widespread in societies around the world since Neolithic times have been used for many purposes in art literature and other expressions of human culture In classical times the Romans considered pork the finest of meats enjoying sausages and depicting them in their art 135 Across Europe pigs have been celebrated in carnivals since the Middle Ages 136 becoming specially important in Medieval Germany in cities such as Nuremberg 137 and in Early Modern Italy in cities such as Bologna 138 139 Pigs especially miniature breeds are occasionally kept as pets 140 141 In literature both for children 142 and adults pig characters appear in allegories comic stories and serious novels 136 143 144 In art pigs have been represented in a wide range of media and styles from the earliest times in many cultures 145 Pig names are used in idioms and animal epithets often derogatory since pigs have long been linked with dirtiness and greed 146 147 while places such as Swindon are named for their association with swine 148 The eating of pork is forbidden in Islam and Judaism 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