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Cornus nuttallii

Cornus nuttallii, the Pacific dogwood,[1][2] western dogwood,[3] or mountain dogwood,[2] is a species of dogwood tree native to western North America.

Pacific Dogwood
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Cornales
Family: Cornaceae
Genus: Cornus
Subgenus: Cornus subg. Cynoxylon
Species:
C. nuttallii
Binomial name
Cornus nuttallii
Natural range
Synonyms

Benthamidia nuttallii (Audubon ex Torr. & A.Gray) Moldenke

Description

 
The small flowers are in a dense cluster surrounded by large white bracts.

It is a small to medium-sized deciduous tree, reaching 6–23 metres (20–75 feet) tall, often with a canopy spread of 6 m (20 ft). Its habit varies based on the level of sunlight; in full sun it will have a short trunk with a crown as wide as it is tall, while under a canopy it will have a tapered trunk with a short, slender crown.[4] The trunk attains 15–30 centimetres (6–12 in) in diameter. The bark is reddish brown.[5]

The branches have fine hairs and the young bark is thin and smooth, becoming scale-like with ridges as it ages.[4]

The leaves are opposite, simple, oval, 5–13 cm (2–5 in) long, and 3.8–7.1 cm (1+12–3 in) broad. They are green with stiff, appressed hairs on top, and hairier and lighter on the bottom.[4][5] They turn orange to purplish in autumn.[5]

The flowers are individually small and inconspicuous, 2–3 millimetres (11618 in) across, produced in a dense, rounded, greenish-white flower head 2 cm (34 in) in diameter; the 4–8 large white 'petals' are actually bracts, each bract 4–7 cm (1+122+34 in) long and broad, creating the appearance of a larger flower head. The flowers commonly bloom twice per season, once in the spring and again in late summer or early fall.[4][3]

The fruit is a compound pink-red or orange drupe about 1–1.5 cm (1212 in) long, in clusters containing 20–40 drupelets, each of which contains two seeds. They appear in September or October.[3][4]

Similar species

The eastern United States' Cornus florida (flowering dogwood) is similar in appearance and possibly in chemical composition. Cornus canadensis has similar blossoms but grows as a groundcover.[5]

Etymology

In 1806, Meriwether Lewis noted that the species is similar in appearance to C. florida.[5] However, when Scottish botanist David Douglas encountered C. nuttalli on his expedition to the Pacific Northwest in the 1820s, he mistook it for C. florida and did not send seeds back to England.[3]

English botanist Thomas Nuttall was the first to describe the species for science while staying at Fort Vancouver in the autumn of 1834.[6] It was named nuttallii after him by his friend John James Audubon.[3]

The common names comes from that of Cornus sanguinea, the hard wood of which Northern Europeans used to make nails ("dags") during the medieval era.[5]

 
British Columbia Dogwood in flower.

Distribution and habitat

 
Pacific Dogwood in the understory of a forest, showing its typical habit.

It occurs from the lowlands of southern British Columbia to the mountains of southern California. There exists an inland population in central Idaho, where it is considered critically imperiled. It occurs predominantly below 1,500 m (4,900 ft) in elevation.[4]

Cultivated examples are found as far north as Haida Gwaii.

It has high flood tolerance, and is common along streams with moist but well-drained soils, often on gentle slopes. Soil composition can range from clay to sandy loam, and it prefers a high humus content, moderate to high nutrient levels, and acidic soils with a pH from 5.5 to 6. It has low frost tolerance, and is usually found in low-elevation temperate or mesothermal climates.[4] It is hardy to USDA zone 7.[3]

Ecology

Value to animals

New sprouts are good browse for both wild and domesticated ungulates, especially after a recent fire, but the mature foliage is usually ignored by all species except slugs.[4]

The fruit are eaten by deer mice, pileated woodpeckers, the band-tailed pigeon,[4][7] and bears.[5]

It provides habitat and cover to small birds such as Wilson's warbler, and small mammals including the red tree vole.[4]

Diseases

Like the related Cornus florida, it is very susceptible to dogwood anthracnose, a disease caused by the fungus Discula destructiva. Fungal activity is greatest from May to July, although it can be active any time conditions are moist and the plant is growing. Infected leaves become blotched and drop, and defoliation can be extreme. Twigs and leaf buds are also impacted. This has killed many of the larger plants in the wild and has also restricted its use as an ornamental tree, to the point where it is considered threatening to the species in its native range.[4]

Successional status

It is present in all stages of both primary and secondary succession – from new colonization on glacial outwash or areas destroyed by the 1980 eruption of Mount St. Helens, to late seral and even climax communities.[4] It is shade tolerant but prefers sunlight in sufficiently humid conditions.[5]

It is adapted to a wide variety of fire regimes, with intervals ranging from just one year on dry sites, to 500 years or more in moist, riparian zones. The tree can survive low severity wildfires which are not hot enough to kill buds protected by bark. After being severely burned, Cornus nuttali typically resprouts from the root crown – however, the resulting shoots are so palatable to mule deer that they are at risk of being killed by over-browsing.[4]

Uses

Some Plateau Indian tribes such as the Nlaka'pamux used the bark as a brown dye. Those groups also used the bark for medicinal purposes as a blood purifier, lung strengthener, stomach treatment, laxative, and emetic.[4][8]

The berries are edible by humans, though not very palatable.

It is mostly prized as a cultivated ornamental.[5]

Culture

It has been the provincial flower of British Columbia[9] since 1956.[10] It was once protected by law in the province (in an act which also protected Rhododendron macrophyllum and Trillium ovatum),[11] but this was repealed in 2002.[12]

References

  1. ^ USDA, NRCS (n.d.). "Cornus nuttallii". The PLANTS Database (plants.usda.gov). Greensboro, North Carolina: National Plant Data Team. Retrieved 11 February 2014.
  2. ^ a b "Cornus nuttallii". Calflora. The Calflora Database. 2014. Retrieved 12 February 2014.
  3. ^ a b c d e f "Cornus nuttallii | Landscape Plants | Oregon State University". landscapeplants.oregonstate.edu. Retrieved 2021-05-12.
  4. ^ a b c d e f g h i j k l m n "Species: Cornus nuttallii". www.fs.usda.gov. Retrieved 2021-05-12.
  5. ^ a b c d e f g h i Arno, Stephen F.; Hammerly, Ramona P. (2020) [1977]. Northwest Trees: Identifying & Understanding the Region's Native Trees (field guide ed.). Seattle: Mountaineers Books. pp. 267–271. ISBN 978-1-68051-329-5. OCLC 1141235469.
  6. ^ "Thomas Nuttall (U.S. National Park Service)". www.nps.gov. Retrieved 2021-05-12.
  7. ^ Peattie, Donald Culross (1953). A Natural History of Western Trees. New York: Bonanza Books. p. 655.
  8. ^ Hunn, Eugene S. (1990). Nch'i-Wana, "The Big River": Mid-Columbia Indians and Their Land. University of Washington Press. p. 353. ISBN 0-295-97119-3.
  9. ^ "Provincial Symbols and Honours Act". Queen's Printer British Columbia. Retrieved 2008-05-06.
  10. ^ "British Columbia (BC) - Facts, Flags and Symbols". 30 April 2010.
  11. ^ . Queen's Printer British Columbia. Archived from the original on 2002-01-02. Retrieved 2008-05-06.
  12. ^ "Dogwood, Rhododendron and Trillium Protection Act". Queen's Printer British Columbia. Retrieved 2008-05-06.

External links

cornus, nuttallii, western, dogwood, redirects, here, shrub, with, same, name, cornus, sericea, pacific, dogwood, western, dogwood, mountain, dogwood, species, dogwood, tree, native, western, north, america, pacific, dogwoodscientific, classificationkingdom, p. Western Dogwood redirects here For the shrub with the same name see Cornus sericea Cornus nuttallii the Pacific dogwood 1 2 western dogwood 3 or mountain dogwood 2 is a species of dogwood tree native to western North America Pacific DogwoodScientific classificationKingdom PlantaeClade TracheophytesClade AngiospermsClade EudicotsClade AsteridsOrder CornalesFamily CornaceaeGenus CornusSubgenus Cornus subg CynoxylonSpecies C nuttalliiBinomial nameCornus nuttalliiAudubon ex Torr amp A GrayNatural rangeSynonymsBenthamidia nuttallii Audubon ex Torr amp A Gray Moldenke Contents 1 Description 1 1 Similar species 2 Etymology 3 Distribution and habitat 4 Ecology 4 1 Value to animals 4 2 Diseases 4 3 Successional status 5 Uses 6 Culture 7 References 8 External linksDescription Edit The small flowers are in a dense cluster surrounded by large white bracts It is a small to medium sized deciduous tree reaching 6 23 metres 20 75 feet tall often with a canopy spread of 6 m 20 ft Its habit varies based on the level of sunlight in full sun it will have a short trunk with a crown as wide as it is tall while under a canopy it will have a tapered trunk with a short slender crown 4 The trunk attains 15 30 centimetres 6 12 in in diameter The bark is reddish brown 5 The branches have fine hairs and the young bark is thin and smooth becoming scale like with ridges as it ages 4 The leaves are opposite simple oval 5 13 cm 2 5 in long and 3 8 7 1 cm 1 1 2 3 in broad They are green with stiff appressed hairs on top and hairier and lighter on the bottom 4 5 They turn orange to purplish in autumn 5 The flowers are individually small and inconspicuous 2 3 millimetres 1 16 1 8 in across produced in a dense rounded greenish white flower head 2 cm 3 4 in in diameter the 4 8 large white petals are actually bracts each bract 4 7 cm 1 1 2 2 3 4 in long and broad creating the appearance of a larger flower head The flowers commonly bloom twice per season once in the spring and again in late summer or early fall 4 3 The fruit is a compound pink red or orange drupe about 1 1 5 cm 1 2 1 2 in long in clusters containing 20 40 drupelets each of which contains two seeds They appear in September or October 3 4 Similar species Edit The eastern United States Cornus florida flowering dogwood is similar in appearance and possibly in chemical composition Cornus canadensis has similar blossoms but grows as a groundcover 5 Etymology EditIn 1806 Meriwether Lewis noted that the species is similar in appearance to C florida 5 However when Scottish botanist David Douglas encountered C nuttalli on his expedition to the Pacific Northwest in the 1820s he mistook it for C florida and did not send seeds back to England 3 English botanist Thomas Nuttall was the first to describe the species for science while staying at Fort Vancouver in the autumn of 1834 6 It was named nuttallii after him by his friend John James Audubon 3 The common names comes from that of Cornus sanguinea the hard wood of which Northern Europeans used to make nails dags during the medieval era 5 British Columbia Dogwood in flower Distribution and habitat Edit Pacific Dogwood in the understory of a forest showing its typical habit It occurs from the lowlands of southern British Columbia to the mountains of southern California There exists an inland population in central Idaho where it is considered critically imperiled It occurs predominantly below 1 500 m 4 900 ft in elevation 4 Cultivated examples are found as far north as Haida Gwaii It has high flood tolerance and is common along streams with moist but well drained soils often on gentle slopes Soil composition can range from clay to sandy loam and it prefers a high humus content moderate to high nutrient levels and acidic soils with a pH from 5 5 to 6 It has low frost tolerance and is usually found in low elevation temperate or mesothermal climates 4 It is hardy to USDA zone 7 3 Ecology EditValue to animals Edit New sprouts are good browse for both wild and domesticated ungulates especially after a recent fire but the mature foliage is usually ignored by all species except slugs 4 The fruit are eaten by deer mice pileated woodpeckers the band tailed pigeon 4 7 and bears 5 It provides habitat and cover to small birds such as Wilson s warbler and small mammals including the red tree vole 4 Diseases Edit Like the related Cornus florida it is very susceptible to dogwood anthracnose a disease caused by the fungus Discula destructiva Fungal activity is greatest from May to July although it can be active any time conditions are moist and the plant is growing Infected leaves become blotched and drop and defoliation can be extreme Twigs and leaf buds are also impacted This has killed many of the larger plants in the wild and has also restricted its use as an ornamental tree to the point where it is considered threatening to the species in its native range 4 Successional status Edit It is present in all stages of both primary and secondary succession from new colonization on glacial outwash or areas destroyed by the 1980 eruption of Mount St Helens to late seral and even climax communities 4 It is shade tolerant but prefers sunlight in sufficiently humid conditions 5 It is adapted to a wide variety of fire regimes with intervals ranging from just one year on dry sites to 500 years or more in moist riparian zones The tree can survive low severity wildfires which are not hot enough to kill buds protected by bark After being severely burned Cornus nuttali typically resprouts from the root crown however the resulting shoots are so palatable to mule deer that they are at risk of being killed by over browsing 4 Uses EditSome Plateau Indian tribes such as the Nlaka pamux used the bark as a brown dye Those groups also used the bark for medicinal purposes as a blood purifier lung strengthener stomach treatment laxative and emetic 4 8 The berries are edible by humans though not very palatable It is mostly prized as a cultivated ornamental 5 Culture EditIt has been the provincial flower of British Columbia 9 since 1956 10 It was once protected by law in the province in an act which also protected Rhododendron macrophyllum and Trillium ovatum 11 but this was repealed in 2002 12 References Edit USDA NRCS n d Cornus nuttallii The PLANTS Database plants usda gov Greensboro North Carolina National Plant Data Team Retrieved 11 February 2014 a b Cornus nuttallii Calflora The Calflora Database 2014 Retrieved 12 February 2014 a b c d e f Cornus nuttallii Landscape Plants Oregon State University landscapeplants oregonstate edu Retrieved 2021 05 12 a b c d e f g h i j k l m n Species Cornus nuttallii www fs usda gov Retrieved 2021 05 12 a b c d e f g h i Arno Stephen F Hammerly Ramona P 2020 1977 Northwest Trees Identifying amp Understanding the Region s Native Trees field guide ed Seattle Mountaineers Books pp 267 271 ISBN 978 1 68051 329 5 OCLC 1141235469 Thomas Nuttall U S National Park Service www nps gov Retrieved 2021 05 12 Peattie Donald Culross 1953 A Natural History of Western Trees New York Bonanza Books p 655 Hunn Eugene S 1990 Nch i Wana The Big River Mid Columbia Indians and Their Land University of Washington Press p 353 ISBN 0 295 97119 3 Provincial Symbols and Honours Act Queen s Printer British Columbia Retrieved 2008 05 06 British Columbia BC Facts Flags and Symbols 30 April 2010 Dogwood Rhododendron and Trillium Protection Act Queen s Printer British Columbia Archived from the original on 2002 01 02 Retrieved 2008 05 06 Dogwood Rhododendron and Trillium Protection Act Queen s Printer British Columbia Retrieved 2008 05 06 External links Edit Wikimedia Commons has media related to Cornus nuttallii Jepson Flora Project Cornus nuttallii Plants of British Columbia Cornus nuttallii Interactive Distribution Map of Cornus nuttallii Cornus nuttallii Plants for a Future Retrieved from https en wikipedia org w index php title Cornus nuttallii amp oldid 1166686729, wikipedia, wiki, book, books, library,

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