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Madygen Formation

The Madygen Formation (Russian: Madygen Svita) is a Late Triassic (Carnian) geologic formation and Lagerstätte in the Batken and Osh Regions of western Kyrgyzstan, with minor outcrops in neighboring Tajikistan and Uzbekistan. The conglomerates, sandstones and mudstones of the 560 m (1,840 ft) thick formation were deposited in terrestrial lacustrine, alluvial, fluvial and deltaic environments.

Madygen Formation
Stratigraphic range: Carnian
~235–222 Ma
TypeGeological formation
OverliesCambrian to Carboniferous rocks
Thickness560 m (1,840 ft)
Lithology
PrimaryConglomerate, sandstone
OtherMudstone
Location
Coordinates40°06′N 70°12′E / 40.1°N 70.2°E / 40.1; 70.2Coordinates: 40°06′N 70°12′E / 40.1°N 70.2°E / 40.1; 70.2
Approximate paleocoordinates41°12′N 60°36′E / 41.2°N 60.6°E / 41.2; 60.6
RegionBatken & Osh Regions
Country Kyrgyzstan
 Tajikistan
 Uzbekistan
ExtentFergana Valley & Range
Type section
Named forMadygen village
Named byEvgeny A. Kochnev
Madygen Formation (Kyrgyzstan)

The formation, extending across the Fergana Valley and Fergana Range, is unique for Central Asia, as it represents one of the few known continental deposits and the Madygen Formation is renowned for the preservation of more than 20,000 fossil insects, making it one of the richest Triassic Lagerstätten in the world. Other vertebrate fossils as fish, amphibians, reptiles and synapsids have been recovered from the formation too, as well as minor fossil flora.

The lake sediments of the Lagerstätte provided fossil cartilaginous fishes and their egg capsules and unusual Triassic reptiles like Sharovipteryx and Longisquama.[1][2] The wide diversity of insect fossils was first discovered in the 1960s and first described by Russian paleontologist Aleksandr Sharov, with a notable example being Gigatitan.[3]

Description

 
Paleogeography of the Late Triassic, around 230 Ma. The Madygen Formation was deposited north of the Paleo-Tethys ocean.

The Madygen Formation is a 560 metres (1,840 ft)[4] thick succession of predominantly siliciclastic rocks accumulated in a tectonically induced basin, covering parts of the Fergana Range and Fergana Valley of Kyrgyzstan with minor outcrops in Tajikistan and Uzbekistan.[5] The Late Triassic layers rest on top of Paleozoic basement with local Permo-Triassic molasse sediments. The section consists of mudstones, sandstones, conglomerates and fanglomerates. This wide variety of siliciclastic rocks reflects the complex spatial and temporal pattern of depositional sub-environments including alluvial fans, sandflats, swamps, back-swamp areas, and littoral to profundal lake zones. The fluvio-lacustrine deposits of the Madygen Formation belong to one of only a few occurrences of continental Triassic beds in Central Asia.[6]

The formation was deposited during the Carnian Pluvial Event (CPE), a global humid event leading to high extinction levels of various groups globally. The CPE led to anoxic conditions, most notably in the South China Block. The area where the Madygen Formation was deposited formed part of the Cimmerian microcontinent, a slab of crust that collided with Laurasia during the Cimmerian orogeny in later Mesozoic times. This orogeny led to the disappearance of the Paleo-Tethys Ocean.

Petroleum geology

The formation grades from bottom to top from alluvial to fluvial into a thick succession of lacustrine mudstones, followed by an alluvial package, on top of which lacustrine, fluvial, deltaic and alluvial layers were deposited.[5]

The hydrocarbon potential of samples of the Madygen Formation ranges from poor to excellent. The sediments containing more than 0.5% Total Organic Carbon (TOC) may be regarded as sources of gaseous hydrocarbons rather than of oil.[7] The Hydrogen Index (HI) of outcrop samples reaches 100 and the maximum recorded maturity (Ro) is 0.8.[8]

Paleontological significance

During the 1960s, Russian paleontologists recovered an unusually rich fossil content in the type strata of the Madygen Formation, including abundant macrophytes, more than 20,000 insect remains[9] and unique small reptiles with well preserved soft tissue.[6] Spirorbis-like polychaete worm tubes, crustaceans (ostracods, kazacharthrans, malacostraca), freshwater pelecypods and gastropods are known from shallow to deeper lake environments. Non-aquatic insects are among the most common fossil remains of the Madygen Formation. These include representatives of the orders Ephemeroptera, Odonata, Notoptera, Blattodea, Titanoptera, Ensifera, Caelifera, Rhynchota, Auchenorrhyncha, Stenorrhyncha, Coleoptera, Hymenoptera, Trichoptera and Diptera. Traces of insect larvae are preserved in near-shore lake deposits.[10]

Fish remains mostly represent endemic genera assigned to the actinopterygian families Evenkiidae (Oshia), Palaeoniscidae (Ferganiscus, Sixtelia) and Megaperleidus and Alvinia. The actinopterygian Saurichthys and the dipnoan Asiatoceratodus are cosmopolitan taxa also recorded in the Madygen Formation. Two distinctive elasmobranch egg capsule types, i.e. Palaeoxyris, indicating a small Lissodus- or Lonchidion-like hybodont shark and an indeterminate capsule type, imply the presence of two different elasmobranch species which used the freshwater environments of the Madygen Formation as spawning grounds. Tetrapods are known from a probably larval urodelan (Triassurus), a small procynosuchid cynodont (Madysaurus), a gliding reptile (Sharovipteryx) and the enigmatic diapsid Longisquama.[10]

Fossil content

Amphibians

Genus Species Material Notes Images
Madygenerpeton M. pustulatus A chroniosuchid reptiliomorph[11]
 
Triassurus T. sixtelae A possible early caudate (salamander) known from fossil larvae[12]
 

Reptiles

Genus Species Material Notes Images
Kyrgyzsaurus K. bukhanchenkoi A single specimen preserving the front half of a skeleton and scale impressions A drepanosaur[13]
 
Longisquama L. insignis A reptile with an elongated row of scales along its back[14]
 
Sharovipteryx S. mirabilis A gliding prolacertiform[15]
 

Synapsids

Genus Species Material Notes Images
Madysaurus M. sharovi A cynodont[16]
 

Cartilaginous fishes

Genus Species Material Notes Images
Fayolia F. sharovi An egg capsule likely belonging to a xenacanthid[1]
Lonchidion L. ferganensis Teeth and egg capsules A hybodontid shark[1]
Palaeoxyris P. alterna Teeth and egg capsules A hybodontid shark[1]

Fishes

The following fish fossils were found in the formation:[10][17][18]

Genus Species Images
Alvinia Alvinia serrata
Ferganiscus Ferganiscus osteolepis
Megaperleidus Megaperleidus lissolepis
Oshia Oshia ferganica
Saurichthys Saurichthys orientalis
 
Sixtelia Sixtelia asiatica

Arthropods

Genus Species Description Notes Images
Gigatitan G. extensus A mantis-like predator with a wingspan of approximately 33 centimetres (13 in). It is the type genus of the family Gigatitanidae. [19]
 
G. magnificus
G. ovatus
G. similis
G. vulgaris
Aiban A. kichineis A member of Cnemidolestida/Cnemidolestodea (an extinct group of insects of uncertain phylogenetic placement, might be related to plecopterans or orthopterans) belonging to the family Sylvabestiidae [20]
Batkentak B. intactus A member of Grylloblattida/Eoblattida belonging to the family Daldubidae [21]
Chubakka C. madygensis A madygelline xyelid sawfly [22]
Locustoblattina L. marginata A member of Eoblattida belonging to the family Mesorthopteridae [23]
L. segmentata
Madygella M. aristovi A madygelline xyelid sawfly [22]
M. bashkuevi
M. kurochkini
M. levivenosa
Nestorembia N. novojilovi A webspinner belonging to the family Alexarasniidae [24]
N. shcherbakovi [23]
Paratitan P. reliquia A right wing belonging to Neopterygota [25]
Sharovites S. alexanderi A member of Grylloblattida/Eoblattida, belonging to the family Mesorthopteridae [26]
Shurabia S. tanga A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae [27]
Guillermia G. lecticula [28]
Madygenorhynchus M. multifidus
Thaumatomerope T. sogdiana
Prochoristella P. longa
Dilemmala D. specula
Sacvoyagea S. ventrosa
Nonescyta N. mala
Maguviopsis M. kotchnevi
Falcarta F. bella
Krendelia K. ansata
Cuanoma C. protracta
Phyllotexta P. latens
Sitechka S. perforata
Tingiopsis T. reticulata
Pelorisca P. connectens
Xamenophlebia X. ornata
Thuringoblatta T. sogdianensis
Sacvoyagea S. ventrosa
Protoblattogryllus P. variabilis [29]
Megablattogryllus M. austerus
M. pinguis
Metakhosara M. sharovi
Batkentak B. intactus
Austroidelia A. asiatica
A. nervosa
Mesoidelia M. faceta
Parastenaropodites P. fluxa
P. longiuscula
Baharellus B. madygensis
Anoblattogryllus A. fundatus
Costatoviblatta C. aenigmatosa
C. conjuncta
Paratitan P. reductus
Prototitan P. sharovi
Mesoedischia M. obliqua
Paragryllavus P. curvatus
Zagryllavus Z. elongatus
Hagloedischia H. primitiva
Voliopus V. ancestralis
Euvoliopus E. giganteus
Macrovoliopus M. declivis
Paravoliopus P. dorsalis
Voliopellus V. latus
Triassaga T. angusta
Zamaraga Z. reticulata
Haglomorpha H. martynovi
Modihagla M. ovalis
Dulcihagla D. mistshenkoi
Lyrohagla L. uvarovi
Sonohagla S. curta
Tinnihagla T. zeuneri
Dolichohagla D. longa
Locustavus L. problematicus
L. intermedius
L. minutus
Miolocustavus M. reductus
Brevilocustavus B. distinctus
Fritaniopsis F. brevicaulis
Sogdoblatta S. nana
S. porrecta
Thuringoblatta T. sogdianensis
Xiphopterum X. sharovi
Sharovoplana S. parallelica
Triassophasma T. intermedium
T. pusillum
Prochresmoda P. parva
Nestorembia N. novojilovi
Madygenophlebia M. bella
M. nana
Gorochovia G. individua
G. minuta
G. bifurca
Gorochoviella G. conjuncta
Pseudoliomopterites P. obscurus
Ideliopsina I. nana
I. stupenda
I. ornata
Pseudoshurabia P. pallidula
Peltosyne P. varyvrosa
Ofthalmopeltos O. synkritos
Obrienia O. illaetabilis
Triassochorista T. kirgizica
Parachorista P. arguta
P. immota
Choristopanorpa C. temperata
Thaumatomerope T. oligoneura
T. sogdiana
Mesageta M. rieki
Liassochorista L. utilis
Agetopanorpa A. deceptoria
Prochoristella P. longa
Mesopsyche M. ordinata
M. justa
M. tortiva
Psychotipa P. predicta
Vymrhyphus V. tuomikoskii
Gnomusca G. molecula
Asiocula A. lima
Fasolinka F. beckermigdisovae
Fulgobole F. evansi
Scytachile S. emeljanovi
Serpentivena S. tigrina
Coccavus C. supercubitus
Kennedya K. carpenteri
K. gracilis
Batkenia B. pusilla
Paurophlebia P. lepida
P. angusta
Neritophlebia N. elegans
N. vicina
N. longa
Mixophlebia M. mixta
Cyrtophlebia C. sinuosa
Zygophlebia Z. ramosa
Reisia R. sogdiana
Shurabia S. serrata
Mesoblattogryllus M. intermedius [19]
Protoblattogryllus P. asiaticus
P. variabilis
Megablattogryllus M. austerus
M. magister
Megakhosarodes M. paulivenosus
Metakhosara M. sharovi
Austroidelia A. asiatica
A. nervosa
Mesoidelia M. ignorata
M. faceta
Parastenaropodites P. longiuscula
P. fluxa
Locustoblattina L. segmentata
Dorniella D. primitiva
Baharellus B. madygensis
Baharellinus B. dimidiatus
B. pectinatus
Costatoviblatta C. aenigmatosa
C. similis
Ferganamadygenia F. plicata
Paratitan P. libelluloides
P. venosus
P. intermedius
P. latispeculum
P. modestus
Mesotitanodes M. tillyardi
M. similis
Mesotitan M. primitivus
Provitimia P. pectinata
Proshiella P. ramivenosa
Mesoedischia M. madygenica
Gryllacrimima G. perfecta
Madygenia M. orientalis
M. ovalis
Kashgarlimahmutia K. reducta
Proxenopterum P. primitivum
Axenopterum A. venosum
Ferganopterus F. clarus
Ferganopterodes F. reductus
Pteroferganodes P. rieki
Tuphella T. rohdendorfi
T. sharovi
Platyvoliopus P. maximus
Stenovoliopus S. elongatus
Zavoliopus Z. densus
Turkestania T. deviata
Triassaga T. tshorkuphlebioides
Eumaraga E. madygenica
Hagloptera H. intermedia
Archihagla A. tenuis
Proisfaroptera P. martynovi
Protshorkuphlebia P. triassica
P. similis
Dulcihagla D. beybienkoi
Lyrohagla L. pravdini
L. decipiens
Sonohagla S. saussurei
S. chopardi
Microhagla M. minuta
Dinohagla D. corrugata
Adzhajloutshella A. plana
Locustavus L. lanceolatus
L. problematicus
L. intermedius
L. deformatus
L. minutus
Dicronemoura D. acaulis
Tritaniella T. mera
T. synneura
Fritaniopsis F. brevicaulis
F. remota
Sogdoblatta S. nana
Thuringoblatta T. sogdianensis
Subioblatta S. madygenica
Xiphopterum X. curvatum
Sharovoplana S. affinis
Triassophasma T. brevipoda
T. intermedium
T. minutissimum
Prochresmoda P. minuta
Madygenophlebia M. bella
M. primitiva
Micromadygenophlebia M. obscura
Gorochovia G. individua
Ideliopsina I. nana
I. ornata
Madygenidelia M. conjuncta
Pseudoshurabia P. pallidula
Hadeocoleus H. gigas
H. pelopius
H. catachtonius
Triassocoleus T. tortulosus
Salebroferus S. confragosus
S. asper
Schizophoroides S. rugosus
Thnesidius T. xyphophorus
Catabrycus C. hoplites
Triaplus T. laticoxa
Dolichosyne D. rostrata
Ademosyne A. kirghizica
Cephalosyne C. capitata
Cupesia C. sepulta
Notocupes N. laticella
N. rostratus
N. tenuis
Cladochorista C. multivenosa
Prophilopotamus P. asiaticus
Triassochorista T. kirgizica
Parachorista P. asiatica
P. multivena
P. arguta
P. sana
Mesageta M. pertrita
M. ignava
Agetopanorpa A. consueta
Prochoristella P. longa
Mesopsyche M. shcherbakovi
M. justa
M. gentica
Psychotipa P. depicta
Nadiptera N. pulchella
Oryctoxyela O. anomala
Madygenius M. primitivus
Ferganoxyela F. sogdiana
F. destructa
Triassoxyela T. orycta
Nevicia N. imitans
Asiocula A. lima
Phyllotexta P. latens
Fulgobole F. evansi
Scytachile S. cf. emeljanovi
Tingiopsis T. reticulata
Coccavus C. supercubitus
Permonka P. unica
P. triassica
Permosialis P. triassica
Triassolestodes T. asiaticus
Terskeja T. pumilio
T. tenuis
Paurophlebia P. lepida
Cladophlebia C. parvula
C. brevis
Nonymophlebia N. venosa
Neritophlebia N. longa
Triadophlebia T. madygenica
T. minuta
T. magna
T. honesta
T. modica
Mitophlebia M. enormis
Xamenophlebia X. ornata
Zygophlebiella Z. curta
Mixophlebia M. mixta
Shurabia S. minuta
S. ferganensis
S. anomala
Maguviopsis M. kotchnevi [30]
Tingiopsis T. reticulata
Dolichosyne D. confragosa [31]
Rhabdocupes R. baculatus
Salebroferus S. confragosus
Asiocula A. lima [32]
Blattomerope B. polyneura
Cuanoma C. protracta
Fasolinka F. beckermigdisovae
Kirgizichorista K. larvata
Krendelia K. ansata
Lithocupes L. incertus
Maguviopsis M. kotchnevi
Megakhosarodes M. paulivenosus
Nonescyta N. mala
Obrienia O. ingurgata
O. kuscheli
Panorpaenigma P. aemulum
Parachorista P. religiosa
Phyllotexta P. latens
Sacvoyagea S. ventrosa
Siberioperla S. ovalis
Sitechka S. perforata
Sogdodromeus S. altus
Triaplus T. macroplatus
Megakhosarodes M. paulivenosus [33]
Choristopanorpa C. opinata [34]
Dzhajloutshella D. arcanum
Kuperwoodia K. benefica
Macrocatinius M. brachycephalus
Neotuphella N. minor
Pesus P. prognathus
Mesageta M. gigantea [35]
M. rieki [34]
Triassoxya T. novozhilovi [36]

Flora

Genus Species Description Notes Images
Mesenteriophyllum M. kotschnevii [14]
Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Insect fauna correlations

Progonocimicidae found in the formation are also recorded in the Carnian Los Rastros Formation of Argentina, the Norian Blackstone and Mount Crosby Formations of Australia, and the Norian to Rhaetian Tologoi Formation of Kazakhstan.[37] Permochoristidae are also known from the Carnian Potrerillos and Cacheuta Formations of Argentina, Huangshanjie Formation of China, the Norian Blackstone and Mount Crosby Formations of Australia; the Norian to Rhaetian Tologoi Formation of Kazakhstan, the Sinemurian Dzhil Formation of Kyrgyzstan and the Toarcian Posidonia Shale of Germany.[38]

Orthophlebia had a relatively broad distribution in the Late Triassic as it is also found in the Sinemurian Badaowan Formation of China and Dzhil Formation of Kyrgyzstan, the Pliensbachian Makarova Formation of Russia and Sulyukta Formation of Tajikistan; the Toarcian Whitby Mudstone Formation of England, Posidonia Shale of Germany, and Cheremkhovo Formation of Russia, and the Early Jurassic Kushmurun Formation of Kazakhstan.[39]

Haglidae were also recorded in the Koldzat and Tologoi Formations of Kazakhstan, in the Carnian Cacheutá Formation of Argentina, the Carnian to Norian Molteno Formation of South Africa and Lesotho, and the Norian Mount Crosby Formation of Australia.[40]

See also

Other Central Asian Lagerstätten

References

  1. ^ a b c d Fischer et al., 2011
  2. ^ Unwin et al., 2003, pp.177-186
  3. ^ Shcherbakov, 2008
  4. ^ Fischer et al., 2018, p.25
  5. ^ a b Berner et al., 2009, p.2
  6. ^ a b Fischer et al., 2007, p.42
  7. ^ Berner et al., 2009, p.5
  8. ^ Berner et al., 2009, p.6
  9. ^ Kelly, 2018, p.62
  10. ^ a b c Voigt et al., 2007, p.162
  11. ^ Schoch et al., 2010
  12. ^ Schoch et al., 2020
  13. ^ Alifanov & Kurochkin, 2011
  14. ^ a b Buchwitz & Voigt, 2012
  15. ^ Sharov, 1971
  16. ^ Tatarinov, 2005
  17. ^ Kogan et al., 2009, p.142
  18. ^ Franeck et al., 2013, p.54
  19. ^ a b Sharov, 1968
  20. ^ Aristov, 2018
  21. ^ Aristov, 2015
  22. ^ a b Kopylov, 2014
  23. ^ a b Aristov, 2017
  24. ^ Shcherbakov, 2015
  25. ^ Béthoux et al., 2010, p.9
  26. ^ Aristov & Storozhenko, 2013
  27. ^ Aristov & Sukatcheva, 2018
  28. ^ Shcherbakov, 2011
  29. ^ Aristov et al., 2009
  30. ^ Becker-Migdisova, 1953
  31. ^ Ponomarenko, 1969
  32. ^ Arnoldi et al., 1977
  33. ^ Storozhenko, 1993
  34. ^ a b Shcherbakov et al., 1995
  35. ^ Novokshonov, 1997
  36. ^ Gorochov, 2005
  37. ^ Kelly, 2018, p.190
  38. ^ Kelly, 2018, p.186
  39. ^ Kelly, 2018, p.185
  40. ^ Kelly, 2018, p.182

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madygen, formation, russian, madygen, svita, late, triassic, carnian, geologic, formation, lagerstätte, batken, regions, western, kyrgyzstan, with, minor, outcrops, neighboring, tajikistan, uzbekistan, conglomerates, sandstones, mudstones, thick, formation, we. The Madygen Formation Russian Madygen Svita is a Late Triassic Carnian geologic formation and Lagerstatte in the Batken and Osh Regions of western Kyrgyzstan with minor outcrops in neighboring Tajikistan and Uzbekistan The conglomerates sandstones and mudstones of the 560 m 1 840 ft thick formation were deposited in terrestrial lacustrine alluvial fluvial and deltaic environments Madygen FormationStratigraphic range Carnian 235 222 Ma PreꞒ Ꞓ O S D C P T J K Pg NTypeGeological formationOverliesCambrian to Carboniferous rocksThickness560 m 1 840 ft LithologyPrimaryConglomerate sandstoneOtherMudstoneLocationCoordinates40 06 N 70 12 E 40 1 N 70 2 E 40 1 70 2 Coordinates 40 06 N 70 12 E 40 1 N 70 2 E 40 1 70 2Approximate paleocoordinates41 12 N 60 36 E 41 2 N 60 6 E 41 2 60 6RegionBatken amp Osh RegionsCountry Kyrgyzstan Tajikistan UzbekistanExtentFergana Valley amp RangeType sectionNamed forMadygen villageNamed byEvgeny A KochnevMadygen Formation Kyrgyzstan The formation extending across the Fergana Valley and Fergana Range is unique for Central Asia as it represents one of the few known continental deposits and the Madygen Formation is renowned for the preservation of more than 20 000 fossil insects making it one of the richest Triassic Lagerstatten in the world Other vertebrate fossils as fish amphibians reptiles and synapsids have been recovered from the formation too as well as minor fossil flora The lake sediments of the Lagerstatte provided fossil cartilaginous fishes and their egg capsules and unusual Triassic reptiles like Sharovipteryx and Longisquama 1 2 The wide diversity of insect fossils was first discovered in the 1960s and first described by Russian paleontologist Aleksandr Sharov with a notable example being Gigatitan 3 Contents 1 Description 1 1 Petroleum geology 2 Paleontological significance 3 Fossil content 3 1 Amphibians 3 2 Reptiles 3 3 Synapsids 3 4 Cartilaginous fishes 3 5 Fishes 3 6 Arthropods 3 7 Flora 3 8 Insect fauna correlations 4 See also 5 References 5 1 BibliographyDescription Edit Paleogeography of the Late Triassic around 230 Ma The Madygen Formation was deposited north of the Paleo Tethys ocean The Madygen Formation is a 560 metres 1 840 ft 4 thick succession of predominantly siliciclastic rocks accumulated in a tectonically induced basin covering parts of the Fergana Range and Fergana Valley of Kyrgyzstan with minor outcrops in Tajikistan and Uzbekistan 5 The Late Triassic layers rest on top of Paleozoic basement with local Permo Triassic molasse sediments The section consists of mudstones sandstones conglomerates and fanglomerates This wide variety of siliciclastic rocks reflects the complex spatial and temporal pattern of depositional sub environments including alluvial fans sandflats swamps back swamp areas and littoral to profundal lake zones The fluvio lacustrine deposits of the Madygen Formation belong to one of only a few occurrences of continental Triassic beds in Central Asia 6 The formation was deposited during the Carnian Pluvial Event CPE a global humid event leading to high extinction levels of various groups globally The CPE led to anoxic conditions most notably in the South China Block The area where the Madygen Formation was deposited formed part of the Cimmerian microcontinent a slab of crust that collided with Laurasia during the Cimmerian orogeny in later Mesozoic times This orogeny led to the disappearance of the Paleo Tethys Ocean Petroleum geology Edit The formation grades from bottom to top from alluvial to fluvial into a thick succession of lacustrine mudstones followed by an alluvial package on top of which lacustrine fluvial deltaic and alluvial layers were deposited 5 The hydrocarbon potential of samples of the Madygen Formation ranges from poor to excellent The sediments containing more than 0 5 Total Organic Carbon TOC may be regarded as sources of gaseous hydrocarbons rather than of oil 7 The Hydrogen Index HI of outcrop samples reaches 100 and the maximum recorded maturity Ro is 0 8 8 Paleontological significance EditDuring the 1960s Russian paleontologists recovered an unusually rich fossil content in the type strata of the Madygen Formation including abundant macrophytes more than 20 000 insect remains 9 and unique small reptiles with well preserved soft tissue 6 Spirorbis like polychaete worm tubes crustaceans ostracods kazacharthrans malacostraca freshwater pelecypods and gastropods are known from shallow to deeper lake environments Non aquatic insects are among the most common fossil remains of the Madygen Formation These include representatives of the orders Ephemeroptera Odonata Notoptera Blattodea Titanoptera Ensifera Caelifera Rhynchota Auchenorrhyncha Stenorrhyncha Coleoptera Hymenoptera Trichoptera and Diptera Traces of insect larvae are preserved in near shore lake deposits 10 Fish remains mostly represent endemic genera assigned to the actinopterygian families Evenkiidae Oshia Palaeoniscidae Ferganiscus Sixtelia and Megaperleidus and Alvinia The actinopterygian Saurichthys and the dipnoan Asiatoceratodus are cosmopolitan taxa also recorded in the Madygen Formation Two distinctive elasmobranch egg capsule types i e Palaeoxyris indicating a small Lissodus or Lonchidion like hybodont shark and an indeterminate capsule type imply the presence of two different elasmobranch species which used the freshwater environments of the Madygen Formation as spawning grounds Tetrapods are known from a probably larval urodelan Triassurus a small procynosuchid cynodont Madysaurus a gliding reptile Sharovipteryx and the enigmatic diapsid Longisquama 10 Fossil content EditAmphibians Edit Genus Species Material Notes ImagesMadygenerpeton M pustulatus A chroniosuchid reptiliomorph 11 Triassurus T sixtelae A possible early caudate salamander known from fossil larvae 12 Reptiles Edit Genus Species Material Notes ImagesKyrgyzsaurus K bukhanchenkoi A single specimen preserving the front half of a skeleton and scale impressions A drepanosaur 13 Longisquama L insignis A reptile with an elongated row of scales along its back 14 Sharovipteryx S mirabilis A gliding prolacertiform 15 Synapsids Edit Genus Species Material Notes ImagesMadysaurus M sharovi A cynodont 16 Cartilaginous fishes Edit Genus Species Material Notes ImagesFayolia F sharovi An egg capsule likely belonging to a xenacanthid 1 Lonchidion L ferganensis Teeth and egg capsules A hybodontid shark 1 Palaeoxyris P alterna Teeth and egg capsules A hybodontid shark 1 Fishes Edit The following fish fossils were found in the formation 10 17 18 Genus Species ImagesAlvinia Alvinia serrataFerganiscus Ferganiscus osteolepisMegaperleidus Megaperleidus lissolepisOshia Oshia ferganicaSaurichthys Saurichthys orientalis Sixtelia Sixtelia asiaticaArthropods Edit Genus Species Description Notes ImagesGigatitan G extensus A mantis like predator with a wingspan of approximately 33 centimetres 13 in It is the type genus of the family Gigatitanidae 19 G magnificusG ovatusG similisG vulgarisAiban A kichineis A member of Cnemidolestida Cnemidolestodea an extinct group of insects of uncertain phylogenetic placement might be related to plecopterans or orthopterans belonging to the family Sylvabestiidae 20 Batkentak B intactus A member of Grylloblattida Eoblattida belonging to the family Daldubidae 21 Chubakka C madygensis A madygelline xyelid sawfly 22 Locustoblattina L marginata A member of Eoblattida belonging to the family Mesorthopteridae 23 L segmentataMadygella M aristovi A madygelline xyelid sawfly 22 M bashkueviM kurochkiniM levivenosaNestorembia N novojilovi A webspinner belonging to the family Alexarasniidae 24 N shcherbakovi 23 Paratitan P reliquia A right wing belonging to Neopterygota 25 Sharovites S alexanderi A member of Grylloblattida Eoblattida belonging to the family Mesorthopteridae 26 Shurabia S tanga A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae 27 Guillermia G lecticula 28 Madygenorhynchus M multifidusThaumatomerope T sogdianaProchoristella P longaDilemmala D speculaSacvoyagea S ventrosaNonescyta N malaMaguviopsis M kotchneviFalcarta F bellaKrendelia K ansataCuanoma C protractaPhyllotexta P latensSitechka S perforataTingiopsis T reticulataPelorisca P connectensXamenophlebia X ornataThuringoblatta T sogdianensisSacvoyagea S ventrosaProtoblattogryllus P variabilis 29 Megablattogryllus M austerusM pinguisMetakhosara M sharoviBatkentak B intactusAustroidelia A asiaticaA nervosaMesoidelia M facetaParastenaropodites P fluxaP longiusculaBaharellus B madygensisAnoblattogryllus A fundatusCostatoviblatta C aenigmatosaC conjunctaParatitan P reductusPrototitan P sharoviMesoedischia M obliquaParagryllavus P curvatusZagryllavus Z elongatusHagloedischia H primitivaVoliopus V ancestralisEuvoliopus E giganteusMacrovoliopus M declivisParavoliopus P dorsalisVoliopellus V latusTriassaga T angustaZamaraga Z reticulataHaglomorpha H martynoviModihagla M ovalisDulcihagla D mistshenkoiLyrohagla L uvaroviSonohagla S curtaTinnihagla T zeuneriDolichohagla D longaLocustavus L problematicusL intermediusL minutusMiolocustavus M reductusBrevilocustavus B distinctusFritaniopsis F brevicaulisSogdoblatta S nanaS porrectaThuringoblatta T sogdianensisXiphopterum X sharoviSharovoplana S parallelicaTriassophasma T intermediumT pusillumProchresmoda P parvaNestorembia N novojiloviMadygenophlebia M bellaM nanaGorochovia G individuaG minutaG bifurcaGorochoviella G conjunctaPseudoliomopterites P obscurusIdeliopsina I nanaI stupendaI ornataPseudoshurabia P pallidulaPeltosyne P varyvrosaOfthalmopeltos O synkritosObrienia O illaetabilisTriassochorista T kirgizicaParachorista P argutaP immotaChoristopanorpa C temperataThaumatomerope T oligoneuraT sogdianaMesageta M riekiLiassochorista L utilisAgetopanorpa A deceptoriaProchoristella P longaMesopsyche M ordinataM justaM tortivaPsychotipa P predictaVymrhyphus V tuomikoskiiGnomusca G moleculaAsiocula A limaFasolinka F beckermigdisovaeFulgobole F evansiScytachile S emeljanoviSerpentivena S tigrinaCoccavus C supercubitusKennedya K carpenteriK gracilisBatkenia B pusillaPaurophlebia P lepidaP angustaNeritophlebia N elegansN vicinaN longaMixophlebia M mixtaCyrtophlebia C sinuosaZygophlebia Z ramosaReisia R sogdianaShurabia S serrataMesoblattogryllus M intermedius 19 Protoblattogryllus P asiaticusP variabilisMegablattogryllus M austerusM magisterMegakhosarodes M paulivenosusMetakhosara M sharoviAustroidelia A asiaticaA nervosaMesoidelia M ignorataM facetaParastenaropodites P longiusculaP fluxaLocustoblattina L segmentataDorniella D primitivaBaharellus B madygensisBaharellinus B dimidiatusB pectinatusCostatoviblatta C aenigmatosaC similisFerganamadygenia F plicataParatitan P libelluloidesP venosusP intermediusP latispeculumP modestusMesotitanodes M tillyardiM similisMesotitan M primitivusProvitimia P pectinataProshiella P ramivenosaMesoedischia M madygenicaGryllacrimima G perfectaMadygenia M orientalisM ovalisKashgarlimahmutia K reductaProxenopterum P primitivumAxenopterum A venosumFerganopterus F clarusFerganopterodes F reductusPteroferganodes P riekiTuphella T rohdendorfiT sharoviPlatyvoliopus P maximusStenovoliopus S elongatusZavoliopus Z densusTurkestania T deviataTriassaga T tshorkuphlebioidesEumaraga E madygenicaHagloptera H intermediaArchihagla A tenuisProisfaroptera P martynoviProtshorkuphlebia P triassicaP similisDulcihagla D beybienkoiLyrohagla L pravdiniL decipiensSonohagla S saussureiS chopardiMicrohagla M minutaDinohagla D corrugataAdzhajloutshella A planaLocustavus L lanceolatusL problematicusL intermediusL deformatusL minutusDicronemoura D acaulisTritaniella T meraT synneuraFritaniopsis F brevicaulisF remotaSogdoblatta S nanaThuringoblatta T sogdianensisSubioblatta S madygenicaXiphopterum X curvatumSharovoplana S affinisTriassophasma T brevipodaT intermediumT minutissimumProchresmoda P minutaMadygenophlebia M bellaM primitivaMicromadygenophlebia M obscuraGorochovia G individuaIdeliopsina I nanaI ornataMadygenidelia M conjunctaPseudoshurabia P pallidulaHadeocoleus H gigasH pelopiusH catachtoniusTriassocoleus T tortulosusSalebroferus S confragosusS asperSchizophoroides S rugosusThnesidius T xyphophorusCatabrycus C hoplitesTriaplus T laticoxaDolichosyne D rostrataAdemosyne A kirghizicaCephalosyne C capitataCupesia C sepultaNotocupes N laticellaN rostratusN tenuisCladochorista C multivenosaProphilopotamus P asiaticusTriassochorista T kirgizicaParachorista P asiaticaP multivenaP argutaP sanaMesageta M pertritaM ignavaAgetopanorpa A consuetaProchoristella P longaMesopsyche M shcherbakoviM justaM genticaPsychotipa P depictaNadiptera N pulchellaOryctoxyela O anomalaMadygenius M primitivusFerganoxyela F sogdianaF destructaTriassoxyela T oryctaNevicia N imitansAsiocula A limaPhyllotexta P latensFulgobole F evansiScytachile S cf emeljanoviTingiopsis T reticulataCoccavus C supercubitusPermonka P unicaP triassicaPermosialis P triassicaTriassolestodes T asiaticusTerskeja T pumilioT tenuisPaurophlebia P lepidaCladophlebia C parvulaC brevisNonymophlebia N venosaNeritophlebia N longaTriadophlebia T madygenicaT minutaT magnaT honestaT modicaMitophlebia M enormisXamenophlebia X ornataZygophlebiella Z curtaMixophlebia M mixtaShurabia S minutaS ferganensisS anomalaMaguviopsis M kotchnevi 30 Tingiopsis T reticulataDolichosyne D confragosa 31 Rhabdocupes R baculatusSalebroferus S confragosusAsiocula A lima 32 Blattomerope B polyneuraCuanoma C protractaFasolinka F beckermigdisovaeKirgizichorista K larvataKrendelia K ansataLithocupes L incertusMaguviopsis M kotchneviMegakhosarodes M paulivenosusNonescyta N malaObrienia O ingurgataO kuscheliPanorpaenigma P aemulumParachorista P religiosaPhyllotexta P latensSacvoyagea S ventrosaSiberioperla S ovalisSitechka S perforataSogdodromeus S altusTriaplus T macroplatusMegakhosarodes M paulivenosus 33 Choristopanorpa C opinata 34 Dzhajloutshella D arcanumKuperwoodia K beneficaMacrocatinius M brachycephalusNeotuphella N minorPesus P prognathusMesageta M gigantea 35 M rieki 34 Triassoxya T novozhilovi 36 Flora Edit Genus Species Description Notes ImagesMesenteriophyllum M kotschnevii 14 Color keyTaxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text crossed out taxa are discredited Insect fauna correlations Edit Progonocimicidae found in the formation are also recorded in the Carnian Los Rastros Formation of Argentina the Norian Blackstone and Mount Crosby Formations of Australia and the Norian to Rhaetian Tologoi Formation of Kazakhstan 37 Permochoristidae are also known from the Carnian Potrerillos and Cacheuta Formations of Argentina Huangshanjie Formation of China the Norian Blackstone and Mount Crosby Formations of Australia the Norian to Rhaetian Tologoi Formation of Kazakhstan the Sinemurian Dzhil Formation of Kyrgyzstan and the Toarcian Posidonia Shale of Germany 38 Orthophlebia had a relatively broad distribution in the Late Triassic as it is also found in the Sinemurian Badaowan Formation of China and Dzhil Formation of Kyrgyzstan the Pliensbachian Makarova Formation of Russia and Sulyukta Formation of Tajikistan the Toarcian Whitby Mudstone Formation of England Posidonia Shale of Germany and Cheremkhovo Formation of Russia and the Early Jurassic Kushmurun Formation of Kazakhstan 39 Haglidae were also recorded in the Koldzat and Tologoi Formations of Kazakhstan in the Carnian Cacheuta Formation of Argentina the Carnian to Norian Molteno Formation of South Africa and Lesotho and the Norian Mount Crosby Formation of Australia 40 See also EditCarnian formations Denmark Hill Insect Bed insect bearing unit of Queensland Australia Candelaria Formation lacustrine formation of the Parana Basin Brazil Ischigualasto Formation alluvial and fluvial Lagerstatte of Argentina Los Rastros Formation insect bearing formation underlying the Ischigualasto Formation Chinle Formation fluvio lacustrine fossiliferous formation of the western United States Doswell Formation continental fossiliferous formation of the eastern United States Lossiemouth Sandstone fossiliferous formation of ScotlandOther Central Asian LagerstattenBissekty Formation Turonian formation of Uzbekistan Bostobe Formation Santonian to Campanian formation of Kazakhstan Karabastau Formation Late Jurassic formation of Kazakhstan Danata Formation Paleogene formation of TurkmenistanReferences Edit a b c d Fischer et al 2011 Unwin et al 2003 pp 177 186 Shcherbakov 2008 Fischer et al 2018 p 25 a b Berner et al 2009 p 2 a b Fischer et al 2007 p 42 Berner et al 2009 p 5 Berner et al 2009 p 6 Kelly 2018 p 62 a b c Voigt et al 2007 p 162 Schoch et al 2010 Schoch et al 2020 Alifanov amp Kurochkin 2011 a b Buchwitz amp Voigt 2012 Sharov 1971 Tatarinov 2005 Kogan et al 2009 p 142 Franeck et al 2013 p 54 a b Sharov 1968 Aristov 2018 Aristov 2015 a b Kopylov 2014 a b Aristov 2017 Shcherbakov 2015 Bethoux et al 2010 p 9 Aristov amp Storozhenko 2013 Aristov amp Sukatcheva 2018 Shcherbakov 2011 Aristov et al 2009 Becker Migdisova 1953 Ponomarenko 1969 Arnoldi et al 1977 Storozhenko 1993 a b Shcherbakov et al 1995 Novokshonov 1997 Gorochov 2005 Kelly 2018 p 190 Kelly 2018 p 186 Kelly 2018 p 185 Kelly 2018 p 182 Bibliography Edit Schoch Rainer R Ralf Werneburg and Sebastian Voigt 2020 A Triassic stem salamander from Kyrgyzstan and the origin of salamanders PNAS 117 11584 11588 Accessed 2020 05 27 Aristov D S and I D Sukatcheva 2018 New insects Insecta Trichoptera Reculida Eoblattida from the Mesozoic of Asia Paleontological Journal 52 4 405 413 Accessed 2020 05 27 doi 10 1134 S0031030118040032 Aristov D S 2018 New and little known cnemidolestid insects Insecta Cnemidolestida from the Middle Permian Middle Triassic of EurasiaPaleontological Journal 52 12 1381 1390 doi 10 1134 S0031030118120079 Fischer Jan Ilja Kogan Sebastian Voigt Michael Buchwitz Jorg W Schneider Philippe Moisan Frederik Spindler Andreas Brosig and Marvin Preusse Frank Scholze amp Ulf Linnemann 2018 The mid Triassic Madygen Lagerstatte Southwest Kyrgyzstan Central Asia 25 26 13th Symposium on Mesozoic Terrestrial Ecosystems and Biota Accessed 2020 05 27 Kelly Richard S 2018 Effects of environmental perturbation during the Late Triassic on the taxic diversity of British insects 1 337 University of Bristol Aristov D S 2017 New insects Insecta Eoblattida Embiida from the Permian of Russia and the Triassic of Kyrgyzstan with observations on the origin of webspinners Paleontological Journal 51 2 161 170 Accessed 2020 05 27 doi 10 1134 S0031030117020046 Aristov D S 2015 Classification of the order Eoblattida Insecta Blattidea with description of new taxa Far Eastern Entomologist 301 1 56 Accessed 2020 05 27 Shcherbakov Dmitry E 2015 Permian and Triassic ancestors of webspinners Embiodea Russian Entomological Journal 24 3 187 200 doi 10 15298 rusentj 24 3 01 Kopylov D S 2014 New sawflies of the subfamily Madygellinae Hymenoptera Xyelidae from the Middle Upper Triassic of KyrgyzstanPaleontological Journal 48 6 610 620 doi 10 1134 S0031030114060070 Aristov D S and S Yu Storozhenko 2013 A new genus of the family Mesorthopteridae Grylloblattida from the Triassic of Kyrgyzstan Far Eastern Entomologist 264 7 12 Accessed 2017 03 26 Franeck Franziska Jan Fischer Ilja Kogan Sebastian Voigt and Jorg W Schneider 2013 Microvertebrate analyses extend the ichthyodiversity of the continental Triassic Madygen Formation Kyrgyzstan in Palaeobiology and Geobiology of Fossil Lagerstatten through Earth History 52 54 Universitat Gottingen Accessed 2020 05 27 Buchwitz M and S Voigt 2012 The dorsal appendages of the Triassic reptile Longisquama insignis reconsideration of a controversial integument typePalaontologische Zeitschrift 86 3 313 331 doi 10 1007 s12542 012 0135 3 Alifanov V R and E N Kurochkin 2011 Kyrgyzsaurus bukhanchenkoi gen et sp nov a new reptile from the triassic of southwestern KyrgyzstanPaleontological Journal 45 6 639 647 doi 10 1134 S0031030111060025 Fischer J S Voigt J W Schneider M Buchwitz and S Voigt 2011 A selachian freshwater fauna from the Triassic of Kyrgyzstan and its implication for Mesozoic shark nurseriesJournal of Vertebrate Paleontology 31 5 937 953 doi 10 1080 02724634 2011 601729 Shcherbakov D E 2011 New and little known families of Hemiptera Cicadomorpha from the Triassic of Central Asian early analogs of treehoppers and planthoppersZootaxa 2836 1 26 Bethoux O S Voigt and J W Schneider 2010 A Triassic palaeodictyopteran from Kyrgyzstan Palaeodiversity 3 9 13 Accessed 2020 05 27 Buchwitz M and S Voigt 2010 Peculiar carapace structure of a Triassic chroniosuchian implies evolutionary shift in trunk flexibilityJournal of Vertebrate Paleontology 30 6 1697 1708 doi 10 1080 02724634 2010 521685 Schoch Rainer R Sebastian Voigt and Michael Buchwitz 2010 A chroniosuchid from the Triassic of Kyrgyzstan and analysis of chroniosuchian relationshipsZoological Journal of the Linnean Society 160 3 515 530 doi 10 1111 j 1096 3642 2009 00613 x Aristov D S T Wappler and A P Rasnitsyn 2009 New and Little Known Grylloblattids of the Family Geinitziidae Insecta Grylloblattida from the Triassic and Jurassic of Europe Asia and South AfricaPaleontological Journal 43 418 424 Berner Ulrich Georg Scheeder Jolanta Kus Sebastian Voigt and Jorg W Schneider 2009 Organic Geochemical Characterization of Terrestrial Source Rocks of the Triassic Madygen Formation Southern Tien Shan Kyrgyzstan 1 8 AAPG Convention Denver Accessed 2020 05 27 Kogan Ilja Katharina Schonberger Jan Fischer and Sebastian Voigt 2009 A nearly complete skeleton of Saurichthys orientalis Pisces Actinopterygii from the Madygen Formation Middle to Late Triassic Kyrgyzstan Central Asia preliminary results Freiberger Forschungshefte C532 139 152 Accessed 2020 05 27 Shcherbakov Dmitry 2008 Madygen Triassic Lagerstatte number one before and after Sharov Alavesia 2 5 125 131 Accessed 2020 05 27 Fischer Jan Sebastian Voigt and Michael Buchwitz 2007 First elasmobranch egg capsules from freshwater lake deposits of the Madygen Formation Middle to Late Triassic Kyrgyzstan Central Asia Freiberger Forschungshefte C526 41 46 Accessed 2020 05 27 Voigt Sebastian Frederik Spindler Jan Fischer Ilja Kogan and Michael Buchwitz 2007 An extraordinary lake basin the Madygen fossil lagerstaette Middle to Upper Triassic Kyrgyzstan Central Asia Wissenschaftliche Mitteilungen des Institutes fur Geologie der TU Bergakademie Freiberg 36 161 163 Accessed 2020 05 27 Tatarinov L P 2005 A new cynodont Reptilia Theriodontia from the Magygen Formation Triassic of Fergana KyrgyzstanPaleontological Journal 39 2 192 198 Gorochov A V 2005 Review of Triassic Orthoptera with Descriptions of New and Little Known Taxa Part 2Paleontological Journal 39 3 272 279 Unwin D M V R Alifanov and M J Benton 2003 Enigmatic small reptiles from the Middle Late Triassic of Kyrgyzstan in The Age of Dinosaurs in Russia and Mongolia 177 186 Cambridge University Press ISBN 9780521545822 Novokshonov V G 1997 New Triassic Scorpionflies Insecta Mecoptera Paleontological Journal 31 628 635 Shcherbakov D E E D Lukashevich and V A Blagoderov 1995 Triassic Diptera and initial radiation of the orderInternational Journal of Dipterological Research 6 75 115 Storozhenko S Y 1993 Reviziya semeystva Megakhosaridae Grylloblattida In A G Ponomarenko ed Mezozoyskie Nasekomye i Ostrakody Azii 100 112 Arnoldi L V V V Zherikhin L M Nikritin and A G Ponomarenko 1977 Mezozoiskie zhestkokryiyeAkademiya Nauk SSSR Trudy Paleontologicheskogo Instituta 161 1 204 Sharov A G 1971 Novye letayushche reptilii is Mesosoya Kazachstana i Kirgizii New Mesozoic flying reptiles from Kazakhstan and Kirgizia Trudy Paleontologicheskiya Instituta Akademiy Nauk SSSR 130 104 113 Ponomarenko A G 1969 Istoricheskoe Razvitie Zhestkokrylykh Arkhostemat Historical Development of the Archostomate Beetles Trudy Akademiya Nauk SSSR 125 1 240 Sharov A G 1968 Filogeniya ortopteroidnykh nasekomykhTrudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118 1 216 Becker Migdisova E E 1953 Dva predstavitelya poluzhestkokrylykh nasekomykh iz urochishcha MadygenDoklady Akademii Nauk SSSR 90 461 464 Retrieved from https en wikipedia org w index php title Madygen Formation amp oldid 1102298127, wikipedia, wiki, book, books, library,

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