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Lake Malawi

Lake Malawi, also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique, is an African Great Lake and the southernmost lake in the East African Rift system, located between Malawi, Mozambique and Tanzania.

Lake Malawi (Lake Nyasa)
View from orbit. North is in upper right corner
Lake Malawi (Lake Nyasa)
Lake Malawi (Lake Nyasa)
Coordinates12°00′S 34°30′E / 12.000°S 34.500°E / -12.000; 34.500
Lake typeAncient lake, Rift lake
Primary inflowsRuhuhu River[1]
Primary outflowsShire River[1]
Basin countriesMalawi, Mozambique, and Tanzania
Max. length560 km (350 mi)[1] to 580[2]
Max. width75 km (47 mi)[1]
Surface area29,600 km2 (11,400 sq mi)[1]
Average depth292 m (958 ft)[3]
Max. depth706 m (2,316 ft)[3]
Water volume8,640 km3 (2,070 cu mi)[3]
Surface elevation468 metres (1,535 ft) above sea level[4]
IslandsLikoma and Chizumulu islets, Mumbo Island
References[1][3]
Official nameLake Niassa and its Coastal Zone
Designated26 April 2011
Reference no.1964[5]

It is the fifth largest fresh water lake in the world by volume, the ninth largest lake in the world by area and the third largest and second deepest lake in Africa. Lake Malawi is home to more species of fish than any other lake in the world,[6] including at least 700 species of cichlids.[7] The Mozambique portion of the lake was officially declared a reserve by the Government of Mozambique on June 10, 2011,[8] while in Malawi a portion of the lake is included in Lake Malawi National Park.[6]

Lake Malawi is a meromictic lake, meaning that its water layers do not mix. The permanent stratification of Lake Malawi's water and the oxic-anoxic boundary are maintained by moderately small chemical and thermal gradients.[9]

Geography edit

Lake Malawi is between 560 kilometres (350 mi)[1] and 580 kilometres (360 mi) long,[2] and about 75 kilometres (47 mi) wide at its widest point. The lake has a total surface area of about 29,600 square kilometres (11,400 sq mi).[1] The lake is 706 m (2,316 ft) at its deepest point, located in a major depression in the north-central part.[10] Another smaller depression in the far north reaches a depth of 528 m (1,732 ft).[10] The southern half of the lake is shallower; less than 400 m (1,300 ft) in the south-central part and less than 200 m (660 ft) in the far south.[10] The lake has shorelines on western Mozambique, eastern Malawi, and southern Tanzania. The largest river flowing into it is the Ruhuhu River, and there is an outlet at its southern end, the Shire River, a tributary that flows into the very large Zambezi River in Mozambique.[2] Evaporation accounts for more than 80% of the water loss from the lake, considerably more than the outflowing Shire River.[11] The outflows from Lake Malawi into the Shire River are vital for the economy as the water resources support hydropower, irrigation and downstream biodiversity.[12] Concerns have been raised over the future climate change impacts of Lake Malawi due to the recent decline in lake levels and the overall drying trend.[13] The climate in the lake region is already experiencing changes, with the temperatures predicted to increase throughout the country.[14]

The lake is about 350 kilometres (220 mi) southeast of Lake Tanganyika, another of the great lakes of the East African Rift. [15]

The Lake Malawi National Park is located at the southern end of the lake.[16]

Geological history edit

 
Topographic profile of Lake Malawi's rift shoulder
 
The East African Rift (red) with the Rift Valley lakes, Malawi being in the south

Malawi is one of the major Rift Valley lakes and an ancient lake. The lake lies in a valley formed by the opening of the East African Rift, where the African tectonic plate is being split into two pieces. This is called a divergent plate tectonics boundary. Malawi has typically been estimated to be 1–2 million years old (mya),[17][18] but more recent evidence points to a considerably older lake with a basin that started to form about 8.6 mya and deep-water condition first appeared 4.5 mya.[19][20]

The water levels have varied dramatically over time, ranging from almost 600 m (2,000 ft) below current level[21] to 10–20 m (33–66 ft) above.[19] During periods the lake dried out almost completely, leaving only one or two relatively small, highly alkaline and saline lakes in what currently are Malawi's deepest parts.[19][21] A water chemistry resembling the current conditions only appeared about 60,000 years ago.[21] Major low-water periods are estimated to have occurred about 1.6 to 1.0–0.57 million years ago (where it might have dried out completely), 420,000 to 250,000–110,000 years ago,[19] about 25,000 years ago and 18,000–10,700 years ago.[20] During the peak of the low-water period between 1390 and 1860 AD, it may have been 120–150 m (390–490 ft) below current water levels.[18]

Water characteristics edit

The lake's water is alkaline (pH 7.7–8.6) and warm with a typical surface temperature between 24 and 29 °C (75–84 °F), while deep sections typically are about 22 °C (72 °F).[22] The thermocline is located at a depth of 40–100 m (130–330 ft).[11] The oxygen limit is at a depth of approximately 250 m (820 ft), effectively restricting fish and other aerobic organisms to the upper part.[23] The water is very clear for a lake and the visibility can be up to 20 m (66 ft), but slightly less than half this figure is more common and it is below 3 m (10 ft) in muddy bays.[10] However, during the rainy season months of January to March, the waters are more muddy due to muddy river inflows.[24]

European discovery and colonisation edit

The Portuguese trader Candido José da Costa Cardoso was the first European to visit the lake in 1846.[25] David Livingstone reached the lake in 1859, and named it Lake Nyasa.[2] He also referred to it by a pair of nicknames: Lake of Stars and Lake of Storms.[26] The Lake of Stars nickname came after Livingstone observed lights from the lanterns of the fishermen in Malawi on their boats, that resemble, from a distance, stars in the sky.[27] Later, after experiencing the unpredictable and extremely violent gales that sweep through the area, he also referred to it as the Lake of Storms.[27]

On 16 August 1914, Lake Malawi was the scene of a brief naval battle during World War I. The British gunboat SS Gwendolen, commanded by a Captain Rhoades, received orders from the British Empire's high command to "sink, burn, or destroy" the German Empire's only gunboat on the lake, the Hermann von Wissmann, commanded by Captain Berndt. Rhoades's crew found the Hermann von Wissmann in a bay near Sphinxhaven, in German East African territorial waters. Gwendolen disabled the German boat with a single cannon shot from a range of about 1,800 metres (2,000 yd). This brief conflict was hailed by The Times in England as the British Empire's first naval victory of World War I.[28][29]

Borders edit

 
Dashed line: current Malawi border
Dotted line: Tanzanian claim

Tanzania–Malawi dispute edit

The partition of the lake's surface area between Malawi and Tanzania is under dispute. Tanzania claims that the international border runs through the middle of the lake.[30] On the other hand, Malawi claims the whole of the surface of this lake that is not in Mozambique, including the waters that are next to the shoreline of Tanzania.[31] Both sides cite the Heligoland Treaty of 1890 between Great Britain and Germany concerning the border. The wrangle in this dispute occurred when the British colonial government, just after they had captured Tanganyika from Germany, placed all of the waters of the lake under a single jurisdiction, that of the territory of Nyasaland, without a separate administration for the Tanganyikan portion of the surface. Later in colonial times, two jurisdictions were established.[32]

The dispute came to a head in 1967 when Tanzania officially protested to Malawi; however nothing was settled.[33] Occasional flare-ups of conflict occurred during the 1990s and in the 21st century.[34] In 2012, Malawi's oil exploration initiative brought the issue to the fore, with Tanzania demanding that exploration cease until the dispute was settled.[35]

Malawi–Mozambique border edit

In 1954, an agreement was signed between the British and the Portuguese making the middle of the lake their boundary with the exception of Chizumulu Island and Likoma Island, which were kept by the British and are now part of Malawi.[32]

Transport edit

 
A jetty juts into the lake at Nkhata Bay

MV Chauncy Maples began service on the lake in 1901 as the SS Chauncy Maples: a floating clinic and church for the Universities' Mission to Central Africa. She later served as a ferry and is currently being renovated into a mobile clinic at Monkey Bay. The renovation was expected to be complete during the first half of 2014, but was halted in 2017.[36] MV Mpasa entered service in 1935.[37] The ferry MV Ilala entered service in 1951. In recent years she has often been out of service, but when operational she runs between Monkey Bay at the southern end of the lake to Karonga on the northern end, and occasionally to the Iringa Region of Tanzania. The ferry MV Mtendere entered service in 1980.[37] By 1982 she was carrying 100,000 passengers each year.,[37] but as of 2014 she was out of service.[38] She normally serves the southern part of the lake but if Ilala was out of service she operated the route to Karonga. The Tanzanian ferry MV Songea was built in 1988.[39] Her operator was the Tanzania Railway Corporation Marine Division until 1997, when it became the Marine Services Company Limited.[40] Songea plies weekly between Liuli and Nkhata Bay via Itungi and Mbamba Bay.[39] The worst Lake disaster was the sinking of the MV Vipya in 1946, which resulted in 145 deaths. [41]

Wildlife edit

 
An African fish eagle catching a fish in Lake Malawi

Wildlife found in and around Lake Malawi or Nyasa includes Nile crocodiles, hippopotamus, monkeys,[42] and a significant population of African fish eagles that feed off fish from the lake.[43]

Fish edit

Fishing edit

 
Lake Malawi sardines (Engraulicypris sardella) spread out to dry on the shore of the lake

Lake Malawi has for millennia provided a major food source to the residents of its shores since its waters are rich in fish. Among the most popular are the four species of chambo, consisting of any one of four species in the subgenus Nyasalapia (Oreochromis karongae, O. lidole, O. saka and O. squamipinnis), as well as the closely related O. shiranus.[44] Other species that support important fisheries include the Lake Malawi sardine (Engraulicypris sardella) and the large kampango catfish (Bagrus meridionalis).[10] Most fishing provides food for the increasing human population near the lake, but some are exported from Malawi. The wild population of fish is increasingly threatened by overfishing and water pollution.[45][46] A drop in the lake's water level represents another threat, and is believed to be driven by water extraction by the increasing human population, climate change and deforestation.[46] The chambo and kampango have been particularly overfished (the kampango declined by about 90% from 2006 to 2016,[47] O. karongae and O. squamipinnis by about 94%, and O. lidole might already be extinct[48][49]) and they are now seriously threatened.[50] The IUCN recognises 117 species of Malawi cichlids as threatened; some of these have tiny ranges and may be restricted to rocky coastlines only a few hundred metres long.[51]

Malawi cichlids
 
 
 
 
 
 
 
Top to bottom:

1. Diplotaxodon, one of the very few cichlid genera that occurs offshore in relatively deep water.[23]
2. Nimbochromis livingstonii is a piscivorous hap that is famous for playing dead to lure prey close.[10][52][53]
3. As typical of utaka, Copadichromis azureus has bright blue males (shown) and duller females that are silvery with dark spots.[54]
4. Aulonocara stuartgranti is part of a group of relatively peaceful species popularly known as peacock cichlids.[55]
5. Fossorochromis rostratus is an "aberrant" hap that often sifts mouthfuls of sand to extract small food organisms.[10][56]
6. Like many mbuna, Pseudotropheus saulosi is a small cichlid where both male (blue and black) and female (yellow) are colorful.[57]

7. Oreochromis squamipinnis is one of only six native tilapia species in the lake, but these are important to fisheries. Notice the white genital tassels, longer when fully extended and unique to male chambo.[10][44]

Cichlids edit

Lake Malawi is noted for being the site of evolutionary radiations among several groups of animals, most notably cichlid fish.[58] There are at least 700 cichlid species in Lake Malawi,[7] with some estimating that the actual figure is as high as 1,000 species.[8][59] The actual number is labelled with some uncertainty because of the many undescribed species and the extreme variation among some species, making the task of delimiting them very complex.[7][10] Except for four species (Astatotilapia calliptera, Coptodon rendalli, Oreochromis shiranus and Serranochromis robustus), all cichlids in the lake are endemic to the Malawi system, which also includes nearby smaller Lake Malombe and the upper Shire River.[10][60][61] Many of these have become popular among aquarium owners due to their bright colors. Recreating a Lake Malawi biotope to host cichlids became quite popular in the aquarium hobby.[62] Most Malawi cichlids are found in relatively shallow coastal waters,[10] but Diplotaxodon has been recorded down to depths of 200–220 m (660–720 ft) and several (especially Diplotaxodon, Rhamphochromis and Copadichromis quadrimaculatus) are known from pelagic waters.[23]

The cichlids of the lake are divided into two groups and the vast majority of the species are haplochromines. The sister species to the Malawi haplochromines is Astatotilapia sp. Ruaha (a currently undescribed species from Great Ruaha River), and these two separated between 2.13 and 6.76 million years ago (mya).[63] The earliest divergence within the Malawi haplochromines occurred between 1.20 and 4.06 mya,[63] but most radiations in this group are far younger; in extreme cases species may have diverged only a few hundred years ago.[18] The Malawi haplochromines are mouthbrooders, but otherwise vary extensively in general behaviour and ecology.[10] Within the Malawi haplochromines there are two main groups, the haps and the mbuna. The haps (they were formerly included in Haplochromis) can be further subdivided into three subgroups: The relatively large, often more than 20 cm (8 in) long, and aggressive piscivores that roam various habitats in pursuit of prey, the open-water (although often not far from sand or rocks) utaka that feed in schools on zooplankton and typically are of medium size, and finally a subgroup of "aberrant" species that essentially are defined by them not fitting clearly into the other subgroups.[10][53][64] Adult male haps generally display bright colors, while juveniles of both sexes and adult females typically show a silvery or grey coloration with sometimes irregular black bars or other markings.[10][53] The second main haplochromine group are the mbuna, a name used both locally and popularly, which means "rockfish" in Tonga.[65] They are found at rocky outcrops, territorially aggressive (although commonly found in high densities) and often specialised aufwuchs feeders.[10][53] The mbuna species tend to be relatively small, mostly less than 13 cm (5 in) long, and often both sexes are brightly colored with males having egg-shaped yellow spots on their anal fin (a feature particularly prevalent in the mbuna, but not exclusive to this group).[10][53]

The second group, the tilapia, comprises only six species in two genera in Lake Malawi: The redbreast tilapia (Coptodon rendalli), a widespread African species, is the only substrate-spawning cichlid in the lake.[10][66] This large cichlid mainly feeds on macrophytes.[10][67] The remaining are five mouthbrooding species of Oreochromis; four chambo in the subgenus Nyasalapia (O. karongae, O. lidole, O. saka and O. squamipinnis) that are endemic to the Lake Malawi system, as well as the closely related O. shiranus, which also is found in Lake Chilwa.[10][44][60] The Malawi Oreochromis mainly feed on phytoplankton, reach lengths up to 26–42 cm (10–17 in) depending on the exact species, and are mostly black or silvery-gray with relatively indistinct dark bars.[10][60][68] Male chambo have unique genital tassels when breeding, which aid in egg fertilisation in a manner comparable to the egg-spots on the anal fin of haplochromines.[10][44]

Non-cichlids edit

 
The kampango (Bagrus meridionalis), one of the largest catfish, reaching up to 1.5 m (4.9 ft) in length[69]

The vast majority of the fish species in the lake are cichlids. Among the non-cichlid native fish are several species of cyprinids (in genera Barbus, Labeo and Opsaridium, and the Lake Malawi sardine Engraulicypris sardella), airbreathing catfish (Bathyclarias and Clarias, and the kampango Bagrus meridionalis), mochokid catfish (Chiloglanis and Malawi squeaker Synodontis njassae), Mastacembelus spiny eel, mormyrids (Marcusenius, Mormyrops and Petrocephalus), the African tetra Brycinus imberi, the poeciliid Aplocheilichthys johnstoni, the spotted killifish (Nothobranchius orthonotus), and the mottled eel (Anguilla nebulosa).[10]

At a genus level, most of these are widespread in Africa, but Bathyclarias is entirely restricted to the lake.[70]

Invertebrates edit

Molluscs edit

Lake Malawi is home to 28 species of freshwater snails (including 16 endemics) and 9 bivalves (2 endemics, Aspatharia subreniformis and the unionid Nyassunio nyassaensis).[71][72] The endemic freshwater snails are all members of the genera Bellamya, Bulinus, Gabbiella, Lanistes and Melanoides.[73]

Lake Malawi is home to a total of four snail species in the genus Bulinus, which is a known intermediate host of bilharzia. A survey in Monkey Bay in 1964 found two endemic species of snails of the genus (B. nyassanus and B. succinoides) in the lake, and two non-endemic species (B. globosus and B. forskalli) in lagoons separated from it. The latter species are known intermediate hosts of bilharzia, and larvae of the parasite were detected in water containing these, but in experiments C. Wright of the British Museum of Natural History was unable to infect the two species endemic to the lake with the parasites. The field workers, who spent many hours on and in the lake, did not find either B. globosus or B. forskalli in the lake itself.[74] More recently, the disease has become a problem in the lake itself as the endemic B. nyassanus has become an intermediate host. This change, first noticed in the mid-1980s, is possibly related to a decline in snail-eating cichlids (for example, Trematocranus placodon) due to overfishing and/or a new strain of the bilharzia parasite.[22]

Crustaceans edit

Unlike Lake Tanganyika with its many endemic freshwater crabs and shrimp, there are few such species in Lake Malawi. The Malawi blue crab, Potamonautes lirrangensis (syn. P. orbitospinus), is the only crab in the lake and it is not endemic.[75][76] The atyid shrimp Caridina malawensis is endemic to the lake, but it is poorly known and has historically been confused with C. nilotica, which is not found in the lake.[77] Pelagic zooplanktonic species include two cladocerans (Diaphanosoma excisum and Bosmina longirostris), three copepods (Tropodiaptomus cunningtoni, Thermocyclops neglectus and Mesocyclops aequatorialis),[78] and several ostracods (including both described and undescribed species).[79]

Lake flies edit

 
Huge swarms of lake flies (Chaoborus edulis), resembling distant plumes of smoke over the water

Lake Malawi is famous for the huge swarms of tiny, harmless lake flies, Chaoborus edulis.[80] These swarms, typically appearing far out over water, can be mistaken for plumes of smoke and were also noticed by David Livingstone when he visited the lake.[80][81][82] The aquatic larvae feed on zooplankton, spending the day at the bottom and the night in the upper water levels.[80] When they pupate they float to the surface and transform into adult flies.[82] The adults are very short-lived and the swarms, which can be several hundred metres tall and often have a spiraling shape, are part of their mating behaviour.[80][83] They lay their eggs at the water's surface and the adults die.[83] The larvae are an important food source for fish,[78][80][84] and the adult flies are important both to birds and local people, who collect them to make kungu cakes/burgers, a local delicacy with a very high protein content.[81][82]

2015 mine leak edit

In January 2015, a sediment control tank collapsed at the Paladin Energy-owned uranium mine in Northern Malawi after a high intensity rain storm hit the area. It was revealed that approximately 50 litres of non radioactive material leaked into a local creek. Despite reports in local media of radioactive contamination the government conducted independent scientific tests on the local river system and found that there was no effect on the environment.[85][86]

Swimming edit

The 25 km solo swim across Lake Malawi between Cape Ngomba and Senga Bay has been accomplished on 5 occasions by 16 swimmers

1992: Lewis Pugh 9hrs 52 minutes (UK/SA)[87] and Otto Thanning (SA) 10hrs 5 minutes

2010: Abigail Brown (UK) 9hrs 45 minutes[88]

2013: Milko van Gool (Netherlands) 8hrs 46 minutes[89] and Kaitlin Harthoorn (US) 9hrs 17 minutes

2016: (current record) Jean Craven (SA), Robert Dunford (Kenya), Michiel Le Roux (SA), Samantha Whelpton (SA), Greig Bannatyne (SA), Haydn Von Maltitz (SA), Douglas Livingstone-Blevins (SA) 7hrs 53 mins [90]

2019: Chris Stapley (Eswatini) and Jay Azran (SA) 8hrs 40 minutes, Andrew Stevens (Australia) 10hrs 50 minutes, and Ruth Azran (SA) 11hrs 8 minutes.[91][92]

In 2019, Martin Hobbs (SA), became the first person to swim the full length of Lake Malawi (54 days), as well as setting the world record for longest solo swim in a lake[93]

See also edit

References edit

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Further reading edit

  • Mayall, James (December 1973). "The Malawi-Tanzania Boundary Dispute". The Journal of Modern African Studies. 11 (4): 611–628. doi:10.1017/S0022278X00008776. S2CID 154785268.
  • Recent study on Lake Malawi water levels reveals drought 100,000 years ago
  • "Freshwater Fish Species in Lake Malawi (Nyasa) [Southeast Africa]". Mongabay. Retrieved 9 December 2016.
  • Growing up in a Border District and Resolving the Tanzania-Malawi Lake Dispute: Compromise and concessions, by Godfrey Mwakikagile, African Renaissance Press, 2022

lake, malawi, other, uses, disambiguation, also, known, lake, nyasa, tanzania, lago, niassa, mozambique, african, great, lake, southernmost, lake, east, african, rift, system, located, between, malawi, mozambique, tanzania, lake, nyasa, view, from, orbit, nort. For other uses see Lake Malawi disambiguation Lake Malawi also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique is an African Great Lake and the southernmost lake in the East African Rift system located between Malawi Mozambique and Tanzania Lake Malawi Lake Nyasa View from orbit North is in upper right cornerLake Malawi Lake Nyasa Show map of MalawiLake Malawi Lake Nyasa Show map of AfricaCoordinates12 00 S 34 30 E 12 000 S 34 500 E 12 000 34 500Lake typeAncient lake Rift lakePrimary inflowsRuhuhu River 1 Primary outflowsShire River 1 Basin countriesMalawi Mozambique and TanzaniaMax length560 km 350 mi 1 to 580 2 Max width75 km 47 mi 1 Surface area29 600 km2 11 400 sq mi 1 Average depth292 m 958 ft 3 Max depth706 m 2 316 ft 3 Water volume8 640 km3 2 070 cu mi 3 Surface elevation468 metres 1 535 ft above sea level 4 IslandsLikoma and Chizumulu islets Mumbo IslandReferences 1 3 Ramsar WetlandOfficial nameLake Niassa and its Coastal ZoneDesignated26 April 2011Reference no 1964 5 It is the fifth largest fresh water lake in the world by volume the ninth largest lake in the world by area and the third largest and second deepest lake in Africa Lake Malawi is home to more species of fish than any other lake in the world 6 including at least 700 species of cichlids 7 The Mozambique portion of the lake was officially declared a reserve by the Government of Mozambique on June 10 2011 8 while in Malawi a portion of the lake is included in Lake Malawi National Park 6 Lake Malawi is a meromictic lake meaning that its water layers do not mix The permanent stratification of Lake Malawi s water and the oxic anoxic boundary are maintained by moderately small chemical and thermal gradients 9 Contents 1 Geography 2 Geological history 3 Water characteristics 4 European discovery and colonisation 5 Borders 5 1 Tanzania Malawi dispute 5 2 Malawi Mozambique border 6 Transport 7 Wildlife 7 1 Fish 7 1 1 Fishing 7 1 2 Cichlids 7 1 3 Non cichlids 7 2 Invertebrates 7 2 1 Molluscs 7 2 2 Crustaceans 7 2 3 Lake flies 8 2015 mine leak 9 Swimming 10 See also 11 References 12 Further readingGeography editLake Malawi is between 560 kilometres 350 mi 1 and 580 kilometres 360 mi long 2 and about 75 kilometres 47 mi wide at its widest point The lake has a total surface area of about 29 600 square kilometres 11 400 sq mi 1 The lake is 706 m 2 316 ft at its deepest point located in a major depression in the north central part 10 Another smaller depression in the far north reaches a depth of 528 m 1 732 ft 10 The southern half of the lake is shallower less than 400 m 1 300 ft in the south central part and less than 200 m 660 ft in the far south 10 The lake has shorelines on western Mozambique eastern Malawi and southern Tanzania The largest river flowing into it is the Ruhuhu River and there is an outlet at its southern end the Shire River a tributary that flows into the very large Zambezi River in Mozambique 2 Evaporation accounts for more than 80 of the water loss from the lake considerably more than the outflowing Shire River 11 The outflows from Lake Malawi into the Shire River are vital for the economy as the water resources support hydropower irrigation and downstream biodiversity 12 Concerns have been raised over the future climate change impacts of Lake Malawi due to the recent decline in lake levels and the overall drying trend 13 The climate in the lake region is already experiencing changes with the temperatures predicted to increase throughout the country 14 The lake is about 350 kilometres 220 mi southeast of Lake Tanganyika another of the great lakes of the East African Rift 15 The Lake Malawi National Park is located at the southern end of the lake 16 nbsp Lake Malawi 1967 nbsp Mwaya Beach nbsp Beach at Cape Maclear near Monkey BayGeological history edit nbsp Topographic profile of Lake Malawi s rift shoulder nbsp The East African Rift red with the Rift Valley lakes Malawi being in the southMalawi is one of the major Rift Valley lakes and an ancient lake The lake lies in a valley formed by the opening of the East African Rift where the African tectonic plate is being split into two pieces This is called a divergent plate tectonics boundary Malawi has typically been estimated to be 1 2 million years old mya 17 18 but more recent evidence points to a considerably older lake with a basin that started to form about 8 6 mya and deep water condition first appeared 4 5 mya 19 20 The water levels have varied dramatically over time ranging from almost 600 m 2 000 ft below current level 21 to 10 20 m 33 66 ft above 19 During periods the lake dried out almost completely leaving only one or two relatively small highly alkaline and saline lakes in what currently are Malawi s deepest parts 19 21 A water chemistry resembling the current conditions only appeared about 60 000 years ago 21 Major low water periods are estimated to have occurred about 1 6 to 1 0 0 57 million years ago where it might have dried out completely 420 000 to 250 000 110 000 years ago 19 about 25 000 years ago and 18 000 10 700 years ago 20 During the peak of the low water period between 1390 and 1860 AD it may have been 120 150 m 390 490 ft below current water levels 18 Water characteristics editThe lake s water is alkaline pH 7 7 8 6 and warm with a typical surface temperature between 24 and 29 C 75 84 F while deep sections typically are about 22 C 72 F 22 The thermocline is located at a depth of 40 100 m 130 330 ft 11 The oxygen limit is at a depth of approximately 250 m 820 ft effectively restricting fish and other aerobic organisms to the upper part 23 The water is very clear for a lake and the visibility can be up to 20 m 66 ft but slightly less than half this figure is more common and it is below 3 m 10 ft in muddy bays 10 However during the rainy season months of January to March the waters are more muddy due to muddy river inflows 24 European discovery and colonisation editThe Portuguese trader Candido Jose da Costa Cardoso was the first European to visit the lake in 1846 25 David Livingstone reached the lake in 1859 and named it Lake Nyasa 2 He also referred to it by a pair of nicknames Lake of Stars and Lake of Storms 26 The Lake of Stars nickname came after Livingstone observed lights from the lanterns of the fishermen in Malawi on their boats that resemble from a distance stars in the sky 27 Later after experiencing the unpredictable and extremely violent gales that sweep through the area he also referred to it as the Lake of Storms 27 On 16 August 1914 Lake Malawi was the scene of a brief naval battle during World War I The British gunboat SS Gwendolen commanded by a Captain Rhoades received orders from the British Empire s high command to sink burn or destroy the German Empire s only gunboat on the lake the Hermann von Wissmann commanded by Captain Berndt Rhoades s crew found the Hermann von Wissmann in a bay near Sphinxhaven in German East African territorial waters Gwendolen disabled the German boat with a single cannon shot from a range of about 1 800 metres 2 000 yd This brief conflict was hailed by The Times in England as the British Empire s first naval victory of World War I 28 29 Borders edit nbsp Dashed line current Malawi border Dotted line Tanzanian claimTanzania Malawi dispute edit The partition of the lake s surface area between Malawi and Tanzania is under dispute Tanzania claims that the international border runs through the middle of the lake 30 On the other hand Malawi claims the whole of the surface of this lake that is not in Mozambique including the waters that are next to the shoreline of Tanzania 31 Both sides cite the Heligoland Treaty of 1890 between Great Britain and Germany concerning the border The wrangle in this dispute occurred when the British colonial government just after they had captured Tanganyika from Germany placed all of the waters of the lake under a single jurisdiction that of the territory of Nyasaland without a separate administration for the Tanganyikan portion of the surface Later in colonial times two jurisdictions were established 32 The dispute came to a head in 1967 when Tanzania officially protested to Malawi however nothing was settled 33 Occasional flare ups of conflict occurred during the 1990s and in the 21st century 34 In 2012 Malawi s oil exploration initiative brought the issue to the fore with Tanzania demanding that exploration cease until the dispute was settled 35 Malawi Mozambique border edit In 1954 an agreement was signed between the British and the Portuguese making the middle of the lake their boundary with the exception of Chizumulu Island and Likoma Island which were kept by the British and are now part of Malawi 32 Transport edit nbsp A jetty juts into the lake at Nkhata BayMV Chauncy Maples began service on the lake in 1901 as the SS Chauncy Maples a floating clinic and church for the Universities Mission to Central Africa She later served as a ferry and is currently being renovated into a mobile clinic at Monkey Bay The renovation was expected to be complete during the first half of 2014 but was halted in 2017 36 MV Mpasa entered service in 1935 37 The ferry MV Ilala entered service in 1951 In recent years she has often been out of service but when operational she runs between Monkey Bay at the southern end of the lake to Karonga on the northern end and occasionally to the Iringa Region of Tanzania The ferry MV Mtendere entered service in 1980 37 By 1982 she was carrying 100 000 passengers each year 37 but as of 2014 she was out of service 38 She normally serves the southern part of the lake but if Ilala was out of service she operated the route to Karonga The Tanzanian ferry MV Songea was built in 1988 39 Her operator was the Tanzania Railway Corporation Marine Division until 1997 when it became the Marine Services Company Limited 40 Songea plies weekly between Liuli and Nkhata Bay via Itungi and Mbamba Bay 39 The worst Lake disaster was the sinking of the MV Vipya in 1946 which resulted in 145 deaths 41 Wildlife edit nbsp An African fish eagle catching a fish in Lake MalawiWildlife found in and around Lake Malawi or Nyasa includes Nile crocodiles hippopotamus monkeys 42 and a significant population of African fish eagles that feed off fish from the lake 43 Fish edit Fishing edit nbsp Lake Malawi sardines Engraulicypris sardella spread out to dry on the shore of the lakeLake Malawi has for millennia provided a major food source to the residents of its shores since its waters are rich in fish Among the most popular are the four species of chambo consisting of any one of four species in the subgenus Nyasalapia Oreochromis karongae O lidole O saka and O squamipinnis as well as the closely related O shiranus 44 Other species that support important fisheries include the Lake Malawi sardine Engraulicypris sardella and the large kampango catfish Bagrus meridionalis 10 Most fishing provides food for the increasing human population near the lake but some are exported from Malawi The wild population of fish is increasingly threatened by overfishing and water pollution 45 46 A drop in the lake s water level represents another threat and is believed to be driven by water extraction by the increasing human population climate change and deforestation 46 The chambo and kampango have been particularly overfished the kampango declined by about 90 from 2006 to 2016 47 O karongae and O squamipinnis by about 94 and O lidole might already be extinct 48 49 and they are now seriously threatened 50 The IUCN recognises 117 species of Malawi cichlids as threatened some of these have tiny ranges and may be restricted to rocky coastlines only a few hundred metres long 51 Malawi cichlids nbsp nbsp nbsp nbsp nbsp nbsp nbsp Top to bottom 1 Diplotaxodon one of the very few cichlid genera that occurs offshore in relatively deep water 23 2 Nimbochromis livingstonii is a piscivorous hap that is famous for playing dead to lure prey close 10 52 53 3 As typical of utaka Copadichromis azureus has bright blue males shown and duller females that are silvery with dark spots 54 4 Aulonocara stuartgranti is part of a group of relatively peaceful species popularly known as peacock cichlids 55 5 Fossorochromis rostratus is an aberrant hap that often sifts mouthfuls of sand to extract small food organisms 10 56 6 Like many mbuna Pseudotropheus saulosi is a small cichlid where both male blue and black and female yellow are colorful 57 7 Oreochromis squamipinnis is one of only six native tilapia species in the lake but these are important to fisheries Notice the white genital tassels longer when fully extended and unique to male chambo 10 44 Cichlids edit Lake Malawi is noted for being the site of evolutionary radiations among several groups of animals most notably cichlid fish 58 There are at least 700 cichlid species in Lake Malawi 7 with some estimating that the actual figure is as high as 1 000 species 8 59 The actual number is labelled with some uncertainty because of the many undescribed species and the extreme variation among some species making the task of delimiting them very complex 7 10 Except for four species Astatotilapia calliptera Coptodon rendalli Oreochromis shiranus and Serranochromis robustus all cichlids in the lake are endemic to the Malawi system which also includes nearby smaller Lake Malombe and the upper Shire River 10 60 61 Many of these have become popular among aquarium owners due to their bright colors Recreating a Lake Malawi biotope to host cichlids became quite popular in the aquarium hobby 62 Most Malawi cichlids are found in relatively shallow coastal waters 10 but Diplotaxodon has been recorded down to depths of 200 220 m 660 720 ft and several especially Diplotaxodon Rhamphochromis and Copadichromis quadrimaculatus are known from pelagic waters 23 The cichlids of the lake are divided into two groups and the vast majority of the species are haplochromines The sister species to the Malawi haplochromines is Astatotilapia sp Ruaha a currently undescribed species from Great Ruaha River and these two separated between 2 13 and 6 76 million years ago mya 63 The earliest divergence within the Malawi haplochromines occurred between 1 20 and 4 06 mya 63 but most radiations in this group are far younger in extreme cases species may have diverged only a few hundred years ago 18 The Malawi haplochromines are mouthbrooders but otherwise vary extensively in general behaviour and ecology 10 Within the Malawi haplochromines there are two main groups the haps and the mbuna The haps they were formerly included in Haplochromis can be further subdivided into three subgroups The relatively large often more than 20 cm 8 in long and aggressive piscivores that roam various habitats in pursuit of prey the open water although often not far from sand or rocks utaka that feed in schools on zooplankton and typically are of medium size and finally a subgroup of aberrant species that essentially are defined by them not fitting clearly into the other subgroups 10 53 64 Adult male haps generally display bright colors while juveniles of both sexes and adult females typically show a silvery or grey coloration with sometimes irregular black bars or other markings 10 53 The second main haplochromine group are the mbuna a name used both locally and popularly which means rockfish in Tonga 65 They are found at rocky outcrops territorially aggressive although commonly found in high densities and often specialised aufwuchs feeders 10 53 The mbuna species tend to be relatively small mostly less than 13 cm 5 in long and often both sexes are brightly colored with males having egg shaped yellow spots on their anal fin a feature particularly prevalent in the mbuna but not exclusive to this group 10 53 The second group the tilapia comprises only six species in two genera in Lake Malawi The redbreast tilapia Coptodon rendalli a widespread African species is the only substrate spawning cichlid in the lake 10 66 This large cichlid mainly feeds on macrophytes 10 67 The remaining are five mouthbrooding species of Oreochromis four chambo in the subgenus Nyasalapia O karongae O lidole O saka and O squamipinnis that are endemic to the Lake Malawi system as well as the closely related O shiranus which also is found in Lake Chilwa 10 44 60 The Malawi Oreochromis mainly feed on phytoplankton reach lengths up to 26 42 cm 10 17 in depending on the exact species and are mostly black or silvery gray with relatively indistinct dark bars 10 60 68 Male chambo have unique genital tassels when breeding which aid in egg fertilisation in a manner comparable to the egg spots on the anal fin of haplochromines 10 44 Non cichlids edit nbsp The kampango Bagrus meridionalis one of the largest catfish reaching up to 1 5 m 4 9 ft in length 69 The vast majority of the fish species in the lake are cichlids Among the non cichlid native fish are several species of cyprinids in genera Barbus Labeo and Opsaridium and the Lake Malawi sardine Engraulicypris sardella airbreathing catfish Bathyclarias and Clarias and the kampango Bagrus meridionalis mochokid catfish Chiloglanis and Malawi squeaker Synodontis njassae Mastacembelus spiny eel mormyrids Marcusenius Mormyrops and Petrocephalus the African tetra Brycinus imberi the poeciliid Aplocheilichthys johnstoni the spotted killifish Nothobranchius orthonotus and the mottled eel Anguilla nebulosa 10 At a genus level most of these are widespread in Africa but Bathyclarias is entirely restricted to the lake 70 Invertebrates edit Molluscs edit Lake Malawi is home to 28 species of freshwater snails including 16 endemics and 9 bivalves 2 endemics Aspatharia subreniformis and the unionid Nyassunio nyassaensis 71 72 The endemic freshwater snails are all members of the genera Bellamya Bulinus Gabbiella Lanistes and Melanoides 73 Lake Malawi is home to a total of four snail species in the genus Bulinus which is a known intermediate host of bilharzia A survey in Monkey Bay in 1964 found two endemic species of snails of the genus B nyassanus and B succinoides in the lake and two non endemic species B globosus and B forskalli in lagoons separated from it The latter species are known intermediate hosts of bilharzia and larvae of the parasite were detected in water containing these but in experiments C Wright of the British Museum of Natural History was unable to infect the two species endemic to the lake with the parasites The field workers who spent many hours on and in the lake did not find either B globosus or B forskalli in the lake itself 74 More recently the disease has become a problem in the lake itself as the endemic B nyassanus has become an intermediate host This change first noticed in the mid 1980s is possibly related to a decline in snail eating cichlids for example Trematocranus placodon due to overfishing and or a new strain of the bilharzia parasite 22 Crustaceans edit Unlike Lake Tanganyika with its many endemic freshwater crabs and shrimp there are few such species in Lake Malawi The Malawi blue crab Potamonautes lirrangensis syn P orbitospinus is the only crab in the lake and it is not endemic 75 76 The atyid shrimp Caridina malawensis is endemic to the lake but it is poorly known and has historically been confused with C nilotica which is not found in the lake 77 Pelagic zooplanktonic species include two cladocerans Diaphanosoma excisum and Bosmina longirostris three copepods Tropodiaptomus cunningtoni Thermocyclops neglectus and Mesocyclops aequatorialis 78 and several ostracods including both described and undescribed species 79 Lake flies edit nbsp Huge swarms of lake flies Chaoborus edulis resembling distant plumes of smoke over the waterLake Malawi is famous for the huge swarms of tiny harmless lake flies Chaoborus edulis 80 These swarms typically appearing far out over water can be mistaken for plumes of smoke and were also noticed by David Livingstone when he visited the lake 80 81 82 The aquatic larvae feed on zooplankton spending the day at the bottom and the night in the upper water levels 80 When they pupate they float to the surface and transform into adult flies 82 The adults are very short lived and the swarms which can be several hundred metres tall and often have a spiraling shape are part of their mating behaviour 80 83 They lay their eggs at the water s surface and the adults die 83 The larvae are an important food source for fish 78 80 84 and the adult flies are important both to birds and local people who collect them to make kungu cakes burgers a local delicacy with a very high protein content 81 82 2015 mine leak editIn January 2015 a sediment control tank collapsed at the Paladin Energy owned uranium mine in Northern Malawi after a high intensity rain storm hit the area It was revealed that approximately 50 litres of non radioactive material leaked into a local creek Despite reports in local media of radioactive contamination the government conducted independent scientific tests on the local river system and found that there was no effect on the environment 85 86 Swimming editThe 25 km solo swim across Lake Malawi between Cape Ngomba and Senga Bay has been accomplished on 5 occasions by 16 swimmers1992 Lewis Pugh 9hrs 52 minutes UK SA 87 and Otto Thanning SA 10hrs 5 minutes2010 Abigail Brown UK 9hrs 45 minutes 88 2013 Milko van Gool Netherlands 8hrs 46 minutes 89 and Kaitlin Harthoorn US 9hrs 17 minutes2016 current record Jean Craven SA Robert Dunford Kenya Michiel Le Roux SA Samantha Whelpton SA Greig Bannatyne SA Haydn Von Maltitz SA Douglas Livingstone Blevins SA 7hrs 53 mins 90 2019 Chris Stapley Eswatini and Jay Azran SA 8hrs 40 minutes Andrew Stevens Australia 10hrs 50 minutes and Ruth Azran SA 11hrs 8 minutes 91 92 In 2019 Martin Hobbs SA became the first person to swim the full length of Lake Malawi 54 days as well as setting the world record for longest solo swim in a lake 93 See also edit nbsp Lakes portal1989 Malawi earthquake 2009 Karonga earthquakes Mtenje village Southeast AfricaReferences edit a b c d e f g h Malawi Cichlids AC Tropical Fish Aquaticcommunity com Retrieved 2007 04 02 a b c d Lake Nyasa Columbia Encyclopedia Online Columbia University Press Retrieved 2011 08 02 a b c d Lake Malawi World Lakes Database International Lake Environment Committee Foundation Archived from the original on February 10 2007 Retrieved 2007 04 02 Scheffel Richard L Wernet Susan J eds 1980 Natural Wonders of the World United States of America Reader s Digest Association Inc p 222 ISBN 0 89577 087 3 Lake Niassa and its Coastal Zone Ramsar Sites Information Service Retrieved 25 April 2018 a b Protected Areas Programme United Nations Environment Programme World Conservation Monitoring Centre UNESCO October 1995 Archived from the original on 2008 05 11 Retrieved 2008 06 26 a b c Turner Seehausen Knight Allender and Robinson 2001 How many species of cichlid fishes are there in African lakes Molecular Ecology 10 793 806 a b WWF 10 June 2011 Mozambique s Lake Niassa declared reserve and Ramsar site Retrieved 17 July 2014 Pilskaln C H 2004 Seasonal and Interannual Particle Export in an African Rift Valley Lake A 5 Yr Record from Lake Malawi Southern East Africa Limnology and Oceanography 49 4 964 977 doi 10 2307 3597647 a b c d e f g h i j k l m n o p q r s t u v Konings Ad 1990 Ad Konings Book of Cichlids and all the other Fishes of Lake Malawi ISBN 978 0866225274 a b Park L E and A S Cohen 2011 Paleoecological response of ostracods to early Late Pleistocene lake level changes in Lake Malawi East Africa Palaeogeography Palaeoclimatology Palaeoecology 303 71 80 doi 10 1016 j palaeo 2010 02 038 Bhave A Vincent K and Mkwambisi D 2019 Projecting future water availability in Lake Malawi and the Shire River basin Future Climate for Africa Brief Cape Town CDKN https futureclimateafrica org resource brief projecting future water availability inlake malawi and the shire river basin permanent dead link Bhave Ajay G Bulcock Lauren Dessai Suraje Conway Declan Jewitt Graham Dougill Andrew J Kolusu Seshagiri Rao Mkwambisi David 2020 05 01 Lake Malawi s threshold behaviour A stakeholder informed model to simulate sensitivity to climate change Journal of Hydrology 584 124671 Bibcode 2020JHyd 58424671B doi 10 1016 j jhydrol 2020 124671 ISSN 0022 1694 S2CID 213751778 Future Climate for Africa How can we improve the use of information for a climate resilient Malawi February 2020 https futureclimateafrica org resource how can we improve the use of information for a climate resilient malawi Aghaindum Ajeagah Gideon 2017 06 15 Water as a weapon of international confrontations Editions L Harmattan ISBN 978 2 14 003961 4 Lake Malawi National Park World Heritage List UNESCO Retrieved 24 October 2015 Wilson Ab Teugels Gg Meyer A Apr 2008 Moritz Craig ed Marine Incursion The Freshwater Herring of Lake Tanganyika Are the Product of a Marine Invasion into West Africa PLOS ONE 3 4 e1979 Bibcode 2008PLoSO 3 1979W doi 10 1371 journal pone 0001979 PMC 2292254 PMID 18431469 a b c Givnish T J and K J Sytsma editors 1997 Molecular Evolution and Adaptive Radiation p 598 ISBN 0 521 57329 7 a b c d Delvaux D 1995 Age of Lake Malawi Nyasa and water level fluctuations Mus roy Afr centr Tervuren Belg Dept Geol Min Rapp ann 1993 amp 1994 99 108 a b Sturmbauer Baric Salzburger Ruber and Verheyen 2001 Lake Level Fluctuations Synchronize Genetic Divergences of Cichlid Fishes in African Lakes Mol Biol Evol 18 2 144 154 doi 10 1093 oxfordjournals molbev a003788 a b c Cohen Stone Beuning Park Reinthal Dettman Scholz Johnson King Talbot Brown and Ivory 2007 Ecological consequences of early Late Pleistocene megadroughts in tropical Africas Proc Natl Acad Sci USA 104 42 16422 16427 doi 10 1073 pnas 0703873104 a b Stauffer J R and H Madsen 2012 Schistosomiasis in Lake Malawi and the Potential Use of Indigenous Fish for Biological Control Pp 119 140 in Rokni M B editor Schistosomiasis ISBN 978 953 307 852 6 a b c Lowe McConnell R H 2003 Recent research in the African Great Lakes Fisheries biodiversity and cichlid evolution Freshwater Forum 20 1 4 64 What Are The Primary Inflows And Outflows Of Lake Malawi WorldAtlas Retrieved 2019 10 24 Jeal Tim 1973 Livingstone New York G P Putnam s Sons ISBN 9780399112157 Bayly Paul 2014 03 27 David Livingstone Africa s Greatest Explorer The Man the Missionary and the Myth Fonthill Media a b R W McColl 2005 Encyclopedia of World Geography Vol 1 Infobase Publishing p 576 Paice Edward 2007 Tip and Run The Untold Tragedy of the Great War in Africa Weidenfeld amp Nicolson p not cited ISBN 978 0 297 84709 0 The Guendolen v Hermann Von Wissmann Clash of Steel Govt clarifies on Tanzania Malawi border Daily News via KForum Dar es Salaam Tanzania 1 August 2007 Archived from the original on 13 July 2011 Retrieved 26 January 2009 Kamlomo Gabriel 27 August 2012 Malawi optimistic on Tanzania border dispute The Daily Times Malawi Archived from the 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Services Company Limited Retrieved 26 June 2011 permanent dead link Home Vessels Marine Services Company Limited Archived from the original on 10 September 2011 Retrieved 26 June 2011 BBC World Service African Perspective MV Vipya Lake Malawi Disaster Lake Malawi National Park UNESCO World Heritage Centre Geography amp Wildlife Malawi Our Africa SOS Children s Villages Retrieved 2015 09 14 a b c d Turner G F and N C Mwanyama July 1992 Distribution and Biology of Chambo Oreochromis spp in Lakes Malawi and Malombe Food and Agriculture Organization Fisheries and Aquaculture Department FI DP MLW 86 013 Field Document 21 Retrieved 13 April 2017 Preserving the Future for Lake Malawi web mit edu Retrieved 2015 09 14 a b Banda M 22 May 2013 Rapid drop in Lake Malawi s water levels drives down fish stocks The Guardian Retrieved 11 April 2017 Phiri T B Gobo E Tweddle D Kanyerere G Z 2019 amended version of 2018 assessment Bagrus meridionalis IUCN Red List of Threatened Species 2019 e 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CichlidForum Retrieved 16 April 2017 Barber P Pseudotropheus saulosi CichlidForum Retrieved 16 April 2017 Svardal Hannes Quah Fu Xiang Malinsky Milan Ngatunga Benjamin P Miska Eric A Salzburger Walter Genner Martin J Turner George F Durbin Richard 2019 08 18 Ancestral hybridisation facilitated species diversification in the Lake Malawi cichlid fish adaptive radiation Molecular Biology and Evolution doi 10 1101 738633 hdl 1983 40d3dafd f1d5 4ea4 a447 126df76a0651 S2CID 202010546 Kornfield I amp P F Smith 2000 African Cichlid Fishes Model Systems for Evolutionary Biology Annual Review of Ecology and Systematics 31 163 196 doi 10 1146 annurev ecolsys 31 1 163 a b c Oliver M K 12 April 2015 The Tilapias of Lake Malawi MalawiCichlids Retrieved 13 April 2017 Oliver M K 12 April 2015 The Nonendemic Haplochromine Cichlids of Lake Malawi MalawiCichlids Retrieved 13 April 2017 Pardee Keith African Cichlids Lake Malawi www aquariumlife net Archived from the original on 12 October 2017 Retrieved 30 September 2017 a b Genner Ngatunga Mzighani Smith and Turner 2015 Geographical ancestry of Lake Malawi s cichlid fish diversity Biol Lett 11 2015023 doi 10 1098 rsbl 2015 0232 Aquaticcommunity 2004 08 Haplochromis Retrieved 13 April 2017 Loiselle P V 1988 A Fishkeepers Guide to African Cichlids p 97 Salamander Books London amp New York ISBN 0 86101 407 3 Oliver M K 12 April 2015 Coptodon rendalli Malawicichlids Retrieved 13 April 2017 Froese Rainer Pauly Daniel eds 2017 Coptodon rendalli in FishBase April 2017 version Froese Rainer and Pauly Daniel eds 2017 Species of Oreochromis in FishBase April 2017 version Froese Rainer Pauly Daniel eds 2017 Bagrus meridionalis in FishBase April 2017 version Anseaume L and G G Teugels 1999 On the rehabilitation of the clariid catfish genus Bathyclarias endemic to the East African Rift Lake Malawi Fish Biology 55 2 405 419 doi 10 1111 j 1095 8649 1999 tb00687 x Segers H and Martens K editors 2005 The Diversity of Aquatic Ecosystems p 46 Developments in Hydrobiology Aquatic Biodiversity ISBN 1 4020 3745 7 Van Damme D 2018 Aspatharia subreniformis IUCN Red List of Threatened Species 2018 e T44266A120109809 doi 10 2305 IUCN UK 2018 2 RLTS T44266A120109809 en Brown D 1994 Freshwater Snails Of Africa And Their Medical Importance p 571 2nd edition ISBN 0 7484 0026 5 Wright C A Klein J Eccles D H 1967 Endemic species of Bulinus Mollusca Planorbidae in Lake Malawi Lake Nyasa Journal of Zoology 151 1 199 209 doi 10 1111 j 1469 7998 1967 tb02873 x Archived from the original on 2012 10 20 Retrieved 2010 05 22 Cumberlidge N and Meyer K S 2011 A revision of the freshwater crabs of Lake Kivu East Africa Journal Articles Paper 30 Dobson M 2004 Freshwater Crabs of Africa Archived 2016 06 23 at the Wayback Machine Freshwater Forum 21 3 26 Richard J and Clark P F 2009 African Caridina Crustacea Decapoda Caridea Atyidae redescriptions of C africana Kingsley 1882 C togoensis Hilgendorf 1893 C natalensis Bouvier 1925 and C roubaudi Bouvier 1925 with descriptions of 14 new species Zootaxa 1995 1 75 a b Darwall Allison Turner and Irvine 2010 Lake of flies or lake of fish A trophic model of Lake Malawi Ecological Modelling 221 713 727 doi 10 1016 j ecolmodel 2009 11 001 Martens K 2003 On the evolution of Gomphocythere Crustacea Ostracoda in Lake Nyassa Malawi East Africa with the description of 5 new species Hydrobiologia 497 1 2 121 144 doi 10 1023 A 1025417822555 a b c d e Morris B 2004 Insects and Human Life pp 73 76 ISBN 1 84520 075 6 a b van Huis A H van Gurp and M Dicke 2012 The Insect Cookbook Food for a Sustainable Planet p 31 ISBN 978 0 231 16684 3 a b c Malawi Tourism Interesting seasonal highlights of Malawi Archived 2014 08 12 at the Wayback Machine Retrieved 8 April 2017 a b Andrew D 30 June 2015 What Are These Strange Looking Clouds IFLScience Retrieved 8 April 2017 Allison Irvine Thompson and Ngatunga 1996 Diets and food consumption rates of pelagic fish in Lake Malawi Africa Freshwater Biology 35 3 489 515 doi 10 1111 j 1365 2427 1996 tb01764 x No harm caused by Paladin mine Kayerekera Malawi govt Malawi Nyasa Times Malawi breaking news in Malawi Archived from the original on 2015 04 01 Retrieved 2016 01 02 Radioactive pollution of Lake Malawi by Australian uranium company Paladin Expeditions Internal Waters Archived from the original on 2020 04 08 Retrieved 2018 12 11 Lake Malawi Crossing Archived from the original on 2015 09 06 Retrieved 2018 12 11 Milko Van Gool thought to be fastest male solo swimmer across North Channel BBC News 2013 07 30 CROSSING LAKE MALAWI TheCows Swimming Cows conquer Lake Malawi in brutal conditions The Cows Archived from the original on 2020 08 04 Retrieved 2020 05 21 Cows members conquer Lake Malawi in tough conditions for CHOC The Witness 10 May 2019 Retrieved 21 May 2020 Thom Liezl 26 April 2019 Man braves crocodiles hippos to set world record in 54 day swim across lake ABC News Further reading edit nbsp Wikimedia Commons has media related to Lake Malawi Mayall James December 1973 The Malawi Tanzania Boundary Dispute The Journal of Modern African Studies 11 4 611 628 doi 10 1017 S0022278X00008776 S2CID 154785268 Recent study on Lake Malawi water levels reveals drought 100 000 years ago Freshwater Fish Species in Lake Malawi Nyasa Southeast Africa Mongabay Retrieved 9 December 2016 Growing up in a Border District and Resolving the Tanzania Malawi Lake Dispute Compromise and concessions by Godfrey Mwakikagile African Renaissance Press 2022 Retrieved from https en wikipedia org w index php title Lake Malawi amp oldid 1200419149, wikipedia, wiki, book, books, library,

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