Pinus brutia
Pinus brutia, commonly known as the Turkish pine, is a species of pine native to the eastern Mediterranean region. The bulk of its range is in Turkey.
| Pinus brutia | |
|---|---|
| Turkish pine foliage and cones | |
| Scientific classification | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Gymnosperms |
| Division: | Pinophyta |
| Class: | Pinopsida |
| Order: | Pinales |
| Family: | Pinaceae |
| Genus: | Pinus |
| Subgenus: | P. subg. Pinus |
| Section: | P. sect. Pinus |
| Subsection: | Pinus subsect. Pinaster |
| Species: | P. brutia |
| Binomial name | |
| Pinus brutia | |
| Distribution | |
Turkish pine is also known by several other common names: Calabrian pine (from a naturalised population of the pine in Calabria in southern Italy, from where the pine was first botanically described), East Mediterranean pine, and Brutia pine.
Description edit
Pinus brutia is a medium-size tree, reaching 20–35 metres (66–115 feet) tall with a trunk diameter of up to 1 m (3+1⁄2 ft), exceptionally 2 m (6+1⁄2 ft). The bark is orange-red, thick and deeply fissured at the base of the trunk, and thin and flaky in the upper crown. The leaves (needles) are in pairs, slender, mostly 10–16 centimetres (4–6+1⁄4 inches) long, bright green to slightly yellowish green.[citation needed]
The cones are stout, heavy and hard, 6–11 cm (2+1⁄4–4+1⁄4 in) long and 4–5 cm (1+1⁄2–2 in) broad at the base when closed, green at first, ripening glossy red-brown when 24 months old. They open slowly over the next year or two to release the seeds, opening to 5–8 cm (2–3+1⁄4 in) broad. The seeds are 7–8 millimetres (1⁄4–3⁄8 in) long, with a 15–20 mm (5⁄8–3⁄4 in) wing, and are mainly wind-dispersed. One can see from gallery below in Flowers image, that this pine is not white pine classification, as it clearly has two needles per bundle, meaning it is either a red or jack pine.
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Bark
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Cone
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Flowers
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Foliage
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Tree
Varieties edit
Turkish pine is closely related to Aleppo pine, Canary Island pine, and Maritime pine, which all share many features with it. Some authors have treated it as a subspecies of Aleppo pine, but it is usually regarded as a distinct species.[citation needed] It is a moderately variable species, and the following subspecies and varieties are named:
| Image | Subspecies | Distribution |
|---|---|---|
| Pinus brutia var. brutia | typical form; most of the range | |
| Pinus brutia var. pityusa | Georgia, adjacent Russian Black Sea coast, and Crimea; barely distinct from the type | |
| Pinus brutia var. pendulifolia | southern coastal Turkey; needles 20–29 cm or 7+3⁄4–11+1⁄2 in, pendulous | |
| Pinus brutia var. eldarica (Afghan pine) | Azerbaijan; Georgia; needles 8–14 cm or 3+1⁄4–5+1⁄2 in, cones 5–9 cm or 2–3+1⁄2 in |
The Eldar pine is treated as a species (Pinus eldarica) by some authors; it is adapted to a drier climate with a summer rainfall peak, whereas var. brutia, var. pityusa, and var. pendulifolia are adapted to a climate with mainly winter rainfall.
Taxonomy edit
Italian botanist Michele Tenore described the species in 1811.
Distribution and habitat edit
The bulk of its range is in Turkey, but it also extends to southeasternmost Bulgaria,[2] the East Aegean Islands of Aegean Sea, Crete, Crimea, Iran, Georgia, Azerbaijan, northern Iraq, western Syria, Lebanon and Cyprus.[3] It generally occurs at low altitudes, mostly from sea level to 600 m (2,000 ft), up to 1,200 m (3,900 ft) in the south of its range.[citation needed]
Ecology edit
Pinus brutia is a diagnostic species of the vegetation class Pinetea halepensis.[4]
The Krüper's nuthatch, a rare nuthatch, is largely restricted to forests of Turkish pine and depends heavily on it for feeding; the ranges of the two species are largely coincident.[citation needed]
P. brutia is resistant to the Israeli pine bast scale insect Matsucoccus josephi and is a major host for Thaumetopoea caterpillars.
The species covers 175,000 hectares (430,000 acres) in Cyprus, roughly ~90% of all woodland coverage on the island.[5] It forms ectomycorrhizal associations with numerous species of fungi, and its logs and branches are excellent substrates for many kinds of decomposing organisms.
Uses edit
Honey edit
Turkish pine is host to a sap-sucking aphid Marchalina hellenica. Under normal circumstances, this insect does no significant damage to the pine, but is of great importance for the excess sugar it secretes. This sugar, "honeydew", is collected by honey bees which make it into a richly flavoured and valuable honey, "pine honey" (Turkish, çam balı), with reputed medicinal benefits.
Landmark edit
The "Lone Pine", a prominent landmark tree at an ANZAC First World War battle at Gallipoli, was this species. Cones from the battlefield were taken home to Australia, and plants sourced from the seeds were planted as living memorials.
"Lone Pine" memorials, based on cones brought back from Gallipoli, may use this species or Aleppo pine. Some memorials utilise other species altogether.[6]
Forestry edit
It is widely planted for timber, both in its native area (it is the most important tree in forestry in Turkey and Cyprus) and elsewhere in the Mediterranean region east to Pakistan. The timber is used for many purposes including carpentry, industry, general constructions, firewood and pulp.[7] In Israel it is sometimes preferred to the wider-used Pinus halepensis (Aleppo pine) because of its resistance to Matsucoccus josephi. It is also known for being well suited to recreational sites.
Cultivation edit
Pinus brutia is a popular ornamental tree, extensively planted in parks and gardens in hot dry areas (such as southern California, Utah, New Mexico and Nevada, as well as throughout Arizona and central Texas in the United States), where its considerable heat and drought tolerance is highly valued. The subspecies eldarica is the most drought tolerant form, used in Afghanistan, Iran and more recently in the Southwestern United States. In this region, P. brutia subsp. eldarica is referred to as "Eldarica pine", "Afghan pine" or "Mondell pine" (after Mondell Bennett, a commercial tree grower in New Mexico who popularized the species starting in 1969).[8]
References edit
- ^ Farjon, A. (2013). "Pinus brutia". IUCN Red List of Threatened Species. 2013: e.T42347A2974345. doi:10.2305/IUCN.UK.2013-1.RLTS.T42347A2974345.en. Retrieved 19 November 2021.
- ^ "Pinus brutia". Red Book of Bulgaria. Retrieved 10 January 2018.
- ^ "Pinus brutia". Germplasm Resources Information Network. Agricultural Research Service, United States Department of Agriculture. Retrieved 14 April 2011.
- ^ Bonari, Gianmaria; Fernández‐González, Federico; Çoban, Süleyman; Monteiro‐Henriques, Tiago; Bergmeier, Erwin; Didukh, Yakiv P.; Xystrakis, Fotios; Angiolini, Claudia; Chytrý, Kryštof; Acosta, Alicia T.R.; Agrillo, Emiliano (January 2021). Ewald, Jörg (ed.). "Classification of the Mediterranean lowland to submontane pine forest vegetation". Applied Vegetation Science. 24 (1). doi:10.1111/avsc.12544. hdl:10400.5/21923. ISSN 1402-2001. S2CID 228839165.
- ^ Loizides, Michael (2021-07-23). "Basidiomycete diversity within Calabrian pine (Pinus brutia) ecosystems on the island of Cyprus". Mycotaxon. 136 (2): 543. doi:10.5248/136.543. S2CID 238789537.
- ^ Wilcox, Mike; Spencer David (May 2007). "Stand up for the real Anzac Lone Pine Of Gallipoli" (PDF). New Zealand Journal of Forestry: 3–9. Retrieved 29 March 2012.
- ^ Fady, B.; Semerci, H. & Vendramin, G.G. (2003). "Aleppo and Brutia pines - Pinus halepensis/Pinus brutia" (PDF). EUFORGEN Technical Guidelines for Genetic Conservation and Use.
- ^ Widmoyer, Fred B. (1984-05-11). "History of Pinus Eldarica in the United States" (PDF). Southwest Christmas Tree Industry Research Needs and Commercial Opportunities: Proceedings of the Symposium held May 11, 1984, Tucson, Arizona. New Mexico State University. Retrieved 5 May 2012.
Further reading edit
- Shayanmehr, F., Jalali, S. G., Ghanati, F., & Kartoolinejad, D. (2008). Discrimination of Pinus eldarica MEDW. and its two new species by epicuticular wax, lignin content, electrophoretic isozyme and activity of peroxidase. Feddes Repertorium, 119(7‐8), 644–654. http://onlinelibrary.wiley.com/doi/10.1002/fedr.200811188/full
- Shayanmehr, F., Jalali, S. G., Ghanati, F., Kartoolinejad, D., & Apple, M. E. (2009). Two new morphotypes of Pinus eldarica: Discrimination by macromorphological and anatomical traits. Dendrobiology, 61, 27–36. http://onlinelibrary.wiley.com/doi/10.1002/fedr.200811188/full
- Frankis, M. P. (1999). Pinus brutia. Curtis's Botanical Magazine 16: 173–184.
External links edit
- Photos of trees in Turkey (scroll down page)
- Gymnosperm Database - Pinus brutia
- Pinus brutia - distribution map, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN)