Pinus halepensis
Pinus halepensis, commonly known as the Aleppo pine, also known as the Jerusalem pine,[2] is a pine native to the Mediterranean region.
| Pinus halepensis | |
|---|---|
| Pinus halepensis in Sounion Natural Park, Greece | |
| Scientific classification | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Gymnospermae |
| Division: | Pinophyta |
| Class: | Pinopsida |
| Order: | Pinales |
| Family: | Pinaceae |
| Genus: | Pinus |
| Subgenus: | P. subg. Pinus |
| Section: | P. sect. Pinus |
| Subsection: | Pinus subsect. Pinaster |
| Species: | P. halepensis |
| Binomial name | |
| Pinus halepensis | |
| Distribution map | |
Description edit
Pinus halepensis is a small to medium-sized tree, 15–25 metres (49–82 feet) tall, with a trunk diameter up to 60 centimetres (24 inches), exceptionally up to 1 m (3 ft 3 in). The bark is orange-red, thick, and deeply fissured at the base of the trunk, and thin and flaky in the upper crown. The leaves ('needles') are very slender, 6–12 cm (2+1⁄4–4+3⁄4 in) long, distinctly yellowish green, and produced in pairs (rarely a few in threes). The cones are narrow conic, 5–12 cm (2–4+3⁄4 in) long and 2–3 cm (3⁄4–1+1⁄4 in) broad at the base when closed, green at first, ripening glossy red-brown when 24 months old. They open slowly over the next few years, a process quickened if they are exposed to heat such as in forest fires. The cones open 5–8 cm (2–3+1⁄4 in) wide to allow the seeds to disperse. The seeds are 5–6 millimetres (3⁄16–1⁄4 in) long, with a 20 mm (13⁄16 in) wing, and are wind-dispersed.[3][4][5]
- Cones
- Foliage
- Bark and trunk
- Plate from Lambert's Description of the Genus Pinus
- Cone of pinus halepensis in Hebron
- Pinus halepensis forest at the island of Mljet
- A dead Aleppo pine in front of the Étang de Thau
- A grove of Aleppo pines in Pinet
Related species edit
The Aleppo pine is closely related to the Turkish pine, Canary Island pine, and maritime pine, which all share many of its characteristics. Some authors include the Turkish pine as a subspecies of the Aleppo pine, as Pinus halepensis subsp. brutia (Ten.) Holmboe,[6] but it is usually regarded as a distinct species.[3][4][5][7] It is a relatively nonvariable species, in that its morphological characteristics stay constant over the entire range.[3]
Distribution and habitat edit
The native range of Pinus halepensis extends from Morocco, Algeria, Tunisia, and Spain north to southern France, Malta, Italy, Croatia, Montenegro, and Albania, and east to Greece. It has been introduced into many parts of the world, including Portugal. There is an outlying population (from which it was first described) in Syria, Lebanon, southern Turkey, Jordan, Israel and Palestine.
The species is generally found at low altitudes, mostly from sea level to 200 m (660 ft), but can grow above 1,000 m (3,300 ft) in southern and eastern Spain, well over 1,200 m (3,900 ft) on Crete, and up to 1,700 m (5,600 ft) in the south, in Morocco, Algeria and Tunisia.[3][4] The tree is able to quickly colonize open and disturbed areas. It is classed as an invasive species in South Africa.[8] It can grow on all substrates and almost in all bioclimates in the Mediterranean.[9]
Pinus halepensis is a diagnostic species of the vegetation class Pinetea halepensis.[10]
Uses edit
The resin of the Aleppo pine is used to flavor the Greek wine retsina.
From the pine nuts of the Aleppo pine is made a pudding called asidet zgougou in the Tunisian dialect; it is served in bowls, covered with cream, and topped with almonds and small candies.
The Maltese dessert prinjolata is also prepared using these pine nuts, both in its filling as well as a topping.
Aleppo pine are used for bonsai.
Forestry edit
In its native area, P. halepensis is widely planted for its fine timber, making it one of the most important forestry trees in Algeria and Morocco.[5]
In Israel, natural patches of Aleppo pine forests can be found in the Carmel and Galilee regions.[11] The Aleppo pine, along with Pinus brutia, has been planted extensively by the Jewish National Fund. It proved very successful in Yatir Forest in the northern Negev (on the edge of the desert), where foresters had not expected it to survive. Many Aleppo pine forests exist today in Israel and are used for recreational purposes. Although it is a local species, some argue that the historical replacement of natural oak maquis shrubland and garrigue with tall stands of pine has created "ecological deserts" and has significantly changed the species assemblage of these regions.[12] The species produces timber which is valued for its hardness, density and unproblematic seasoning. Seasoned timber is inclined to tear out with planing, but this can be avoided by using sharp blades or adjusting the sharpening angle of tools.[13]
The Aleppo pine is considered an invasive species though useful in South Africa; in South Australia, a control program is in place on Eyre Peninsula.
Landscape edit
Pinus halepensis is a popular ornamental tree, extensively planted in gardens, parks, and private and agency landscapes in hot dry areas such as Southern California and the Karoo in South Africa, where the Aleppo pine's considerable heat and drought tolerance, fast growth, and aesthetic qualities are highly valued.
In culture edit
Paul Cézanne had an Aleppo pine in his garden at Aix-en-Provence; this tree was the inspiration and model for his painting The Big Trees. As of 2005, the tree is still growing in Cézanne's garden.[14]
References edit
- ^ Farjon, A. (2013). "Pinus halepensis". IUCN Red List of Threatened Species. 2013: e.T42366A2975569. doi:10.2305/IUCN.UK.2013-1.RLTS.T42366A2975569.en. Retrieved 19 November 2021.
- ^ Aisner, R.; Terkel, J. (1992-08-01). "Ontogeny of pine cone opening behaviour in the black rat, Rattus rattus". Animal Behaviour. 44: 327–336. doi:10.1016/0003-3472(92)90038-B. ISSN 0003-3472. S2CID 53148456.
- ^ a b c d Farjon, A. (2005). Pines. Drawings and Descriptions of the genus Pinus. Brill, Leiden. ISBN 90-04-13916-8.
- ^ a b c Rushforth, K. (1999). Trees of Britain and Europe. Collins ISBN 0-00-220013-9.
- ^ a b c Nahal, I. (1962). Le Pin d'Alep (Pinus halepensis Miller). Étude taxonomique, phytogéographique, écologique et sylvicole. Annales de l'École National des Eaux et Forêts (Nancy) 19: 1–207.
- ^ Christensen, K. I. (1997). Gymnospermae. Pp. 1–17 in Strid, A., & Tan, K., eds., Flora Hellenica 1. Königstein.
- ^ Richardson, D. M., ed. (1998). Ecology and Biogeography of Pinus. Cambridge University Press ISBN 0-521-55176-5.
- ^ "Aleppo pine – Invasive Species South Africa".
- ^ Facy, B.; Semerci, H. & Vendramin, G.G. (2003). (PDF). EUFORGEN Technical Guidelines for Genetic Conservation and Use. Archived from the original (PDF) on 2018-09-30. Retrieved 2016-10-24.
- ^ Bonari, Gianmaria; Fernández‐González, Federico; Çoban, Süleyman; Monteiro‐Henriques, Tiago; Bergmeier, Erwin; Didukh, Yakiv P.; Xystrakis, Fotios; Angiolini, Claudia; Chytrý, Kryštof; Acosta, Alicia T.R.; Agrillo, Emiliano (January 2021). Ewald, Jörg (ed.). "Classification of the Mediterranean lowland to submontane pine forest vegetation". Applied Vegetation Science. 24 (1). doi:10.1111/avsc.12544. hdl:10400.5/21923. ISSN 1402-2001. S2CID 228839165.
- ^
- ^ F.T. Maestre, J. Cortina . "Are Pinus halepensis plantations useful as a restoration tool in semiarid Mediterranean areas?" Forest Ecology and Management, 2004 (Elsevier).
- ^ Reducing Tear Out when Wood Planing
- ^ Cézanne, P. "Visions". In Architectural Digest, December 2005: 117.
External links edit
- Gymnosperm Database: Pinus halepensis
- Pinus halepensis—distribution map, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN)