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Passerida

Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri (standard taxonomic practice would place them at the rank of infraorder). While more recent research suggests that its sister parvorder, Corvida, is not a monophyletic grouping, the Passerida as a distinct clade are widely accepted.

Passerida
Eurasian tree sparrow (Passer montanus)
Grey-streaked flycatcher (Muscicapa griseisticta)
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Suborder: Passeri
Infraorder: Passerida
Superfamilies

and see text

Systematics and phylogeny edit

The Passerida quite certainly consist of the 3 major subclades outlined by Sibley & Ahlquist (1990). However, their content has been much revised. In addition, it has turned out that not all passeridan lineages neatly fit into this arrangement. The kinglets are so distinct that they might actually form a separate infraorder, as they are only slightly less basal than the Corvoidea or the Picathartidae. See Jønsson & Fjeldså (2006) for details on phylogeny.

Superfamily Sylvioidea edit

Mostly smallish insectivores, distribution centered on the Indo-Pacific region. Few occur in the Americas,[1][2] highest diversity of families probably in subtropical East Asia and tropical Africa. Relationships of the latter are still not well-resolved as of 2019.

Includes the "Old World babblers" and "Old World warbler", two highly paraphyletic "wastebin taxa" which for long united the bulk of the thrush-sized and sparrow-sized sylvioids, respectively. Sometimes, they were even united with the muscicapoids as one huge "family" including most "songsters". Usually skulking in shrubby vegetation, many are extremely drab (most of birdwatchers' "little brown jobs" belong here) and rely on complex and often melodious vocalizations as social signals; others are less accomplished singers but produce a diversity of squeaking and twittering calls. The sexes usually look alike, though in some the males are noticeably brighter, typically with vivid yellow, green and blackish hues. Red plumage is usually due to phaeomelanins rather than carotins, and blue coloration is rarely found in this superfamily. Even in the more colorful species the plumage is usually quite cryptic in the natural habitat, but numerous have contrasting facial patterns.

Superfamily Muscicapoidea edit

Generally middle-sized insectivores, with frugivory also very important; near-global distribution centered on Old World tropics. One family is endemic to the Americas, two are almost cosmopolitan, but half the families are absent or nearly so from the Americas (and Australia). Many have strong legs and are capable of running on the ground quickly. Some brightly colored (often with dark bluish hues and/or iridescence) and in such cases usually strongly sexually dimorphic; more often, however, sexes rather alike, with drab brownish plumage spotted and streaked (particularly on the underside) for camouflage. Many have highly accomplished, complex, melodious and loud songs; a considerable number is capable of sophisticated vocal mimicry.

  • Cinclidae: dippers
  • Muscicapidae: Old World flycatchers and chats. Monophyly needs confirmation.
  • Turdidae: thrushes and allies. Monophyly needs confirmation.
  • Buphagidae: oxpeckers. Formerly usually included in Sturnidae.
  • Sturnidae: starlings and possibly Philippine creepers. Placement of latter in Muscicapoidea seems robust, but inclusion in Sturnidae requires confirmation; possibly distinct family Rhabdornithidae.
  • Mimidae: mockingbirds and thrashers

Superfamily Passeroidea edit

Mostly smallish herbivores, near-global distribution centered on Palearctic and Americas. Often pronounced sexual dimorphism with males among the most colorful birds alive. Songs tend to be fairly simple warbling and chirping, with many species relying as much or more on visual mating displays. Includes the nine-primaried oscines (probably a subclade). The basal radiation is mostly found in the Old World, with only Motacillidae naturally occurring in the Americas and Estrildidae in Australia.

The nine-primaried oscines unite most birds commonly called "sparrows" in North American and "finches" in European English, as well as a number of other mostly American groups. They are divided into the fringillid radiation which is largely restricted to the Old World, and the numerous emberizoid families of the Americas, of which in turn only Emberizidae and the Arctic circumpolar Calcariidae have reached the Old World unaided by humans. Besides these, the singular olive warbler from North to Central America apparently represents a very ancient "living fossil" passeroid; its relationships were long disputed as its outward appearance and ecology resemble Setophaga warblers, but its anatomy is in some aspects convergent or symplesiomorphic with sylvioids.

Passerida incertae sedis edit

Rather basal Passerida, most of which seem to constitute several small but distinct superfamilies. Most occur in Asia, Africa and North America.

  • Possible superfamily "Dicaeoidea" – sunbirds and flowerpeckers. Small frugivores/nectarivores of the Old World tropics, typically sexually dimorphic, with bright and/or iridescent colors in males. Songs are simple chirping whistles.
  • Possible superfamily Bombycilloidea – waxwings and allies. Mid-sized, mostly Holarctic frugivores; plumage silky and dark to greyish-brownish, with little if any sexual dimorphism. Ringing calls and usually quite vocal, but no dedicated song.
  • Possible superfamily Paroidea – titmice and allies. Small, round-headed, with tiny pointed bills. Forage acrobatically among twigs, mostly eating small insects and seeds. Generally Palaearctic, ranging into the Old World tropics and North America. Little if any sexual dimorphism; may be brownish-grey or fairly bright and multicolored. In any case head plumage usually either fairly uniform and greyish, or with black markings, and/or crested. Songs usually repetitive chirped phrases.
    • Paridae: tits, chickadees and titmice
    • Remizidae: penduline tits. Sometimes included in Paridae.
    • Stenostiridae: stenostirids ("flycatcher-tits"). A newly assembled family; sometimes included in Paridae.
  • Possible superfamily Certhioidea (or Sittoidea) – wrens and allies. Insectivores, usually tiny. Expert climbers, most are capable of running up vertical trees or cliffs, some can even climb head-downwards. Predominantly palearctic, but two families entirely or almost so American, and one restricted to the Old World tropics. Little sexual dimorphism; plumage either greyish and fairly uniform at least on the upperside, or brown above, lighter below, and heavily streaked. Clear whistled vocalizations, usually melodic and louder than one would expect from birds of their size. Songs often complex, e.g. with social duetting, and apparently very important in species recognition.
  • Possible monotypic superfamily N.N.: sugarbirds (Promeropidae).
    Relatives of the Australasian honeyeaters; 2 species restricted to the Cape Floral Region and mainly feeding on Proteaceae nectar and associated insects. Medium-sized, with extremely long tails; drab coloration resembling a muscicapoid, sexes almost alike. Vocalizations similar to honeyeaters; males have specialized wing feathers that produce noise during courtship flights.
  • Possible monotypic superfamily N.N.: hyliotas (Hyliotidae; formerly in Sylviidae).
    4 species of smallish insectivorous "warblers" from tropical African woodlands. Two- or three-toned, medium grey to blackish above, more or less intense yellowish below, some species with white wing markings. Sexual dimorphism slight; whistling calls.
  • Possible monotypic superfamily Reguloidea – kinglets (Regulidae).
    Tentatively placed here; may belong in Certhioidea. Some 5 species of tiny rotund Holarctic woodland insectivores. The smallest songbirds, and as a family the smallest living birds altogether by average length.[7] Greenish-brownish above, dull whitish below, and with a bright yellow to red central patch on the top of the head. Rapid twittering high-pitched chirps, easier heard than seen. Unlike hummingbirds which lay only 2 eggs per clutch and can live up to a dozen years or more, kinglets are (together with some small quails) the most r-selected birds alive, with clutch sizes of around 10 eggs, a maximum lifespan of merely around 5 years even in captivity, and an annual mortality of 80%.

Probably not Passerida edit

These lineages have been assigned to the Passerida in recent times, often based on DNA-DNA hybridization data. However, they are probably more basal among the songbirds and would belong either to the Corvoidea or the allied basal lineages. Most of them are either African or Wallacean groups.

  • Aegithinidae: ioras. Formerly in Irenidae, and may be closely related; possibly Corvoidea closely related to cuckooshrikes.
  • Chloropseidae: leafbirds. Formerly in Irenidae, and may be closely related.
  • Irenidae; fairy-bluebirds. Formerly in "Timaliidae" or Pycnonotidae.
  • Melanocharitidae: berrypeckers and longbills. Formerly in Dicaeidae; possibly Corvoidea closely related to cuckooshrikes.
  • Paramythiidae: tit berrypecker and crested berrypeckers. Formerly in Dicaeidae or Melanocharitidae; possibly Corvoidea closely related to whipbirds.
  • Picathartidae: rockfowl. Formerly in "Timaliidae", but possibly close to rockjumpers (Chaetops) and sometimes considered the basal living branch of the Passerida.
  • Chaetopidae: rockjumpers. Possibly close to rockfowls (Picathartes).
  • Platysteiridae: wattle-eyes or puffback flycatchers. Formerly in Muscicapidae; probably Corvoidea closely related to bush-shrikes.

See also edit

References edit

  1. ^ Fregin, Silke; Haase, Martin; Olsson, Urban; Alström, Per (2012). "New insights into family relationships within the avian superfamily Sylvioidea (Passeriformes) based on seven molecular markers". BMC Evolutionary Biology. 12 (157): 1–12. doi:10.1186/1471-2148-12-157. PMC 3462691. PMID 22920688.
  2. ^ Alström, Per; Olsson, Urban; Lei, Fumin (2013). "A review of the recent advances in the systematics of the avian superfamily Sylvioidea". Chinese Birds. 4 (2): 99–131. doi:10.5122/cbirds.2013.0016.
  3. ^ Gill, Frank; Donsker, David, eds. (2018). "Family index". World Bird List Version 8.2. International Ornithologists' Union. Retrieved 24 July 2018.
  4. ^ Barker, F. K.; Burns, K. J.; Klicka, J.; Lanyon, S. M.; Lovette, I. J. (2013). "Going to extremes: Contrasting rates of diversification in a recent radiation of New World passerine birds". Systematic Biology. 62 (2): 298–320. doi:10.1093/sysbio/sys094. PMID 23229025.
  5. ^ Barker, F. K.; Burns, K. J.; Klicka, J.; Lanyon, S. M.; Lovette, I. J. (2015). "New insights into New World biogeography: An integrated view from the phylogeny of blackbirds, cardinals, sparrows, tanagers, warblers, and allies". The Auk. 132 (2): 333–348. doi:10.1642/AUK-14-110.1. S2CID 53058340.
  6. ^ del Hoyo, J.; Elliott, A.; Sargatal, J.; Christie, D.A.; de Juana, E., eds. (2019). "Passeriformes". Handbook of the Birds of the World Alive. Lynx Edicions, Barcelona. Retrieved 5 January 2019.
  7. ^ Many species of hummingbirds are - at least excepting the bill - shorter and much lighter than kinglets. But while the latter are one small genus and differ little in size among each other, hummingbirds are a diverse order of nonpasseriform birds, including dozens of species which far exceed kinglets in length, and in many cases even in weight.
  • Jønsson, Knud A.; Fjeldså, Jon (2006). "A phylogenetic supertree of oscine passerine birds (Aves: Passeri)". Zool. Scripta. 35 (2): 149–186. doi:10.1111/j.1463-6409.2006.00221.x. S2CID 85317440.
  • Harshman, John (2008). Passerida. Version 23 June 2008 (under construction). in The Tree of Life Web Project. Retrieved 24 January 2011.
  • Sibley, Charles Gald & Ahlquist, Jon Edward (1990): Phylogeny and classification of birds. Yale University Press, New Haven, Conn.

passerida, under, sibley, ahlquist, taxonomy, parvorders, contained, within, suborder, passeri, standard, taxonomic, practice, would, place, them, rank, infraorder, while, more, recent, research, suggests, that, sister, parvorder, corvida, monophyletic, groupi. Passerida is under the Sibley Ahlquist taxonomy one of two parvorders contained within the suborder Passeri standard taxonomic practice would place them at the rank of infraorder While more recent research suggests that its sister parvorder Corvida is not a monophyletic grouping the Passerida as a distinct clade are widely accepted Passerida Eurasian tree sparrow Passer montanus Grey streaked flycatcher Muscicapa griseisticta Scientific classification Domain Eukaryota Kingdom Animalia Phylum Chordata Class Aves Order Passeriformes Suborder Passeri Infraorder Passerida Superfamilies Sylvioidea Muscicapoidea Passeroidea and see text Contents 1 Systematics and phylogeny 1 1 Superfamily Sylvioidea 1 2 Superfamily Muscicapoidea 1 3 Superfamily Passeroidea 1 4 Passerida incertae sedis 1 5 Probably not Passerida 2 See also 3 ReferencesSystematics and phylogeny editThe Passerida quite certainly consist of the 3 major subclades outlined by Sibley amp Ahlquist 1990 However their content has been much revised In addition it has turned out that not all passeridan lineages neatly fit into this arrangement The kinglets are so distinct that they might actually form a separate infraorder as they are only slightly less basal than the Corvoidea or the Picathartidae See Jonsson amp Fjeldsa 2006 for details on phylogeny Superfamily Sylvioidea edit Mostly smallish insectivores distribution centered on the Indo Pacific region Few occur in the Americas 1 2 highest diversity of families probably in subtropical East Asia and tropical Africa Relationships of the latter are still not well resolved as of 2019 Includes the Old World babblers and Old World warbler two highly paraphyletic wastebin taxa which for long united the bulk of the thrush sized and sparrow sized sylvioids respectively Sometimes they were even united with the muscicapoids as one huge family including most songsters Usually skulking in shrubby vegetation many are extremely drab most of birdwatchers little brown jobs belong here and rely on complex and often melodious vocalizations as social signals others are less accomplished singers but produce a diversity of squeaking and twittering calls The sexes usually look alike though in some the males are noticeably brighter typically with vivid yellow green and blackish hues Red plumage is usually due to phaeomelanins rather than carotins and blue coloration is rarely found in this superfamily Even in the more colorful species the plumage is usually quite cryptic in the natural habitat but numerous have contrasting facial patterns Leiothrichidae laughingthrushes and allies Pellorneidae fulvettas ground babblers Timaliidae babblers scimitar babblers Zosteropidae white eyes Sylviidae sylviid babblers Cettiidae cettia bush warblers and allies Scotocercidae streaked scrub warbler Erythrocercidae yellow flycatchers Aegithalidae bushtits Hyliidae doubtfully distinct from Cettiidae not recognised by Gill and Donsker who consider the genera Hylia and Pholidornis as Incertae sedis 3 Phylloscopidae leaf warblers and allies Pycnonotidae bulbuls Hirundinidae swallows martins Bernieridae Madagascan warblers Donacobiidae black capped donacobius Locustellidae grassbirds and allies Acrocephalidae reed warblers and allies Pnoepygidae cupwings Cisticolidae cisticolas and allies Macrosphenidae crombecs African warblers Alaudidae larks Panuridae bearded reedling Nicatoridae nicators Superfamily Muscicapoidea edit Generally middle sized insectivores with frugivory also very important near global distribution centered on Old World tropics One family is endemic to the Americas two are almost cosmopolitan but half the families are absent or nearly so from the Americas and Australia Many have strong legs and are capable of running on the ground quickly Some brightly colored often with dark bluish hues and or iridescence and in such cases usually strongly sexually dimorphic more often however sexes rather alike with drab brownish plumage spotted and streaked particularly on the underside for camouflage Many have highly accomplished complex melodious and loud songs a considerable number is capable of sophisticated vocal mimicry Cinclidae dippers Muscicapidae Old World flycatchers and chats Monophyly needs confirmation Turdidae thrushes and allies Monophyly needs confirmation Buphagidae oxpeckers Formerly usually included in Sturnidae Sturnidae starlings and possibly Philippine creepers Placement of latter in Muscicapoidea seems robust but inclusion in Sturnidae requires confirmation possibly distinct family Rhabdornithidae Mimidae mockingbirds and thrashers Superfamily Passeroidea edit Mostly smallish herbivores near global distribution centered on Palearctic and Americas Often pronounced sexual dimorphism with males among the most colorful birds alive Songs tend to be fairly simple warbling and chirping with many species relying as much or more on visual mating displays Includes the nine primaried oscines probably a subclade The basal radiation is mostly found in the Old World with only Motacillidae naturally occurring in the Americas and Estrildidae in Australia The nine primaried oscines unite most birds commonly called sparrows in North American and finches in European English as well as a number of other mostly American groups They are divided into the fringillid radiation which is largely restricted to the Old World and the numerous emberizoid families of the Americas of which in turn only Emberizidae and the Arctic circumpolar Calcariidae have reached the Old World unaided by humans Besides these the singular olive warbler from North to Central America apparently represents a very ancient living fossil passeroid its relationships were long disputed as its outward appearance and ecology resemble Setophaga warblers but its anatomy is in some aspects convergent or symplesiomorphic with sylvioids Passeridae true sparrows Prunellidae accentors Urocynchramidae Przewalski s finch Recently split from Fringillidae tentatively placed here Estrildidae estrildid finches waxbills munias etc Ploceidae weavers Certain members of Ploceidae such as the long tailed widowbird are well known for their elaborate sexual ornaments Viduidae indigobirds and whydahs Nine primaried oscines Motacillidae wagtails and pipits Peucedramidae olive warbler Fringillidae true finches Includes the Hawaiian honeycreepers Emberizoidea New World nine primaried oscine radiation 4 5 6 Icteridae grackles New World blackbirds and New World orioles Parulidae New World warblers Icteriidae yellow breasted chat Phaenicophilidae Hispaniolan tanagers Zeledoniidae wrenthrush Teretistridae Cuban warblers Thraupidae tanagers and allies Mitrospingidae mitrospingid tanagers and allies Rhodinocichlidae rosy thrush tanager Calyptophilidae chat tanagers Nesospingidae Puerto Rican tanager Spindalidae spindalises Cardinalidae cardinals Emberizidae buntings Passerellidae American sparrows Calcariidae snow buntings and longspurs Passerida incertae sedis edit Rather basal Passerida most of which seem to constitute several small but distinct superfamilies Most occur in Asia Africa and North America Possible superfamily Dicaeoidea sunbirds and flowerpeckers Small frugivores nectarivores of the Old World tropics typically sexually dimorphic with bright and or iridescent colors in males Songs are simple chirping whistles Nectariniidae sunbirds Dicaeidae flowerpeckers Possible superfamily Bombycilloidea waxwings and allies Mid sized mostly Holarctic frugivores plumage silky and dark to greyish brownish with little if any sexual dimorphism Ringing calls and usually quite vocal but no dedicated song Bombycillidae waxwings Dulidae palmchat Tentatively placed here Ptiliogonatidae silky flycatchers Tentatively placed here Hypocoliidae hypocolius Tentatively placed here Mohoidae Hawaiian honeyeaters Tentatively placed here Possible superfamily Paroidea titmice and allies Small round headed with tiny pointed bills Forage acrobatically among twigs mostly eating small insects and seeds Generally Palaearctic ranging into the Old World tropics and North America Little if any sexual dimorphism may be brownish grey or fairly bright and multicolored In any case head plumage usually either fairly uniform and greyish or with black markings and or crested Songs usually repetitive chirped phrases Paridae tits chickadees and titmice Remizidae penduline tits Sometimes included in Paridae Stenostiridae stenostirids flycatcher tits A newly assembled family sometimes included in Paridae Possible superfamily Certhioidea or Sittoidea wrens and allies Insectivores usually tiny Expert climbers most are capable of running up vertical trees or cliffs some can even climb head downwards Predominantly palearctic but two families entirely or almost so American and one restricted to the Old World tropics Little sexual dimorphism plumage either greyish and fairly uniform at least on the upperside or brown above lighter below and heavily streaked Clear whistled vocalizations usually melodic and louder than one would expect from birds of their size Songs often complex e g with social duetting and apparently very important in species recognition Sittidae nuthatches Tichodromadidae wallcreeper Tentatively placed here Certhiidae treecreepers Troglodytidae wrens Polioptilidae gnatcatchers Possible monotypic superfamily N N sugarbirds Promeropidae Relatives of the Australasian honeyeaters 2 species restricted to the Cape Floral Region and mainly feeding on Proteaceae nectar and associated insects Medium sized with extremely long tails drab coloration resembling a muscicapoid sexes almost alike Vocalizations similar to honeyeaters males have specialized wing feathers that produce noise during courtship flights Possible monotypic superfamily N N hyliotas Hyliotidae formerly in Sylviidae 4 species of smallish insectivorous warblers from tropical African woodlands Two or three toned medium grey to blackish above more or less intense yellowish below some species with white wing markings Sexual dimorphism slight whistling calls Possible monotypic superfamily Reguloidea kinglets Regulidae Tentatively placed here may belong in Certhioidea Some 5 species of tiny rotund Holarctic woodland insectivores The smallest songbirds and as a family the smallest living birds altogether by average length 7 Greenish brownish above dull whitish below and with a bright yellow to red central patch on the top of the head Rapid twittering high pitched chirps easier heard than seen Unlike hummingbirds which lay only 2 eggs per clutch and can live up to a dozen years or more kinglets are together with some small quails the most r selected birds alive with clutch sizes of around 10 eggs a maximum lifespan of merely around 5 years even in captivity and an annual mortality of 80 Probably not Passerida edit These lineages have been assigned to the Passerida in recent times often based on DNA DNA hybridization data However they are probably more basal among the songbirds and would belong either to the Corvoidea or the allied basal lineages Most of them are either African or Wallacean groups Aegithinidae ioras Formerly in Irenidae and may be closely related possibly Corvoidea closely related to cuckooshrikes Chloropseidae leafbirds Formerly in Irenidae and may be closely related Irenidae fairy bluebirds Formerly in Timaliidae or Pycnonotidae Melanocharitidae berrypeckers and longbills Formerly in Dicaeidae possibly Corvoidea closely related to cuckooshrikes Paramythiidae tit berrypecker and crested berrypeckers Formerly in Dicaeidae or Melanocharitidae possibly Corvoidea closely related to whipbirds Picathartidae rockfowl Formerly in Timaliidae but possibly close to rockjumpers Chaetops and sometimes considered the basal living branch of the Passerida Chaetopidae rockjumpers Possibly close to rockfowls Picathartes Platysteiridae wattle eyes or puffback flycatchers Formerly in Muscicapidae probably Corvoidea closely related to bush shrikes See also editList of birdsReferences edit nbsp Wikimedia Commons has media related to Passerida nbsp Wikispecies has information related to Passerida Fregin Silke Haase Martin Olsson Urban Alstrom Per 2012 New insights into family relationships within the avian superfamily Sylvioidea Passeriformes based on seven molecular markers BMC Evolutionary Biology 12 157 1 12 doi 10 1186 1471 2148 12 157 PMC 3462691 PMID 22920688 Alstrom Per Olsson Urban Lei Fumin 2013 A review of the recent advances in the systematics of the avian superfamily Sylvioidea Chinese Birds 4 2 99 131 doi 10 5122 cbirds 2013 0016 Gill Frank Donsker David eds 2018 Family index World Bird List Version 8 2 International Ornithologists Union Retrieved 24 July 2018 Barker F K Burns K J Klicka J Lanyon S M Lovette I J 2013 Going to extremes Contrasting rates of diversification in a recent radiation of New World passerine birds Systematic Biology 62 2 298 320 doi 10 1093 sysbio sys094 PMID 23229025 Barker F K Burns K J Klicka J Lanyon S M Lovette I J 2015 New insights into New World biogeography An integrated view from the phylogeny of blackbirds cardinals sparrows tanagers warblers and allies The Auk 132 2 333 348 doi 10 1642 AUK 14 110 1 S2CID 53058340 del Hoyo J Elliott A Sargatal J Christie D A de Juana E eds 2019 Passeriformes Handbook of the Birds of the World Alive Lynx Edicions Barcelona Retrieved 5 January 2019 Many species of hummingbirds are at least excepting the bill shorter and much lighter than kinglets But while the latter are one small genus and differ little in size among each other hummingbirds are a diverse order of nonpasseriform birds including dozens of species which far exceed kinglets in length and in many cases even in weight Jonsson Knud A Fjeldsa Jon 2006 A phylogenetic supertree of oscine passerine birds Aves Passeri Zool Scripta 35 2 149 186 doi 10 1111 j 1463 6409 2006 00221 x S2CID 85317440 Harshman John 2008 Passerida Version 23 June 2008 under construction in The Tree of Life Web Project Retrieved 24 January 2011 Sibley Charles Gald amp Ahlquist Jon Edward 1990 Phylogeny and classification of birds Yale University Press New Haven Conn Retrieved from https en wikipedia org w index php title Passerida amp oldid 1206091621 Superfamily Passeroidea, wikipedia, wiki, book, books, library,

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