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Yamnaya culture

The Yamnaya culture or the Yamna culture (Russian: Ямная культура, Ukrainian: Ямна культура lit. 'culture of pits'), also known as the Pit Grave culture or Ochre Grave culture, was a late Copper Age to early Bronze Age archaeological culture of the region between the Southern Bug, Dniester, and Ural rivers (the Pontic steppe), dating to 3300–2600 BCE.[2] It was discovered by Vasily Gorodtsov following his archaelogical excavations near Siversky Donets in 1901—1903. Its name derives from its characteristic burial tradition: Я́мная (romanization: yamnaya) is a Russian adjective that means 'related to pits (yama)', as these people used to bury their dead in tumuli (kurgans) containing simple pit chambers.

Yamnaya culture
Alternative names
  • Pit Grave culture
  • Yamna culture
  • Ochre Grave culture
  • Yamnaya Horizon
Geographical rangeEurasia
PeriodCopper Age, Bronze Age
Datesc. 3300–2600 BC
Preceded bySamara culture, Khvalynsk culture, Dnieper–Donets culture, Sredny Stog culture, Repin culture, Maykop culture, Cucuteni–Trypillia culture
Followed by
Defined byVasily Gorodtsov

The people of the Yamnaya culture were likely the result of a genetic admixture between the descendants of Eastern European Hunter-Gatherers (EHG)[a] and people related to hunter-gatherers from the Caucasus (CHG),[3] an ancestral component which is often named "Steppe ancestry", with additional admixture of up to 18% from Early European Farmers.[4] Their material culture was very similar to the Afanasevo culture, and the populations of both cultures are genetically indistinguishable.[1] They lived primarily as nomads, with a chiefdom system and wheeled carts and wagons that allowed them to manage large herds.

The people of the Yamnaya culture are also closely connected to Final Neolithic cultures, which later spread throughout Europe and Central Asia, especially the Corded Ware people and the Bell Beaker culture, as well as the peoples of the Sintashta, Andronovo, and Srubnaya cultures. Back migration from Corded Ware also contributed to Sintashta and Andronovo.[5] In these groups, several aspects of the Yamnaya culture are present.[b] Genetic studies have also indicated that these populations derived large parts of their ancestry from the steppes.[1][6][7][8]

The Yamnaya culture is identified with the late Proto-Indo-Europeans, and the Pontic-Caspian steppe is the strongest candidate for the Urheimat (original homeland) of the Proto-Indo-European language.

Origins

 
Corded ware pot, Yamnaya culture
 
Largest expansion of the Yamnaya culture.[image reference needed] C. 3500 origins of Usatovo culture; 3400 origins of Yamnaya; c. 3400–3200 expansion of Yamnaya across the Pontic-Caspian steppe; c. 3000 end of Tripolye culture,[citation needed] and transformation of Yamnaya into Corded Ware in the contact zone east of the Carpathian mountains; 3100–2600 Yamnaya expansion into the Danube Valley.[9][10][11]

Yamnaya culture was defined by Vasily Gorodtsov in order to differentiate it from catacomb and srubnaya cultures that existed in the area but considered to be of later period. Owing to the historical distance of Yamnaya culture, and reliance on archaeological findings, debate continues as to its origin. In 1996 Pavel Dolukhanov suggested that the emergence of the Pit-Grave culture represents a social development of various local Bronze Age cultures,[citation needed] representing "an expression of social stratification and the emergence of chiefdom-type nomadic social structures", which in turn intensified inter-group contacts between essentially heterogeneous social groups.[12]

According to Mallory (1999), "The origin of the Yamnaya culture is still a topic of debate," with proposals for its origins pointing to both Khvalynsk and Sredny Stog.[13] The Khvalynsk culture (4700–3800 BC)[14] (middle Volga) and the Don-based Repin culture (c. 3950–3300 BC)[15] in the eastern Pontic-Caspian steppe, and the closely related Sredny Stog culture (c. 4500–3500 BC) in the western Pontic-Caspian steppe, preceded the Yamnaya culture (3300–2500 BC).[16][17]

Further efforts to pinpoint the location came from Anthony (2007), suggesting from his research that the Yamnaya culture (3300–2600 BC) originated in the DonVolga area at c. 3400 BC,[18][2] preceded by the middle Volga-based Khvalynsk culture and the Don-based Repin culture (c. 3950–3300 BC),[15][2] arguing that late pottery from these two cultures can barely be distinguished from early Yamnaya pottery.[19] Earlier continuity from eneolithic but largely hunter-gatherer Samara culture and influences from the more agricultural Dnieper–Donets II are apparent.

He argues that the early Yamnaya horizon spread quickly across the Pontic–Caspian steppes between c. 3400 and 3200 BC:[18]

The spread of the Yamnaya horizon was the material expression of the spread of late Proto-Indo-European across the Pontic–Caspian steppes.[20]
[...] The Yamnaya horizon is the visible archaeological expression of a social adjustment to high mobility – the invention of the political infrastructure to manage larger herds from mobile homes based in the steppes.[21]

Alternatively, Parpola (2015) relates both the Corded ware culture and the Yamnaya culture to the late Tripolye culture.[22] He hypothesizes that "the Tripolye culture was taken over by PIE speakers by c. 4000 BC,"[23] and that in its final phase the Tripolye culture expanded to the steppes, morphing into various regional cultures which fused with the late Sredny Stog pastoralist cultures, which, he suggests, gave rise to the Yamnaya culture.[24]

The Yamnaya (Pit-grave) culture was succeeded in its western range by the Catacomb culture (2800–2200 BC); in the east, by the Poltavka culture (2700–2100 BC) at the middle Volga. These two cultures were followed by the Srubnaya culture (18th–12th century BC).

Characteristics

 
Yamnaya culture grave, Volgograd Oblast

The Yamnaya culture was semi-nomadic, with some agriculture practiced near rivers, and a few fortified sites, the largest of which is Mikhaylovka.[25]

Characteristic for the culture are the burials in pit graves under kurgans (tumuli), often accompanied with animal offerings. Some graves contain large anthropomorphic stelae, with carved human heads, arms, hands, belts, and weapons.[26] The dead bodies were placed in a supine position with bent knees and covered in ochre. Some kurgans contained "stratified sequences of graves".[27] It has been argued that kurgan burials were rare, and reserved for special adults, who were predominantly, but not necessarily, male.[28] Status and gender are marked by grave goods and position, and in some areas, elite individuals are buried with complete wooden wagons.[29] Grave goods are more common in eastern Yamnaya burials, which are also characterized by a higher proportion of male burials and more male-centred rituals than western areas.[30]

In the northern Pontic steppes were excavated the oldest wheels in the world, which may tentatively be associated with the Indo-Europeans.[31] The Yamnaya culture had and used two-wheeled carts and four-wheeled wagons, which are thought to have been oxen-drawn at this time, and there is evidence that they rode horses.[32]

Metallurgists and other craftsmen are given a special status in Yamnaya society, and metal objects are sometimes found in large quantities in elite graves. New metalworking technologies and weapon designs are used.[29]

Anthony[33] speculates that the Yamnaya ate a diet consisting of meat, milk, yogurt, cheese, and soups made from seeds and wild vegetables, and probably consumed mead.

Mallory and Adams suggest that Yamnaya society may have had a tripartite structure of three differentiated social classes, although the evidence available does not demonstrate the existence of specific classes such as priests, warriors, and farmers.[34]

Archaeogenetics

According to Jones et al. (2015) and Haak et al. (2015), autosomal tests indicate that the Yamnaya people were the result of a genetic admixture between two different hunter-gatherer populations: distinctive "Eastern Hunter-Gatherers" (EHG), from Eastern Europe, with high affinity to the Mal'ta–Buret' culture or other, closely related people from Siberia[6] and a population of "Caucasus hunter-gatherers" (CHG) who probably arrived from the Caucasus[35][3] or Iran.[36] Each of those two populations contributed about half the Yamnaya DNA.[7][3] This admixture is referred to in archaeogenetics as Western Steppe Herder (WSH) ancestry.

Admixture between EHGs and CHGs is believed to have occurred on the eastern Pontic-Caspian steppe starting around 5,000 BC, while admixture with Early European Farmers (EEF) happened in the southern parts of the Pontic-Caspian steppe sometime later. More recent genetic studies have found that the Yamnaya were a mixture of EHGs, CHGs, and to a lesser degree Anatolian farmers and Levantine farmers, but not EEFs from Europe due to lack of WHG DNA in the Yamnaya. This occurred in two distinct admixture events from West Asia into the Pontic-Caspian steppe.[37][38]

Haplogroup R1b, especially subclades of R1b-M269, is the most common Y-DNA haplogroup found among both the Yamnaya and modern-day Western Europeans.[1][6] Additionally, a minority are found to belong to haplogroup I2.[7] They are found to belong to a wider variety of mtDNA haplogroups, including U, T, and haplogroups associated with Caucasus Hunter-Gatherers and Early European Farmers.[39][40]

People of the Yamnaya culture are believed to have had mostly brown eye colour, light to intermediate skin, and brown hair colour, with some variation.[41][42] A 2022 study by Lazaridis et al. found that the typical phenotype among the Yamnaya population was brown eyes, brown hair, and intermediate skin colour. None of the Yamnaya samples were predicted to have either blue eyes or blonde hair.[43] Some individuals are believed to have carried a mutation to the KITLG gene associated with blond hair, as several individuals with Steppe ancestry are later found to carry this mutation. The Ancient North Eurasian population, who contributed significant ancestry to Western Steppe Herders, are believed to be the source of this mutation.[44] A study in 2015 found that Yamnaya had the highest ever calculated genetic selection for height of any of the ancient populations tested.[45][46] It has been hypothesized that an allele associated with lactase persistence (conferring lactose tolerance into adulthood) was brought to Europe from the steppe by Yamnaya-related migrations.[47][48][49][50]

The geneticist David Reich has argued that the genetic data supports the likelihood that the people of the Yamnaya culture were a "single, genetically coherent group" who were responsible for spreading many Indo-European languages.[51] Reich also argues that the genetic evidence shows that Yamnaya society was an oligarchy dominated by a small number of elite males.[52]

The genetic evidence for the extent of the role of the Yamnaya culture in the spread of Indo-European languages has however been questioned by Russian archaeologist Leo Klejn[53] and Balanovsky et al.,[54] who note a lack of male haplogroup continuity between the people of the Yamnaya culture and the contemporary populations of Europe. Klejn has also suggested that the autosomal evidence does not support a Yamnaya migration, arguing that Western Steppe Herder ancestry in both contemporary and Bronze Age samples is lowest around the Danube in Hungary, near the western limits of the Yamnaya culture, and highest in Northern Europe, which Klejn argues is the opposite of what would be expected if the geneticists' hypothesis is correct.[55]

Language

Marija Gimbutas identified the Yamnaya culture with the late Proto-Indo-Europeans (PIE) in her Kurgan hypothesis. The Pontic-Caspian steppe is the strongest candidate for the Urheimat (original homeland) of the Proto-Indo-European language, and evidence from linguistics[56] and genetics[6][57] suggests that the Yamnaya culture may be the homeland of the core Indo-European languages, excluding the Anatolian languages.[58]

According to David W. Anthony, the genetic evidence suggests that the leading clans of the Yamnaya were of EHG (Eastern European hunter-gatherer) and WHG (Western European hunter-gatherer) paternal origin[59] and implies that the Indo-European languages were the result of "a dominant language spoken by EHGs that absorbed Caucasus-like elements in phonology, morphology, and lexicon."[60]

Yamnaya-related migrations

 
Expansion of Yamnaya-related people, according to Anthony (2007),[56] 2017;[61] Narasimhan et al. (2019);[62] Nordqvist and Heyd (2020):[10] * 3000 BC: Initial eastward migration initiating the Afanasievo culture, possibly Proto-Tocharian. * 2900 BC: North-westward migrations carrying Corded Ware culture, transforming into Bell Beaker; according to Anthony, westward migration west of Carpatians into Hungary as Yamnaya, transforming into Bell Beaker, possibly ancestral to Italo-Celtic (disputed). * 2700 BC: Second eastward migration starting east of Carpatian mountains as Corded Ware, transforming into Fatyanovo-Balanova (2800 BC) → Abashevo (2200 BC) → Sintashta (2100–1900 BC) → Andronovo (1900–1700 BC) → Indo-Aryans.

Western Europe

Haak et al. (2015) conducted a genome-wide study of 69 ancient skeletons from Europe and Russia. They concluded that Yamnaya autosomal characteristics are very close to the Corded Ware culture people, with an estimated 73% ancestral contribution from the Yamnaya DNA in the DNA of Corded Ware skeletons from Germany. The same study estimated a (38.8–50.4 %) ancestral contribution of the Yamnaya in the DNA of modern Central, and Northern Europeans, and an 18.5–32.6 % contribution in modern Southern Europeans; this contribution is found to a lesser extent in Sardinians (2.4–7.1 %) and Sicilians (5.9–11.6 %).[63][57][8] Haak et al. also note that their results state that haplogroup R-M269 spread into Europe from the East after 3000 BC.[64] Studies that analysed ancient human remains in Ireland and Portugal support the thesis that R-M269 was introduced in these places along with autosomal DNA from the Eastern European steppes.[65][66]

Autosomal tests also indicate that the Yamnaya are the most likely vector for "Ancient North Eurasian" admixture into Europe.[6] "Ancient North Eurasian" is the name given in literature to a genetic component that represents descent from the people of the Mal'ta–Buret' culture[6] or a population closely related to them. That genetic component is visible in tests of the Yamnaya people[6] as well as modern-day Europeans, but not of Europeans predating the Bronze Age.[67]

Eastern Europe and Finland

 
According to Allentoft (2015), the Sintashta culture probably derived from the Corded Ware Culture.

In the Baltic, Jones et al. (2017) found that the Neolithic transition – the passage from a hunter-gatherer economy to a farming-based economy – coincided with the arrival en masse of individuals with Yamnaya-like ancestry. This is different from what happened in Western and Southern Europe, where the Neolithic transition was caused by a population that came from Anatolia, with Pontic steppe ancestry being detected from only the late Neolithic onward.[68]

Per Haak et al. (2015), the Yamnaya contribution in the modern populations of Eastern Europe ranges from 46.8% among Russians to 42.8% in Ukrainians. Finland has one of the highest Yamnaya contributions in all of Europe (50.4%).[69][c]

Central and South Asia

 
Map of the approximate maximal extent of the Andronovo culture. The formative Sintashta-Petrovka culture is shown in darker red. The location of the earliest spoke-wheeled chariot finds is indicated in purple. Adjacent and overlapping cultures (Afanasevo, Srubna, Bactria-Margiana Culture are shown in green.
 
Archaeological cultures associated with Indo-Iranian migrations and Indo-Aryan migrations (after EIEC). The Andronovo, BMAC and Yaz cultures have often been associated with Indo-Iranian migrations. The GGC, Cemetery H, Copper Hoard and PGW cultures are candidates for cultures associated with Indo-Aryan migrations.

Studies also point to the strong presence of Yamnaya descent in the current nations of South Asia, especially in groups that are referred to as Indo-Aryans.[62][70] According to Pathak et al. (2018), the "North-Western Indian & Pakistani" populations (PNWI) showed significant Middle-Late Bronze Age Steppe (Steppe_MLBA) ancestry along with Yamnaya Early-Middle Bronze Age (Steppe_EMBA) ancestry, but the Indo-Europeans of Gangetic Plains and Dravidian people only showed significant Yamnaya (Steppe_EMBA) ancestry and no Steppe_MLBA. The study also noted that ancient south Asian samples had significantly higher Steppe_MLBA than Steppe_EMBA (or Yamnaya). The study identified the Rors and Jats as the population in South Asia with the highest proportion of Steppe ancestry.[70] Lazaridis et al. (2016) estimated (6.5–50.2 %) steppe-related admixture in South Asians, though the proportion of Steppe ancestry varies widely across ethnic groups.[36][d] According to Narasimhan et al. (2019), the Yamnaya-related ancestry, termed Western_Steppe_EMBA, that reached central and south Asia was not the initial expansion from the steppe to the east, but a secondary expansion that involved a group possessing ~67% Western_Steppe_EMBA ancestry and ~33% ancestry from the European cline. This group included people similar to that of Corded Ware, Srubnaya, Petrovka, and Sintashta. Moving further east in the central steppe, it acquired ~9% ancestry from a group of people that possessed West Siberian Hunter Gatherer ancestry, thus forming the Central Steppe MLBA cluster, which is the primary source of steppe ancestry in South Asia, contributing up to 30% of the ancestry of the modern groups in the region.[62]

According to Unterländer et al. (2017), all Iron Age Scythian Steppe nomads can best be described as a mixture of Yamnaya-related ancestry and an East Asian-related component, which most closely corresponds to the modern North Siberian Nganasan people of the lower Yenisey River, to varying degrees, but generally higher among Eastern Scythians, while Bronze Age Steppe nomads of Central Asia had the East Asian-related component at only trace levels.[71]

Artifacts

See also

Notes

  1. ^ The Eastern European hunter-gatherers were themselves mostly descended from ancient North Eurasians, related to the palaeolithic Mal'ta–Buret' culture.
  2. ^ Yamnayan cultural aspects, for example, were horse-riding, burial styles, and to some extent the pastoralist economy.
  3. ^ Per Haak et al. (2015), adding a north-Siberian people as a fourth reference population improves residuals for northeastern European populations. This accounts for the higher than expected Yamnaya contribution and brings it down to expected levels (67.8–50.4 % in Finns, 64.9–46.8 % in Russians).
  4. ^ Lazaridis et al. (2016) Supplementary Information, Table S9.1: "Kalash – 50.2 %, Tiwari Brahmins – 44.1 %, Gujarati (four samples) – 46.1 % to 27.5 %, Pathan – 44.6 %, Burusho – 42.5 %, Sindhi – 37.7 %, Punjabi – 32.6 %, Balochi – 32.4 %, Brahui – 30.2 %, Lodhi – 29.3 %, Bengali – 24.6 %, Vishwabhramin – 20.4 %, Makrani – 19.2 %, Mala – 18.4 %, Kusunda – 8.9 %, Kharia – 6.5 %."

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  65. ^ Cassidy et al. 2016.
  66. ^ Martiniano, et al. 2017.
  67. ^ Lazaridis et al. 2014.
  68. ^ Jones et al. 2017.
  69. ^ Haak et al. 2015, pp. 121–122.
  70. ^ a b Pathak et al. 2018.
  71. ^ Unterländer et al. 2017Genomic inference reveals that Scythians in the east and the west of the steppe zone can best be described as a mixture of Yamnaya-related ancestry and an East Asian component. Demographic modelling suggests independent origins for eastern and western groups with ongoing gene-flow between them, plausibly explaining the striking uniformity of their material culture. We also find evidence that significant gene-flow from east to west Eurasia must have occurred early during the Iron Age." and "The blend of EHG [European hunter-gatherer] and Caucasian elements in carriers of the Yamnaya culture was formed on the European steppe and exported into Central Asia and Siberia". We therefore considered an alternative model in which we treat them as a mix of Yamnaya and the Han (Supplementary Table 25). This model fits all of the Iron Age Scythian groups, consistent with these groups having ancestry related to East Asians not found in the other populations. Alternatively, the Iron Age Scythian groups can also be modelled as a mix of Yamnaya and the north Siberian Nganasan (Supplementary Note 2, Supplementary Table 26).

Notes

Sources

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External links

  • "Genetic study revives debate on origin and expansion of Indo-European Languages". Science Daily. March 2015.
  • "First Horse Warriors". PBS.org NOVA series. 15 May 2019.

yamnaya, culture, yamna, culture, redirects, here, confused, with, yamna, language, papua, indonesia, yamna, culture, russian, Ямная, культура, ukrainian, Ямна, культура, culture, pits, also, known, grave, culture, ochre, grave, culture, late, copper, early, b. Yamna culture redirects here Not to be confused with the Yamna language of Papua Indonesia The Yamnaya culture or the Yamna culture Russian Yamnaya kultura Ukrainian Yamna kultura lit culture of pits also known as the Pit Grave culture or Ochre Grave culture was a late Copper Age to early Bronze Age archaeological culture of the region between the Southern Bug Dniester and Ural rivers the Pontic steppe dating to 3300 2600 BCE 2 It was discovered by Vasily Gorodtsov following his archaelogical excavations near Siversky Donets in 1901 1903 Its name derives from its characteristic burial tradition Ya mnaya romanization yamnaya is a Russian adjective that means related to pits yama as these people used to bury their dead in tumuli kurgans containing simple pit chambers Yamnaya cultureAlternative namesPit Grave culture Yamna culture Ochre Grave culture Yamnaya HorizonGeographical rangeEurasiaPeriodCopper Age Bronze AgeDatesc 3300 2600 BCPreceded bySamara culture Khvalynsk culture Dnieper Donets culture Sredny Stog culture Repin culture Maykop culture Cucuteni Trypillia cultureFollowed byWest Catacomb culture East Poltavka culture North Corded Ware culture possibly from Yamnaya culture 1 Defined byVasily GorodtsovThe people of the Yamnaya culture were likely the result of a genetic admixture between the descendants of Eastern European Hunter Gatherers EHG a and people related to hunter gatherers from the Caucasus CHG 3 an ancestral component which is often named Steppe ancestry with additional admixture of up to 18 from Early European Farmers 4 Their material culture was very similar to the Afanasevo culture and the populations of both cultures are genetically indistinguishable 1 They lived primarily as nomads with a chiefdom system and wheeled carts and wagons that allowed them to manage large herds The people of the Yamnaya culture are also closely connected to Final Neolithic cultures which later spread throughout Europe and Central Asia especially the Corded Ware people and the Bell Beaker culture as well as the peoples of the Sintashta Andronovo and Srubnaya cultures Back migration from Corded Ware also contributed to Sintashta and Andronovo 5 In these groups several aspects of the Yamnaya culture are present b Genetic studies have also indicated that these populations derived large parts of their ancestry from the steppes 1 6 7 8 The Yamnaya culture is identified with the late Proto Indo Europeans and the Pontic Caspian steppe is the strongest candidate for the Urheimat original homeland of the Proto Indo European language Contents 1 Origins 2 Characteristics 3 Archaeogenetics 4 Language 5 Yamnaya related migrations 5 1 Western Europe 5 2 Eastern Europe and Finland 5 3 Central and South Asia 6 Artifacts 7 See also 8 Notes 9 References 10 Notes 11 Sources 12 External linksOrigins Edit Corded ware pot Yamnaya culture Largest expansion of the Yamnaya culture image reference needed C 3500 origins of Usatovo culture 3400 origins of Yamnaya c 3400 3200 expansion of Yamnaya across the Pontic Caspian steppe c 3000 end of Tripolye culture citation needed and transformation of Yamnaya into Corded Ware in the contact zone east of the Carpathian mountains 3100 2600 Yamnaya expansion into the Danube Valley 9 10 11 See also Kurgan hypothesis and Marija Gimbutas Yamnaya culture was defined by Vasily Gorodtsov in order to differentiate it from catacomb and srubnaya cultures that existed in the area but considered to be of later period Owing to the historical distance of Yamnaya culture and reliance on archaeological findings debate continues as to its origin In 1996 Pavel Dolukhanov suggested that the emergence of the Pit Grave culture represents a social development of various local Bronze Age cultures citation needed representing an expression of social stratification and the emergence of chiefdom type nomadic social structures which in turn intensified inter group contacts between essentially heterogeneous social groups 12 According to Mallory 1999 The origin of the Yamnaya culture is still a topic of debate with proposals for its origins pointing to both Khvalynsk and Sredny Stog 13 The Khvalynsk culture 4700 3800 BC 14 middle Volga and the Don based Repin culture c 3950 3300 BC 15 in the eastern Pontic Caspian steppe and the closely related Sredny Stog culture c 4500 3500 BC in the western Pontic Caspian steppe preceded the Yamnaya culture 3300 2500 BC 16 17 Further efforts to pinpoint the location came from Anthony 2007 suggesting from his research that the Yamnaya culture 3300 2600 BC originated in the Don Volga area at c 3400 BC 18 2 preceded by the middle Volga based Khvalynsk culture and the Don based Repin culture c 3950 3300 BC 15 2 arguing that late pottery from these two cultures can barely be distinguished from early Yamnaya pottery 19 Earlier continuity from eneolithic but largely hunter gatherer Samara culture and influences from the more agricultural Dnieper Donets II are apparent He argues that the early Yamnaya horizon spread quickly across the Pontic Caspian steppes between c 3400 and 3200 BC 18 The spread of the Yamnaya horizon was the material expression of the spread of late Proto Indo European across the Pontic Caspian steppes 20 The Yamnaya horizon is the visible archaeological expression of a social adjustment to high mobility the invention of the political infrastructure to manage larger herds from mobile homes based in the steppes 21 Alternatively Parpola 2015 relates both the Corded ware culture and the Yamnaya culture to the late Tripolye culture 22 He hypothesizes that the Tripolye culture was taken over by PIE speakers by c 4000 BC 23 and that in its final phase the Tripolye culture expanded to the steppes morphing into various regional cultures which fused with the late Sredny Stog pastoralist cultures which he suggests gave rise to the Yamnaya culture 24 The Yamnaya Pit grave culture was succeeded in its western range by the Catacomb culture 2800 2200 BC in the east by the Poltavka culture 2700 2100 BC at the middle Volga These two cultures were followed by the Srubnaya culture 18th 12th century BC Maps of the origins of Yamnaya culture Sredny Stog culture c 4500 3500 BC Usatovo culture c 3500 3000 BC Khvalynsk culture c 4900 3500 BC Early Yamnaya culture 3400 BC according to Anthony 2007 Characteristics Edit Yamnaya culture grave Volgograd Oblast The Yamnaya culture was semi nomadic with some agriculture practiced near rivers and a few fortified sites the largest of which is Mikhaylovka 25 Characteristic for the culture are the burials in pit graves under kurgans tumuli often accompanied with animal offerings Some graves contain large anthropomorphic stelae with carved human heads arms hands belts and weapons 26 The dead bodies were placed in a supine position with bent knees and covered in ochre Some kurgans contained stratified sequences of graves 27 It has been argued that kurgan burials were rare and reserved for special adults who were predominantly but not necessarily male 28 Status and gender are marked by grave goods and position and in some areas elite individuals are buried with complete wooden wagons 29 Grave goods are more common in eastern Yamnaya burials which are also characterized by a higher proportion of male burials and more male centred rituals than western areas 30 In the northern Pontic steppes were excavated the oldest wheels in the world which may tentatively be associated with the Indo Europeans 31 The Yamnaya culture had and used two wheeled carts and four wheeled wagons which are thought to have been oxen drawn at this time and there is evidence that they rode horses 32 Metallurgists and other craftsmen are given a special status in Yamnaya society and metal objects are sometimes found in large quantities in elite graves New metalworking technologies and weapon designs are used 29 Anthony 33 speculates that the Yamnaya ate a diet consisting of meat milk yogurt cheese and soups made from seeds and wild vegetables and probably consumed mead Mallory and Adams suggest that Yamnaya society may have had a tripartite structure of three differentiated social classes although the evidence available does not demonstrate the existence of specific classes such as priests warriors and farmers 34 Archaeogenetics EditFurther information Western Steppe Herders The Kernosivsky idol According to Jones et al 2015 and Haak et al 2015 autosomal tests indicate that the Yamnaya people were the result of a genetic admixture between two different hunter gatherer populations distinctive Eastern Hunter Gatherers EHG from Eastern Europe with high affinity to the Mal ta Buret culture or other closely related people from Siberia 6 and a population of Caucasus hunter gatherers CHG who probably arrived from the Caucasus 35 3 or Iran 36 Each of those two populations contributed about half the Yamnaya DNA 7 3 This admixture is referred to in archaeogenetics as Western Steppe Herder WSH ancestry Admixture between EHGs and CHGs is believed to have occurred on the eastern Pontic Caspian steppe starting around 5 000 BC while admixture with Early European Farmers EEF happened in the southern parts of the Pontic Caspian steppe sometime later More recent genetic studies have found that the Yamnaya were a mixture of EHGs CHGs and to a lesser degree Anatolian farmers and Levantine farmers but not EEFs from Europe due to lack of WHG DNA in the Yamnaya This occurred in two distinct admixture events from West Asia into the Pontic Caspian steppe 37 38 Haplogroup R1b especially subclades of R1b M269 is the most common Y DNA haplogroup found among both the Yamnaya and modern day Western Europeans 1 6 Additionally a minority are found to belong to haplogroup I2 7 They are found to belong to a wider variety of mtDNA haplogroups including U T and haplogroups associated with Caucasus Hunter Gatherers and Early European Farmers 39 40 People of the Yamnaya culture are believed to have had mostly brown eye colour light to intermediate skin and brown hair colour with some variation 41 42 A 2022 study by Lazaridis et al found that the typical phenotype among the Yamnaya population was brown eyes brown hair and intermediate skin colour None of the Yamnaya samples were predicted to have either blue eyes or blonde hair 43 Some individuals are believed to have carried a mutation to the KITLG gene associated with blond hair as several individuals with Steppe ancestry are later found to carry this mutation The Ancient North Eurasian population who contributed significant ancestry to Western Steppe Herders are believed to be the source of this mutation 44 A study in 2015 found that Yamnaya had the highest ever calculated genetic selection for height of any of the ancient populations tested 45 46 It has been hypothesized that an allele associated with lactase persistence conferring lactose tolerance into adulthood was brought to Europe from the steppe by Yamnaya related migrations 47 48 49 50 The geneticist David Reich has argued that the genetic data supports the likelihood that the people of the Yamnaya culture were a single genetically coherent group who were responsible for spreading many Indo European languages 51 Reich also argues that the genetic evidence shows that Yamnaya society was an oligarchy dominated by a small number of elite males 52 The genetic evidence for the extent of the role of the Yamnaya culture in the spread of Indo European languages has however been questioned by Russian archaeologist Leo Klejn 53 and Balanovsky et al 54 who note a lack of male haplogroup continuity between the people of the Yamnaya culture and the contemporary populations of Europe Klejn has also suggested that the autosomal evidence does not support a Yamnaya migration arguing that Western Steppe Herder ancestry in both contemporary and Bronze Age samples is lowest around the Danube in Hungary near the western limits of the Yamnaya culture and highest in Northern Europe which Klejn argues is the opposite of what would be expected if the geneticists hypothesis is correct 55 Language EditMarija Gimbutas identified the Yamnaya culture with the late Proto Indo Europeans PIE in her Kurgan hypothesis The Pontic Caspian steppe is the strongest candidate for the Urheimat original homeland of the Proto Indo European language and evidence from linguistics 56 and genetics 6 57 suggests that the Yamnaya culture may be the homeland of the core Indo European languages excluding the Anatolian languages 58 According to David W Anthony the genetic evidence suggests that the leading clans of the Yamnaya were of EHG Eastern European hunter gatherer and WHG Western European hunter gatherer paternal origin 59 and implies that the Indo European languages were the result of a dominant language spoken by EHGs that absorbed Caucasus like elements in phonology morphology and lexicon 60 Yamnaya related migrations EditSee also Indo European migrations Expansion of Yamnaya related people according to Anthony 2007 56 2017 61 Narasimhan et al 2019 62 Nordqvist and Heyd 2020 10 3000 BC Initial eastward migration initiating the Afanasievo culture possibly Proto Tocharian 2900 BC North westward migrations carrying Corded Ware culture transforming into Bell Beaker according to Anthony westward migration west of Carpatians into Hungary as Yamnaya transforming into Bell Beaker possibly ancestral to Italo Celtic disputed 2700 BC Second eastward migration starting east of Carpatian mountains as Corded Ware transforming into Fatyanovo Balanova 2800 BC Abashevo 2200 BC Sintashta 2100 1900 BC Andronovo 1900 1700 BC Indo Aryans Western Europe Edit See also Corded Ware culture Haak et al 2015 conducted a genome wide study of 69 ancient skeletons from Europe and Russia They concluded that Yamnaya autosomal characteristics are very close to the Corded Ware culture people with an estimated 73 ancestral contribution from the Yamnaya DNA in the DNA of Corded Ware skeletons from Germany The same study estimated a 38 8 50 4 ancestral contribution of the Yamnaya in the DNA of modern Central and Northern Europeans and an 18 5 32 6 contribution in modern Southern Europeans this contribution is found to a lesser extent in Sardinians 2 4 7 1 and Sicilians 5 9 11 6 63 57 8 Haak et al also note that their results state that haplogroup R M269 spread into Europe from the East after 3000 BC 64 Studies that analysed ancient human remains in Ireland and Portugal support the thesis that R M269 was introduced in these places along with autosomal DNA from the Eastern European steppes 65 66 Autosomal tests also indicate that the Yamnaya are the most likely vector for Ancient North Eurasian admixture into Europe 6 Ancient North Eurasian is the name given in literature to a genetic component that represents descent from the people of the Mal ta Buret culture 6 or a population closely related to them That genetic component is visible in tests of the Yamnaya people 6 as well as modern day Europeans but not of Europeans predating the Bronze Age 67 Eastern Europe and Finland Edit According to Allentoft 2015 the Sintashta culture probably derived from the Corded Ware Culture In the Baltic Jones et al 2017 found that the Neolithic transition the passage from a hunter gatherer economy to a farming based economy coincided with the arrival en masse of individuals with Yamnaya like ancestry This is different from what happened in Western and Southern Europe where the Neolithic transition was caused by a population that came from Anatolia with Pontic steppe ancestry being detected from only the late Neolithic onward 68 Per Haak et al 2015 the Yamnaya contribution in the modern populations of Eastern Europe ranges from 46 8 among Russians to 42 8 in Ukrainians Finland has one of the highest Yamnaya contributions in all of Europe 50 4 69 c Central and South Asia Edit See also Sintashta culture Map of the approximate maximal extent of the Andronovo culture The formative Sintashta Petrovka culture is shown in darker red The location of the earliest spoke wheeled chariot finds is indicated in purple Adjacent and overlapping cultures Afanasevo Srubna Bactria Margiana Culture are shown in green Archaeological cultures associated with Indo Iranian migrations and Indo Aryan migrations after EIEC The Andronovo BMAC and Yaz cultures have often been associated with Indo Iranian migrations The GGC Cemetery H Copper Hoard and PGW cultures are candidates for cultures associated with Indo Aryan migrations Studies also point to the strong presence of Yamnaya descent in the current nations of South Asia especially in groups that are referred to as Indo Aryans 62 70 According to Pathak et al 2018 the North Western Indian amp Pakistani populations PNWI showed significant Middle Late Bronze Age Steppe Steppe MLBA ancestry along with Yamnaya Early Middle Bronze Age Steppe EMBA ancestry but the Indo Europeans of Gangetic Plains and Dravidian people only showed significant Yamnaya Steppe EMBA ancestry and no Steppe MLBA The study also noted that ancient south Asian samples had significantly higher Steppe MLBA than Steppe EMBA or Yamnaya The study identified the Rors and Jats as the population in South Asia with the highest proportion of Steppe ancestry 70 Lazaridis et al 2016 estimated 6 5 50 2 steppe related admixture in South Asians though the proportion of Steppe ancestry varies widely across ethnic groups 36 d According to Narasimhan et al 2019 the Yamnaya related ancestry termed Western Steppe EMBA that reached central and south Asia was not the initial expansion from the steppe to the east but a secondary expansion that involved a group possessing 67 Western Steppe EMBA ancestry and 33 ancestry from the European cline This group included people similar to that of Corded Ware Srubnaya Petrovka and Sintashta Moving further east in the central steppe it acquired 9 ancestry from a group of people that possessed West Siberian Hunter Gatherer ancestry thus forming the Central Steppe MLBA cluster which is the primary source of steppe ancestry in South Asia contributing up to 30 of the ancestry of the modern groups in the region 62 According to Unterlander et al 2017 all Iron Age Scythian Steppe nomads can best be described as a mixture of Yamnaya related ancestry and an East Asian related component which most closely corresponds to the modern North Siberian Nganasan people of the lower Yenisey River to varying degrees but generally higher among Eastern Scythians while Bronze Age Steppe nomads of Central Asia had the East Asian related component at only trace levels 71 Artifacts EditFrom the Hermitage Museum collections Corded ware pot Bronze daggers arrowheads and bone artefactsSee also EditKurgan Kurgan stelae Butmir culture Vinca culture Beaker culture Baden culture Botai culture Khvalynsk culture Mamai Hora Samara culture Sintashta culture Yersinia pestis Proto Indo Europeans Indigenous AryanismNotes Edit The Eastern European hunter gatherers were themselves mostly descended from ancient North Eurasians related to the palaeolithic Mal ta Buret culture Yamnayan cultural aspects for example were horse riding burial styles and to some extent the pastoralist economy Per Haak et al 2015 adding a north Siberian people as a fourth reference population improves residuals for northeastern European populations This accounts for the higher than expected Yamnaya contribution and brings it down to expected levels 67 8 50 4 in Finns 64 9 46 8 in Russians Lazaridis et al 2016 Supplementary Information Table S9 1 Kalash 50 2 Tiwari Brahmins 44 1 Gujarati four samples 46 1 to 27 5 Pathan 44 6 Burusho 42 5 Sindhi 37 7 Punjabi 32 6 Balochi 32 4 Brahui 30 2 Lodhi 29 3 Bengali 24 6 Vishwabhramin 20 4 Makrani 19 2 Mala 18 4 Kusunda 8 9 Kharia 6 5 References Edit a b c d Allentoft 2015 a b c Morgunova amp Khokhlova 2013 a b c Europe s fourth ancestral tribe uncovered BBC 16 November 2015 Wang Chuan Chao Reinhold Sabine Kalmykov Alexey Wissgott Antje Brandt Guido Jeong Choongwon Cheronet Olivia Ferry Matthew Harney Eadaoin Keating Denise Mallick Swapan 4 February 2019 Ancient human genome wide data from a 3000 year interval in the Caucasus corresponds with eco geographic regions Nature Communications 10 1 590 Bibcode 2019NatCo 10 590W doi 10 1038 s41467 018 08220 8 ISSN 2041 1723 PMC 6360191 PMID 30713341 Novembre 2015 evidence to support theories of a back migration from Corded Ware related populations that contributed to the origins of the Sintashta culture in the Urals and their descendants the Andronovo a b c d e f g Haak et al 2015 a b c Mathieson et al 2015 a b Gibbons Ann 10 June 2015 Nomadic herders left a strong genetic mark on Europeans and Asians Science AAAS Anthony 2007 p 300 370 a b Nordgvist amp Heyd 2020 Mallory 1999 Dolukhanov 1996 p 94 Mallory 1999 p 215 Anthony 2007 p 182 a b Anthony 2007 p 275 Anthony 2007 p 300 Mallory 1999 p 210 211 a b Anthony 2007 p 321 Anthony 2007 pp 274 277 317 320 Anthony 2007 pp 301 302 Anthony 2007 p 303 Parpola 2015 p 49 Parpola 2015 p 45 Parpola 2015 p 47 Mallory 1997 p 212 Anthony 2007 p 339 Anthony 2007 p 319 author Anthony David The Horse the Wheel and Language OCLC 1102387902 a href Template Cite book html title Template Cite book cite book a last has generic name help a b J Harrison Richard The transformation of Europe in the third millennium BC the example of Le Petit Chasseur I II Sion Valais Switzerland p 196 OCLC 718304072 Anthony 2007 p 305 Holm Hans J J G 2019 The Earliest Wheel Finds their Archeology and Indo European Terminology in Time and Space and Early Migrations around the Caucasus Series Minor 43 Budapest ARCHAEOLINGUA ALAPITVANY ISBN 978 615 5766 30 5 With 306 references six greyscaled and coloured images and miniature images within the table of 130 representative finds including new finds in Germany and Western China P Mallory J 2003 1989 In search of the Indo Europeans language archaeology and myth Thames and Hudson p 213 ISBN 0 500 27616 1 OCLC 886668216 Anthony 2007 p 430 J P Mallory Douglas Q Adams eds 1997 Encyclopedia of Indo European culture London Fitzroy Dearborn p 653 ISBN 1 884964 98 2 OCLC 37931209 Jones et al 2015 a b Lazaridis et al 2016 Lazaridis Iosif Alpaslan Roodenberg Songul Acar Ayse Acikkol Aysen Agelarakis Anagnostis Aghikyan Levon Akyuz Ugur Andreeva Desislava Andrijasevic Gojko Antonovic Dragana Armit Ian Atmaca Alper Avetisyan Pavel Aytek Ahmet Ihsan Bacvarov Krum 26 August 2022 The genetic history of the Southern Arc A bridge between West Asia and Europe Science 377 6609 eabm4247 doi 10 1126 science abm4247 ISSN 0036 8075 PMID 36007055 S2CID 251843620 Chintalapati Manjusha Patterson Nick Moorjani Priya 30 May 2022 Perry George H ed The spatiotemporal patterns of major human admixture events during the European Holocene eLife 11 e77625 doi 10 7554 eLife 77625 ISSN 2050 084X PMC 9293011 PMID 35635751 Wang 2019 Eske Allentoft Morten E Pokutta Dalia Willerslev 2015 Population genomics of Bronze Age Eurasia University of Copenhagen Denmark OCLC 1234973657 Gibbons A 24 July 2015 Revolution in human evolution Science 349 6246 362 366 Bibcode 2015Sci 349 362G doi 10 1126 science 349 6246 362 ISSN 0036 8075 PMID 26206910 Hanel Andrea Carlberg Carsten September 2020 Skin colour and vitamin D An update Experimental Dermatology 29 9 864 875 doi 10 1111 exd 14142 ISSN 0906 6705 PMID 32621306 Lazaridis et al 2022c Mathieson Iain Alpaslan Roodenberg Songul Posth Cosimo Szecsenyi Nagy Anna Rohland Nadin Mallick Swapan Olalde Inigo Broomandkhoshbacht Nasreen Candilio Francesca Cheronet Olivia Fernandes Daniel March 2018 The genomic history of southeastern Europe Nature 555 7695 197 203 Bibcode 2018Natur 555 197M doi 10 1038 nature25778 ISSN 1476 4687 PMC 6091220 PMID 29466330 Heyd Volker April 2017 Kossinna s smile Antiquity 91 356 348 359 doi 10 15184 aqy 2017 21 hdl 10138 255652 ISSN 0003 598X S2CID 164376362 Mathieson Iain Lazaridis Iosif Rohland Nadin Mallick Swapan Patterson Nick Roodenberg Songul Alpaslan Harney Eadaoin Stewardson Kristin Fernandes Daniel Novak Mario Sirak Kendra 24 December 2015 Genome wide patterns of selection in 230 ancient Eurasians Nature 528 7583 499 503 Bibcode 2015Natur 528 499M doi 10 1038 nature16152 ISSN 0028 0836 PMC 4918750 PMID 26595274 Segurel Laure 2020 Why and when was lactase persistence selected for Insights from Central Asian herders and ancient DNA PLOS Biology 18 6 e3000742 doi 10 1371 journal pbio 3000742 PMC 7302802 PMID 32511234 Furthermore ancient DNA studies found that the LP mutation was absent or very rare in Europe until the end of the Bronze Age 26 29 and appeared first in individuals with steppe ancestry 19 20 Thus it was proposed that the mutation originated in Yamnaya associated populations and arrived later in Europe by migration of these steppe herders Callaway Ewen DNA data explosion lights up the Bronze Age Nature the 101 sequenced individuals the Yamnaya were most likely to have the DNA variation responsible for lactose tolerance hinting that the steppe migrants might have eventually introduced the trait to Europe Furholt Martin 2018 Massive Migrations The Impact of Recent DNA Studies on our View of Third Millennium Europe European Journal of Archaeology 21 2 159 191 doi 10 1017 eaa 2017 43 For example one lineage could have a biological evolutionary advantage over the other Allentoft et al 2015 171 have found a remarkably high rate of lactose tolerance among individuals connected to Yamnaya and to Corded Ware as opposed to the majority of Late Neolithic individuals Saag L 2020 Human Genetics Lactase Persistence in a Battlefield Current Biology 30 21 R1311 R1313 doi 10 1016 j cub 2020 08 087 PMID 33142099 S2CID 226229587 Reich David 2018 Who we are and how we got here ancient DNA and the new science of the human past Oxford United Kingdom p 121 ISBN 978 0 19 882125 0 OCLC 1006478846 Reich David 2018 Who we are and how we got here ancient DNA and the new science of the human past Oxford United Kingdom p 239 ISBN 978 0 19 882125 0 OCLC 1006478846 Klejn Leo 2017 The Steppe Hypothesis of Indo European Origins Remains to be Proven Acta Archaeologica 88 1 193 204 doi 10 1111 j 1600 0390 2017 12184 x Balanovsky O Chukhryaeva M Zaporozhchenko V 2017 Genetic differentiation between upland and lowland populations shapes the Y chromosomal landscape of West Asia Human Genetics 136 4 437 450 doi 10 1007 s00439 017 1770 2 PMID 28281087 S2CID 3735168 The ancient Yamnaya samples are located on the eastern R GG400 branch of haplogroup R1b L23 showing that the paternal descendants of the Yamnaya still live in the Pontic steppe and that the ancient Yamnaya population was not an important source of paternal lineages in present day West Europeans Klejn 2017 p 201 In the tables presented in the article by Reichs team Haak et al 2015 the genetic pool connecting the Yamnaya culture with the Corded Ware people is shown to be more intense in Northern Europe Norway and Sweden and decreases gradually from the North to the South Fig 6 It is weakest around the Danube in Hungary i e areas neighbouring the western branch of the Yamnaya culture This is the reverse image to what the proposed hypothesis by the geneticists would lead us to expect It is true that this gradient is traced back from the contemporary materials but it was already present during the Bronze Age a b Anthony 2007 p page needed a b Zimmer Carl 10 June 2015 DNA Deciphers Roots of Modern Europeans New York Times Retrieved 2020 12 12 Olsen Birgit A Olander Thomas Kristiansen Kristian 23 August 2019 Tracing the Indo Europeans Oxbow Books pp 1 6 doi 10 2307 j ctvmx3k2h 6 ISBN 978 1 78925 273 6 S2CID 202354040 Anthony 2019b p 36 Anthony 2019a p 1 19 Anthony 2017 a b c Narasimhan et al 2019 Haak et al 2015 pp 121 124 Haak et al 2015 p 5 Cassidy et al 2016 Martiniano et al 2017 Lazaridis et al 2014 Jones et al 2017 Haak et al 2015 pp 121 122 a b Pathak et al 2018 Unterlander et al 2017Genomic inference reveals that Scythians in the east and the west of the steppe zone can best be described as a mixture of Yamnaya related ancestry and an East Asian component Demographic modelling suggests independent origins for eastern and western groups with ongoing gene flow between them plausibly explaining the striking uniformity of their material culture We also find evidence that significant gene flow from east to west Eurasia must have occurred early during the Iron Age and The blend of EHG European hunter gatherer and Caucasian elements in carriers of the Yamnaya culture was formed on the European steppe and exported into Central Asia and Siberia We therefore considered an alternative model in which we treat them as a mix of Yamnaya and the Han Supplementary Table 25 This model fits all of the Iron Age Scythian groups consistent with these groups having ancestry related to East Asians not found in the other populations Alternatively the Iron Age Scythian groups can also be modelled as a mix of Yamnaya and the north Siberian Nganasan Supplementary Note 2 Supplementary Table 26 Notes EditSources EditAllentoft Morten E et al 2015 Population genomics of Bronze Age Eurasia Nature 522 7555 167 172 Bibcode 2015Natur 522 167A doi 10 1038 nature14507 PMID 26062507 S2CID 4399103 Anthony David W 2007 The Horse the Wheel and Language How Bronze Age Riders from the Eurasian Steppes Shaped the Modern World Princeton University Press ISBN 978 0 691 05887 0 Anthony David 2017 Archaeology and Language Why Archaeologists Care About the Indo European Problem In Crabtree P J Bogucki P eds European Archaeology as Anthropology Essays in Memory of Bernard Wailes Anthony David Spring Summer 2019a Archaeology Genetics and Language in the Steppes A Comment on Bomhard Journal of Indo European Studies 47 1 2 Retrieved 2020 01 09 Anthony David W 2019b Ancient DNA Mating Networks and the Anatolian Split In Serangeli Matilde Olander Thomas eds Dispersals and Diversification Linguistic and Archaeological Perspectives on the Early Stages of Indo European BRILL pp 21 54 ISBN 978 9004416192 Cassidy LM Martiniano R Murphy EM Teasdale MD Mallory J Hartwell B Bradley DG 2016 Neolithic and Bronze Age migration to Ireland and establishment of the insular Atlantic genome PNAS 113 2 368 373 Bibcode 2016PNAS 113 368C doi 10 1073 pnas 1518445113 PMC 4720318 PMID 26712024 Dolukhanov Pavel M 1996 The Early Slavs Eastern Europe from the Initial Settlement to the Kievan Rus New York Longman ISBN 0 582 23627 4 Fortson Benjamin W 2004 Indo European Language and Culture An Introduction Blackwell Publishing Gallego Llorente M Connell S Jones ER Merrett DC Jeon Y Eriksson A et al 2016 The genetics of an early Neolithic pastoralist from the Zagros Iran Scientific Reports 6 31326 Bibcode 2016NatSR 631326G doi 10 1038 srep31326 PMC 4977546 PMID 27502179 Haak W Lazaridis I Patterson N Rohland N Mallick S Llamas B et al 2015 Massive migration from the steppe was a source for Indo European 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The Origin of the Indo Iranians BRILL ISBN 978 9004160545 Lazaridis Iosif Patterson Nick Mittnik Alissa Renaud Gabriel et al 2014 Ancient human genomes suggest three ancestral populations for present day Europeans Nature 513 7518 409 413 arXiv 1312 6639 Bibcode 2014Natur 513 409L doi 10 1038 nature13673 PMC 4170574 PMID 25230663 Lazaridis I Nadel D Rollefson G Merrett DC Rohland N Mallick S et al 16 June 2016 The genetic structure of the world s first farmers bioRxiv 10 1101 059311 Supplementary Information Lazaridis I Nadel D Rollefson G Merrett DC Rohland N Mallick S et al 25 July 2016 Genomic insights into the origin of farming in the ancient Near East Nature published August 2016 536 7617 419 424 Bibcode 2016Natur 536 419L doi 10 1038 nature19310 PMC 5003663 PMID 27459054 Lazaridis Iosif Alpaslan Roodenberg Songul Acar Ayse Acikkol Aysen Agelarakis Anagnostis Aghikyan Levon Akyuz Ugur Andreeva Desislava Andrijasevic Gojko Antonovic Dragana Armit Ian Atmaca Alper Avetisyan Pavel Aytek Ahmet Ihsan Bacvarov Krum 26 August 2022 A genetic probe into the ancient and medieval history of Southern Europe and West Asia Science 377 6609 940 951 doi 10 1126 science abq0755 ISSN 0036 8075 Mallory J P 1997 Yamna Culture Encyclopedia of Indo European Culture Fitzroy Dearborn Mallory J P 1999 In Search of the Indo Europeans Language Archaeology and Myth reprint ed London Thames amp Hudson ISBN 978 0 500 27616 7 Martiniano R et al 2017 The population genomics of archaeological transition in west Iberia Investigation of ancient substructure using imputation and haplotype based methods PLOS Genet 13 7 e1006852 doi 10 1371 journal pgen 1006852 PMC 5531429 PMID 28749934 Mathieson Iain et al 10 October 2015 Eight thousand years of natural selection in Europe bioRxiv 10 1101 016477 Mathieson Iain et al 23 November 2015 Genome wide patterns of selection in 230 ancient Eurasians Nature published 24 December 2015 528 7583 499 503 Bibcode 2015Natur 528 499M doi 10 1038 nature16152 PMC 4918750 PMID 26595274 Mathieson Iain et al 21 February 2018 The Genomic History of Southeastern Europe Nature Nature Research 555 7695 197 203 Bibcode 2018Natur 555 197M doi 10 1038 nature25778 PMC 6091220 PMID 29466330 Morgunova Nina Khokhlova Olga 2013 Chronology and Periodization of the Pit Grave Culture in the Area Between the Volga and Ural Rivers Based on 14C Dating and Paleopedological Research Radiocarbon 55 2 3 1286 1296 doi 10 2458 azu js rc 55 16087 ISSN 0033 8222 Narasimhan VM Patterson N Moorjani P Rohland N et al 2019 The formation of human populations in South and Central Asia Science 365 6457 eaat7487 doi 10 1126 science aat7487 PMC 6822619 PMID 31488661 Nordgvist Kerkko Heyd Volker 2020 The Forgotten Child of the Wider Corded Ware Family Russian Fatyanovo Culture in Context Proceedings of the Prehistoric Society 86 65 93 doi 10 1017 ppr 2020 9 S2CID 228923806 Novembre John 11 June 2015 Ancient DNA steps into the language debate PDF Nature 522 7555 164 165 doi 10 1038 522164a PMID 26062506 S2CID 205085294 Parpola Asko 2015 The Roots of Hinduism Oxford University Press Pashnick Jeff August 2014 Genetic Analysis of Ancient Human Remains from the Early Bronze Age Cultures of the North PonticSteppe Region Masters Theses thesis Vol 737 Grand Valley State University Retrieved 2020 01 12 Pathak AK Kadian A Kushniarevich A Montinaro F Mondal M Ongaro L et al 6 December 2018 The Genetic Ancestry of Modern Indus Valley Populations from Northwest India The American Journal of Human Genetics 103 6 918 929 doi 10 1016 j ajhg 2018 10 022 PMC 6288199 PMID 30526867 Unterlander Martina Palstra Friso Lazaridis Iosif Pilipenko Aleksandr Hofmanova Zuzana Gross Melanie et al 2017 Ancestry and demography and descendants of Iron Age nomads of the Eurasian Steppe Nature Communications 8 14615 Bibcode 2017NatCo 814615U doi 10 1038 ncomms14615 ISSN 2041 1723 PMC 5337992 PMID 28256537 Wang Chuan Chao 4 February 2019 Ancient human genome wide data from a 3000 year interval in the Caucasus corresponds with eco geographic regions Eurasia Nature Communications Nature Research 10 1 590 Bibcode 2019NatCo 10 590W bioRxiv 10 1101 322347 doi 10 1038 s41467 018 08220 8 PMC 6360191 PMID 30713341 Wilde S Timpson A Kirsanow K Kaiser E Kayser M Unterlander M et al 2014 Direct evidence for positive selection of skin hair and eye pigmentation in Europeans during the last 5 000 y Proceedings of the National Academy of Sciences of the United States of America 111 13 4832 4837 Bibcode 2014PNAS 111 4832W doi 10 1073 pnas 1316513111 PMC 3977302 PMID 24616518 External links Edit Wikimedia Commons has media related to Yamna culture Genetic study revives debate on origin and expansion of Indo European Languages Science Daily March 2015 First Horse Warriors PBS org NOVA series 15 May 2019 Retrieved from https en wikipedia org w index php title Yamnaya culture amp oldid 1133977048, wikipedia, wiki, book, books, library,

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