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Right whale

Right whales are three species of large baleen whales of the genus Eubalaena: the North Atlantic right whale (E. glacialis), the North Pacific right whale (E. japonica) and the Southern right whale (E. australis). They are classified in the family Balaenidae with the bowhead whale. Right whales have rotund bodies with arching rostrums, V-shaped blowholes and dark gray or black skin. The most distinguishing feature of a right whale is the rough patches of skin on its head, which appear white due to parasitism by whale lice. Right whales are typically 13–17 m (43–56 ft) long and weigh up to 100 short tons (91 t; 89 long tons) or more.

Right whales[2]
Temporal range: Miocene–recent[1]
Southern right whale breaching
Size compared to an average human
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Infraorder: Cetacea
Family: Balaenidae
Genus: Eubalaena
Gray, 1864
Type species
Balaena australis [2]
Desmoulins, 1822
Species
Range map of Eubalaena species:
  E. glacialis[5]
  E. australis[3]
  E. japonica[6]
Synonyms[8]
  • Baloena Robineau, 1989
  • Halibalaena Gray, 1873
  • Hunterius Gray, 1866
  • Hunterus Gray, 1864
  • Macleayanus Marschall, 1873
  • Macleayius Gray, 1865

All three species are migratory, moving seasonally to feed or give birth. The warm equatorial waters form a barrier that isolates the northern and southern species from one another although the southern species, at least, has been known to cross the equator. In the Northern Hemisphere, right whales tend to avoid open waters and stay close to peninsulas and bays and on continental shelves, as these areas offer greater shelter and an abundance of their preferred foods. In the Southern Hemisphere, right whales feed far offshore in summer, but a large portion of the population occur in near-shore waters in winter. Right whales feed mainly on copepods but also consume krill and pteropods. They may forage the surface, underwater or even the ocean bottom. During courtship, males gather into large groups to compete for a single female, suggesting that sperm competition is an important factor in mating behavior. Gestation tends to last a year, and calves are weaned at eight months old.

Right whales were a preferred target for whalers because of their docile nature, their slow surface-skimming feeding behaviors, their tendency to stay close to the coast, and their high blubber content (which makes them float when they are killed, and which produced high yields of whale oil). Although the whales no longer face pressure from commercial whaling, humans remain by far the greatest threat to these species: the two leading causes of death are being struck by ships and entanglement in fishing gear. Today, the North Atlantic and North Pacific right whales are among the most endangered whales in the world.

Naming edit

A common explanation for the name right whales is that they were regarded as the right ones to hunt,[9] as they float when killed and often swim within sight of shore. They are quite docile and do not tend to shy away from approaching boats. As a result, they were hunted nearly to extinction during the active years of the whaling industry. However, this origin is questionable: in his history of American whaling, Eric Jay Dolin writes:

Despite this highly plausible rationale, nobody actually knows how the right whale got its name. The earliest references to the right whale offer no indication why it was called that, and some who have studied the issue point out that the word "right" in this context might just as likely be intended "to connote 'true' or 'proper,' meaning typical of the group."

— E.J. Dolin, Leviathan: The History of Whaling in America, quoting a 1766 Connecticut Courant newspaper article.[10]

For the scientific names, the generic name Eubalaena means "good or true whales", and specific names include glacialis ("ice") for North Atlantic species, australis ("southern") for Southern Hemisphere species, and japonica ("Japanese") for North Pacific species. [11]

Taxonomy edit

 
North Pacific right whale in Half Moon Bay, California, 20 March 1982, photo by Jim Scarff

The right whales were first classified in the genus Balaena in 1758 by Carl Linnaeus, who at the time considered all of the right whales (including the bowhead) as a single species. Through the 1800s and 1900s, in fact, the family Balaenidae has been the subject of great taxonometric debate. Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale, as one, two, three or four species, either in a single genus or in two separate genera. In the early whaling days, they were all thought to be a single species, Balaena mysticetus. Eventually, it was recognized that bowheads and right whales were in fact different, and John Edward Gray proposed the genus Eubalaena for the right whale in 1864. Later, morphological factors such as differences in the skull shape of northern and southern right whales indicated at least two species of right whale – one in the Northern Hemisphere, the other in the Southern Ocean.[12] As recently as 1998, Rice, in his comprehensive and otherwise authoritative classification listed just two species: Balaena glacialis (the right whales) and Balaena mysticetus (the bowheads).[13]

 
Southern right whale in the breeding grounds at Peninsula Valdés, Patagonia

In 2000, two studies of DNA samples from each of the whale populations concluded the northern and southern populations of right whale should be considered separate species. What some scientists found more surprising was the discovery that the North Pacific and North Atlantic populations are also distinct, and that the North Pacific species is more closely related to the southern right whale than to the North Atlantic right whale.[14][15] The authors of one of these studies concluded that these species have not interbred for between 3 million and 12 million years.[15]

In 2001, Brownell et al. reevaluated the conservation status of the North Pacific right whale as a distinct species,[16] and in 2002, the Scientific Committee of the International Whaling Commission (IWC) accepted Rosenbaum's findings, and recommended that the Eubalaena nomenclature be retained for this genus.[17]

A 2007 study by Churchill provided further evidence to conclude that the three different living right whale species constitute a distinct phylogenetic lineage from the bowhead, and properly belong to a separate genus.[18]

The following cladogram of the family Balaenidae serves to illustrate the current scientific consensus as to the relationships between the three right whales and the bowhead whale.

Family Balaenidae
  Balaenidae  
Eubalaena
  

 E. glacialis   (North Atlantic right whale)

  

 E. japonica   (North Pacific right whale)

 E. australis   (Southern right whale)

(right whales)
Balaena

 B. mysticetus bowhead whale

 (bowhead whales) 
The right whales, genus Eubalaena, in the family Balaenidae[14]

A cladogram is a tool for visualizing and comparing the evolutionary relationships between taxa; the point where each node branches is analogous to an evolutionary branching – the diagram can be read left-to-right, much like a timeline.

 
Orange whale lice on a right whale

Whale lice, parasitic cyamid crustaceans that live off skin debris, offer further information through their own genetics. Because these lice reproduce much more quickly than whales, their genetic diversity is greater. Marine biologists at the University of Utah examined these louse genes and determined their hosts split into three species 5–6 million years ago, and these species were all equally abundant before whaling began in the 11th century.[19] The communities first split because of the joining of North and South America. The rising temperatures of the equator then created a second split, into northern and southern groups, preventing them from interbreeding.[20] "This puts an end to the long debate about whether there are three Eubalaena species of right whale. They really are separate beyond a doubt", Jon Seger, the project's leader, told BBC News.[21]

Others edit

The pygmy right whale (Caperea marginata), a much smaller whale of the Southern Hemisphere, was until recently considered a member of the Family Balaenidae. However, they are not right whales at all, and their taxonomy is presently in doubt. Most recent authors place this species into the monotypic Family Neobalaenidae,[22] but a 2012 study suggests that it is instead the last living member of the Family Cetotheriidae, a family previously considered extinct.[23]

Yet another species of right whale was proposed by Emanuel Swedenborg in the 18th century—the so-called Swedenborg whale. The description of this species was based on a collection of fossil bones unearthed at Norra Vånga, Sweden, in 1705 and believed to be those of giants. The bones were examined by Swedenborg, who realized they belong to a species of whale. The existence of this species has been debated, and further evidence for this species was discovered during the construction of a motorway in Strömstad, Sweden in 2009.[24] To date, however, scientific consensus still considers Hunterius swedenborgii to be a North Atlantic right whale.[25] According to a DNA analysis conducted, it was later confirmed that the fossil bones are actually from a bowhead whale.[26]

Characteristics edit

 
Right whale eye
 
An example of baleen plates; there are about 50 plates in this photo.

Adult right whales are typically 13–16 m (43–52 ft) long. They have extremely thick bodies with a girth as much as 60% of total body length in some cases. They have large, broad and blunt pectoral flippers and the deeply notched, smoothly tipped tail flukes make up to 40% of their body length. The North Pacific species is on average the largest of the three species. weigh 100 short tons (91 t; 89 long tons). The upper jaw of a right whale is a bit arched, and the lower lip is strongly curved. On each side of the upper jaw are 200–270 baleen plates. These are narrow and approximately 2–2.8 m (6.6–9.2 ft) long, and are covered in very thin hairs.[27] Right whales have a distinctive wide V-shaped blow, caused by the widely spaced blowholes on the top of the head. The blow rises 5 m (16 ft) above the surface.[28]

The skin is generally black with occasional white blotches on the body, while some individuals have mottled patterns.[27] Unlike other whales, a right whale has distinctive callosities (roughened patches of skin) on its head. The callosities appear white due to large colonies of cyamids (whale lice).[12][29] Each individual has a unique callosities pattern. In 2016, a competitive effort resulted in the use of facial recognition software to derive a process to uniquely identify right whales with about 87% accuracy based on their callosities.[30] The primary role of callosities has been considered to be protection against predators. Right whale declines might have also reduced barnacles.[31]

An unusually large 40% of their body weight is blubber, which is of relatively low density. Consequently, unlike many other species of whale, dead right whales tend to float.[32][33] Many southern right whales are seen with rolls of fats behind blowholes that northern species often lack, and these are regarded as a sign of better health condition due to sufficient nutrition supply, and could have contributed in vast differences in recovery status between right whales in the southern and northern hemisphere, other than direct impacts by humankind.[34]

The penis on a right whale can be up to 2.7 m (8.9 ft) – the testes, at up to 2 m (6.6 ft) in length, 78 cm (2.56 ft) in diameter, and weighing up to 525 kg (1157 lbs), are also by far the largest of any animal on Earth.[35] The blue whale may be the largest animal on the planet, yet the testicles of the right whale are ten times the size of those of the blue whale. They also exceed predictions in terms of relative size, being six times larger than would be expected on the basis of body mass. Together, the testicles make up nearly 1% of the right whale's total body weight. This strongly suggests sperm competition is important in mating, which correlates to the fact that right whales are highly promiscuous.[28][36]

Range and habitat edit

 
The distinctive V-shaped blow of a right whale

The three Eubalaena species inhabit three distinct areas of the globe: the North Atlantic in the western Atlantic Ocean, the North Pacific in a band from Japan to Alaska and all areas of the Southern Ocean. The whales can only cope with the moderate temperatures found between 20 and 60 degrees in latitude. The warm equatorial waters form a barrier that prevents mixing between the northern and southern groups with minor exclusions.[37] Although the southern species in particular must travel across open ocean to reach its feeding grounds, the species is not considered to be pelagic. In general, they prefer to stay close to peninsulas and bays and on continental shelves, as these areas offer greater shelter and an abundance of their preferred foods.[20]

Because the oceans are so large, it is very difficult to accurately gauge whale population sizes. Approximate figures:[18]

North Atlantic right whale edit

Almost all of the 400 North Atlantic right whales live in the western North Atlantic Ocean. In northern spring, summer and autumn, they feed in areas off the Canadian and northeast U.S. coasts in a range stretching from New York to Newfoundland. Particularly popular feeding areas are the Bay of Fundy and Cape Cod Bay. In winter, they head south towards Georgia and Florida to give birth.[38] There have been a smattering of sightings further east over the past few decades; several sightings were made close to Iceland in 2003. These are possibly the remains of a virtually extinct eastern Atlantic stock, but examination of old whalers' records suggests they are more likely to be strays.[18] However, a few sightings have happened between Norway, Ireland, Spain, Portugal, the Canary Islands and Italy;[39][40] at least the Norway individuals come from the Western stock.[41]

North Pacific right whale edit

The North Pacific right whale appears to occur in two populations. The population in the eastern North Pacific/Bering Sea is extremely low, numbering about 30 individuals.[42] A larger western population of 100–200 appears to be surviving in the Sea of Okhotsk, but very little is known about this population. Thus, the two northern right whale species are the most endangered of all large whales and two of the most endangered animal species in the world. Based on current population density trends, both species are predicted to become extinct within 200 years.[43] The Pacific species was historically found in summer from the Sea of Okhotsk in the west to the Gulf of Alaska in the east, generally north of 50°N. Today, sightings are very rare and generally occur in the mouth of the Sea of Okhotsk and in the eastern Bering Sea. Although this species is very likely to be migratory like the other two species, its movement patterns are not known.[44]

Southern right whale edit

The last major population review of southern right whales by the International Whaling Commission was in 1998. Researchers used data about adult female populations from three surveys (one in each of Argentina, South Africa and Australia) and extrapolated to include unsurveyed areas and estimated counts of males and calves (using available male:female and adult:calf ratios), giving an estimated 1997 population of 7,500 animals. More recent data from 2007 indicate those survey areas have shown evidence of strong recovery, with a population approaching twice that of a decade earlier. However, other breeding populations are still very small, and data are insufficient to determine whether they, too, are recovering.[3]

The southern right whale spends the summer months in the far Southern Ocean feeding, probably close to Antarctica. It migrates north in winter for breeding, and can be seen around the coasts of Argentina, Australia, Brazil, Chile, Mozambique, New Zealand, South Africa and Uruguay.[45] The South American, South African and Australasian groups apparently intermix very little, if at all, because of the strong fidelity of mothers to their feeding and calving grounds. The mother passes these instincts to her calves.[18]

Life history edit

Right whales swim slowly, reaching only 5 kn (9.3 km/h) at top speed. However, they are highly acrobatic and frequently breach (jump clear of the sea surface), tail-slap and lobtail.[27]

Diet and predation edit

The right whales' diets consist primarily of zooplankton, primarily the tiny crustaceans called copepods, as well as krill, and pteropods, although they are occasionally opportunistic feeders. As with other baleens, they feed by filtering prey from the water. They swim with an open mouth, filling it with water and prey. The whale then expels the water, using its baleen plates to retain the prey. Prey must occur in sufficient numbers to trigger the whale's interest, be large enough that the baleen plates can filter it, and be slow enough that it cannot escape. The "skimming" may take place on the surface, underwater, or even at the seabed, indicated by mud occasionally observed on right whales' bodies.[18]

The right whales' two known predators are humans and orcas. When danger lurks, a group of right whales may cluster into a circle, and thrash their outwards-pointing tails. They may also head for shallow water, which sometimes proves to be an ineffective defense. Aside from the strong tails and massive heads equipped with callosities,[31] the sheer size of this animal is its best defense, making young calves the most vulnerable to orca and shark attacks.[28]

Vocalization and hearing edit

Vocalizations made by right whales are not elaborate compared to those made by other whale species. The whales make groans, pops and belches typically at frequencies around 500 Hz. The purpose of the sounds is not known but may be a form of communication between whales within the same group. Northern right whales responded to sounds similar to police sirens—sounds of much higher frequency than their own. On hearing the sounds, they moved rapidly to the surface. The research was of particular interest because northern rights ignore most sounds, including those of approaching boats. Researchers speculate this information may be useful in attempts to reduce the number of ship-whale collisions or to encourage the whales to surface for ease of harvesting.[43][46]

Courtship and reproduction edit

 
A female North Atlantic right whale with her calf.

During the mating season, which can occur at any time in the North Atlantic, right whales gather into "surface-active groups" made up of as many as 20 males consorting a single female. The female has her belly to the surface while the males stroke her with their flippers or keep her underwater. The males do not compete as aggressively against each other as male humpbacks. The female may not become pregnant but she is still able to assess the condition of potential mates.[18] The mean age of first parturition in North Atlantic right whales is estimated at between 7.5[47] and 9[48] years. Females breed every 3–5 years;[47][49] the most commonly seen calving intervals are 3 years and may vary from 2 up to 21 years due to multiple factors.[50][51]

Both reproduction and calving take place during the winter months.[52] Calves are approximately 1 short ton (0.91 t; 0.89 long tons) in weight and 4–6 m (13–20 ft) in length at birth following a gestation period of 1 year. The right whale grows rapidly in its first year, typically doubling in length. Weaning occurs after eight months to one year and the growth rate in later years is not well understood—it may be highly dependent on whether a calf stays with its mother for a second year.[18]

Respective congregation areas in the same region may function as for different objectives for whales.[53]

Lifespan edit

Very little is known about the life span of right whales. One of the few well-documented cases is of a female North Atlantic right whale that was photographed with a baby in 1935, then photographed again in 1959, 1980, 1985, and 1992. Consistent callosity patterns ensured it was the same animal. She was last photographed in 1995 with a seemingly fatal head wound, presumably from a ship strike. By conservative estimates (e.g. she was a new mother who had just reached sexual maturity in 1935), she was nearly 70 years to more than 100 years of age, if not older.[54] Research on the closely related bowhead whale exceeding 210 years suggests this lifespan is not uncommon and may even be exceeded.[18][55]

Relationship to humans edit

Whaling edit

 
Whaling in small wooden boats with hand harpoons was a hazardous enterprise, even when hunting the "right" whale.

In the early centuries of shore-based whaling before 1712, right whales were virtually the only catchable large whales, for three reasons:

  • They often swam close to shore where they could be spotted by beach lookouts, and hunted from beach-based whaleboats.
  • They are relatively slow swimmers, allowing whalers to catch up to them in their whaleboats.
  • Once killed by harpoons, they were more likely to float, and thus could be retrieved. However, some did sink when killed (10–30% in the North Pacific) and were lost unless they later stranded or surfaced.[56]

Basque people were the first to hunt right whales commercially, beginning as early as the 11th century in the Bay of Biscay. They initially sought oil, but as meat preservation technology improved, the animal was also used for food. Basque whalers reached eastern Canada by 1530[18] and the shores of Todos os Santos Bay (in Bahia, Brazil) by 1602. The last Basque voyages were made before the Seven Years' War (1756–1763). All attempts to revive the trade after the war failed. Basque shore whaling continued sporadically into the 19th century.

"Yankee whalers" from the new American colonies replaced the Basques. Setting out from Nantucket, Massachusetts, and Long Island, New York, they took up to a hundred animals in good years. By 1750, the commercial hunt of the North Atlantic right whale was essentially over. The Yankee whalers moved into the South Atlantic before the end of the 18th century. The southernmost Brazilian whaling station was established in 1796, in Imbituba. Over the next hundred years, Yankee whaling spread into the Southern and Pacific Oceans, where the Americans were joined by fleets from several European nations. The beginning of the 20th century saw much greater industrialization of whaling, and the harvest grew rapidly. According to whalers' records, by 1937 there had been 38,000 takes in the South Atlantic, 39,000 in the South Pacific, 1,300 in the Indian Ocean, and 15,000 in the North Pacific. The incompleteness of these records means the actual take was somewhat higher.[57]

As it became clear the stocks were nearly depleted, the world banned right whaling in 1937. The ban was largely successful, although violations continued for several decades. Madeira took its last two right whales in 1968. Japan took twenty-three Pacific right whales in the 1940s and more under scientific permit in the 1960s. Illegal whaling continued off the coast of Brazil for many years, and the Imbituba land station processed right whales until 1973. The Soviet Union illegally took at least 3,212 southern right whales during the 1950s and '60s, although it reported taking only four.[58]

 
A southern right whale approaches close to whale watchers near Península Valdés in Patagonia

Whale watching edit

The southern right whale has made Hermanus, South Africa, one of the world centers for whale watching. During the winter months (July–October), southern right whales come so close to the shoreline, visitors can watch whales from strategically placed hotels.[59] The town employs a "whale crier" (cf. town crier) to walk through the town announcing where whales have been seen.[60] In Brazil, Imbituba in Santa Catarina has been recognized as the National Right Whale Capital and holds annual Right Whale Week celebrations in September[61] when mothers and calves are more often seen. The old whaling station there has been converted to a museum dedicated to the whales.[62] In winter in Argentina, Península Valdés in Patagonia hosts the largest breeding population of the species, with more than 2,000 animals catalogued by the Whale Conservation Institute and Ocean Alliance.[63]

Conservation edit

 
North Atlantic right whale on a Faroese stamp

Both the North Atlantic and North Pacific species are listed as a "species threatened with extinction which [is] or may be affected by trade" (Appendix I) by CITES, and as "endangered" by the IUCN Red List. In the United States, the National Marine Fisheries Service (NMFS), a subagency of the National Oceanic and Atmospheric Administration (NOAA) has classified all three species as "endangered" under the Endangered Species Act. Under the Marine Mammal Protection Act, they are listed as "depleted".[64][65][66]

The southern right whale is listed as "endangered" under the Australian Environment Protection and Biodiversity Conservation Act, as "nationally endangered" under the New Zealand Threat Classification System, as a "natural monument" by the Argentine National Congress, and as a "State Natural Monument" under the Brazilian National Endangered Species List.[66]

The US and Brazil added new protections for right whales in the 2000s to address the two primary hazards. While environmental campaigners were, as reported in 2001, pleased about the plan's positive effects, they attempted to force the US government to do more.[67] In particular, they advocated 12 knots (22 km/h) speed limits for ships within 40 km (25 mi) of US ports in times of high right whale presence. Citing concerns about excessive trade disruption, it did not institute greater protections. The Defenders of Wildlife, the Humane Society of the United States and the Ocean Conservancy sued the NMFS in September 2005 for "failing to protect the critically endangered North Atlantic Right Whale, which the agency acknowledges is 'the rarest of all large whale species' and which federal agencies are required to protect by both the Marine Mammal Protection Act and the Endangered Species Act", demanding emergency protection measures.[68]

The southern right whale, listed as "endangered" by CITES and "lower risk - conservation dependent" by the IUCN, is protected in the jurisdictional waters of all countries with known breeding populations (Argentina, Australia, Brazil, Chile, New Zealand, South Africa and Uruguay). In Brazil, a federal Environmental Protection Area encompassing some 1,560 km2 (600 sq mi) and 130 km (81 mi) of coastline in Santa Catarina State was established in 2000 to protect the species' main breeding grounds in Brazil and promote whale watching.[69]

On February 6, 2006, NOAA proposed its Strategy to Reduce Ship Strikes to North Atlantic Right Whales.[70] The proposal, opposed by some shipping interests, limited ship speeds during calving season. The proposal was made official when on December 8, 2008, NOAA issued a press release that included the following:[71]

  • Effective January 2009, ships 65 feet (20 m) or longer are limited to 10 knots (19 km/h) in waters off New England when whales begin gathering in this area as part of their annual migration. The restriction extends to 20 nautical miles (37 km) around major mid-Atlantic ports.
  • The speed restriction applies in waters off New England and the southeastern US, where whales gather seasonally:
    • Southeastern US from St. Augustine, Florida to Brunswick, Georgia from Nov 15 to April 15
    • Mid-Atlantic U.S. areas from Rhode Island to Georgia from Nov 1 to April 30.
    • Cape Cod Bay from Jan 1 to May 15
    • Off Race Point at the northern end of Cape Cod from March 1 to April 30
    • Great South Channel of New England from April 1 to July 31
  • Temporary voluntary speed limits in other areas or times when a group of three or more right whales is confirmed
  • Scientists would assess the rule's effectiveness before the rule expires in 2013.

In 2020, NOAA published its assessment and found that since the speed rule was adopted, the total number of documented deaths from vessel strike decreased but serious and non-serious injuries have increased.[72] A report by the organization Oceana found that between 2017 and 2020, disobedience of the rule reached close to 90% in mandatory speed zones while in voluntary areas, disobedience neared 85%.[73]

Threats edit

 
The remains of a North Atlantic right whale after it collided with a ship propeller

The leading cause of death among the North Atlantic right whale, which migrates through some of the world's busiest shipping lanes while journeying off the east coast of the United States and Canada, is being struck by ships.[note 1][74] At least sixteen ship-strike deaths were reported between 1970 and 1999, and probably more remain unreported.[18] According to NOAA, twenty-five of the seventy-one right whale deaths reported since 1970 resulted from ship strikes.[71]

A second major cause of morbidity and mortality in the North Atlantic right whale is entanglement in plastic fishing gear. Right whales ingest plankton with wide-open mouths, risking entanglement in any rope or net fixed in the water column. Rope wraps around their upper jaws, flippers and tails. Some are able to escape, but others remain tangled.[75] Whales can be successfully disentangled, if observed and aided. In July 1997, the U.S. NOAA introduced the Atlantic Large Whale Take Reduction Plan, which seeks to minimize whale entanglement in fishing gear and record large whale sightings in an attempt to estimate numbers and distribution.[76]

In 2012, the U.S. Navy proposed to create a new undersea naval training range immediately adjacent to northern right whale calving grounds in shallow waters off the Florida/Georgia border. Legal challenges by leading environmental groups including the Natural Resources Defense Council were denied in federal court, allowing the Navy to proceed.[77][78] These rulings were made despite the extremely low numbers (as low as 313 by some estimates) of right whales in existence at this time, and a very poor calving season.[79]

Notes edit

  1. ^ The Lloyd's mirror effect results in low frequency propeller sounds not being discernible near the surface, where most accidents occur. Combined with spreading and acoustic shadowing effects, the result is that the animal is unable to hear an approaching vessel before it has been run over or entrapped by the hydrodynamic forces of the vessel's passage.

References edit

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Further reading edit

  • Kraus, Scott D.; Rolland, Rosalind, eds. (2010). The Urban Whale: North Atlantic Right Whales at the Crossroads. Harvard University Press. ISBN 978-0-674-03475-4.

External links edit

  • IUCN Red List entry
  • Right Whale Lesson Plan from Smithsonian Education
  • North Atlantic Right Whale on the Smithsonian Ocean Portal
  • National Library for the Environment
  • The Fall and Rise of the Right Whale, New York Times, March 16, 2009

right, whale, this, article, about, three, species, whale, genus, eubalaena, close, relative, that, recently, excluded, from, right, whales, bowhead, whale, balaena, similarly, named, dolphin, dolphin, other, uses, disambiguation, three, species, large, baleen. This article is about three species of whale in the genus Eubalaena For a close relative that was recently excluded from the right whales see bowhead whale Balaena For the similarly named dolphin see Right whale dolphin For other uses see Right whale disambiguation Right whales are three species of large baleen whales of the genus Eubalaena the North Atlantic right whale E glacialis the North Pacific right whale E japonica and the Southern right whale E australis They are classified in the family Balaenidae with the bowhead whale Right whales have rotund bodies with arching rostrums V shaped blowholes and dark gray or black skin The most distinguishing feature of a right whale is the rough patches of skin on its head which appear white due to parasitism by whale lice Right whales are typically 13 17 m 43 56 ft long and weigh up to 100 short tons 91 t 89 long tons or more Right whales 2 Temporal range Miocene recent 1 PreꞒ Ꞓ O S D C P T J K Pg NSouthern right whale breachingSize compared to an average humanScientific classificationDomain EukaryotaKingdom AnimaliaPhylum ChordataClass MammaliaOrder ArtiodactylaInfraorder CetaceaFamily BalaenidaeGenus EubalaenaGray 1864Type speciesBalaena australis 2 Desmoulins 1822SpeciesE australis Southern Desmoulins 1822 3 E ianitrix Bisconti Lambert and Bosselaers 2017 4 E glacialis North Atlantic Muller 1776 5 E japonica North Pacific Lacepede 1818 6 E shinshuensisKimura amp Narita 2007 7 Range map of Eubalaena species E glacialis 5 E australis 3 E japonica 6 Synonyms 8 Baloena Robineau 1989 Halibalaena Gray 1873 Hunterius Gray 1866 Hunterus Gray 1864 Macleayanus Marschall 1873 Macleayius Gray 1865All three species are migratory moving seasonally to feed or give birth The warm equatorial waters form a barrier that isolates the northern and southern species from one another although the southern species at least has been known to cross the equator In the Northern Hemisphere right whales tend to avoid open waters and stay close to peninsulas and bays and on continental shelves as these areas offer greater shelter and an abundance of their preferred foods In the Southern Hemisphere right whales feed far offshore in summer but a large portion of the population occur in near shore waters in winter Right whales feed mainly on copepods but also consume krill and pteropods They may forage the surface underwater or even the ocean bottom During courtship males gather into large groups to compete for a single female suggesting that sperm competition is an important factor in mating behavior Gestation tends to last a year and calves are weaned at eight months old Right whales were a preferred target for whalers because of their docile nature their slow surface skimming feeding behaviors their tendency to stay close to the coast and their high blubber content which makes them float when they are killed and which produced high yields of whale oil Although the whales no longer face pressure from commercial whaling humans remain by far the greatest threat to these species the two leading causes of death are being struck by ships and entanglement in fishing gear Today the North Atlantic and North Pacific right whales are among the most endangered whales in the world Contents 1 Naming 2 Taxonomy 2 1 Others 3 Characteristics 4 Range and habitat 4 1 North Atlantic right whale 4 2 North Pacific right whale 4 3 Southern right whale 5 Life history 5 1 Diet and predation 5 2 Vocalization and hearing 5 3 Courtship and reproduction 5 4 Lifespan 6 Relationship to humans 6 1 Whaling 6 2 Whale watching 7 Conservation 7 1 Threats 8 Notes 9 References 10 Further reading 11 External linksNaming editA common explanation for the name right whales is that they were regarded as the right ones to hunt 9 as they float when killed and often swim within sight of shore They are quite docile and do not tend to shy away from approaching boats As a result they were hunted nearly to extinction during the active years of the whaling industry However this origin is questionable in his history of American whaling Eric Jay Dolin writes Despite this highly plausible rationale nobody actually knows how the right whale got its name The earliest references to the right whale offer no indication why it was called that and some who have studied the issue point out that the word right in this context might just as likely be intended to connote true or proper meaning typical of the group E J Dolin Leviathan The History of Whaling in America quoting a 1766 Connecticut Courant newspaper article 10 For the scientific names the generic name Eubalaena means good or true whales and specific names include glacialis ice for North Atlantic species australis southern for Southern Hemisphere species and japonica Japanese for North Pacific species 11 Taxonomy edit nbsp North Pacific right whale in Half Moon Bay California 20 March 1982 photo by Jim ScarffThe right whales were first classified in the genus Balaena in 1758 by Carl Linnaeus who at the time considered all of the right whales including the bowhead as a single species Through the 1800s and 1900s in fact the family Balaenidae has been the subject of great taxonometric debate Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale as one two three or four species either in a single genus or in two separate genera In the early whaling days they were all thought to be a single species Balaena mysticetus Eventually it was recognized that bowheads and right whales were in fact different and John Edward Gray proposed the genus Eubalaena for the right whale in 1864 Later morphological factors such as differences in the skull shape of northern and southern right whales indicated at least two species of right whale one in the Northern Hemisphere the other in the Southern Ocean 12 As recently as 1998 Rice in his comprehensive and otherwise authoritative classification listed just two species Balaena glacialis the right whales and Balaena mysticetus the bowheads 13 nbsp Southern right whale in the breeding grounds at Peninsula Valdes PatagoniaIn 2000 two studies of DNA samples from each of the whale populations concluded the northern and southern populations of right whale should be considered separate species What some scientists found more surprising was the discovery that the North Pacific and North Atlantic populations are also distinct and that the North Pacific species is more closely related to the southern right whale than to the North Atlantic right whale 14 15 The authors of one of these studies concluded that these species have not interbred for between 3 million and 12 million years 15 In 2001 Brownell et al reevaluated the conservation status of the North Pacific right whale as a distinct species 16 and in 2002 the Scientific Committee of the International Whaling Commission IWC accepted Rosenbaum s findings and recommended that the Eubalaena nomenclature be retained for this genus 17 A 2007 study by Churchill provided further evidence to conclude that the three different living right whale species constitute a distinct phylogenetic lineage from the bowhead and properly belong to a separate genus 18 The following cladogram of the family Balaenidae serves to illustrate the current scientific consensus as to the relationships between the three right whales and the bowhead whale Family Balaenidae Balaenidae Eubalaena E glacialis North Atlantic right whale E japonica North Pacific right whale E australis Southern right whale right whales Balaena B mysticetus bowhead whale bowhead whales The right whales genus Eubalaena in the family Balaenidae 14 A cladogram is a tool for visualizing and comparing the evolutionary relationships between taxa the point where each node branches is analogous to an evolutionary branching the diagram can be read left to right much like a timeline nbsp Orange whale lice on a right whaleWhale lice parasitic cyamid crustaceans that live off skin debris offer further information through their own genetics Because these lice reproduce much more quickly than whales their genetic diversity is greater Marine biologists at the University of Utah examined these louse genes and determined their hosts split into three species 5 6 million years ago and these species were all equally abundant before whaling began in the 11th century 19 The communities first split because of the joining of North and South America The rising temperatures of the equator then created a second split into northern and southern groups preventing them from interbreeding 20 This puts an end to the long debate about whether there are three Eubalaena species of right whale They really are separate beyond a doubt Jon Seger the project s leader told BBC News 21 Others edit The pygmy right whale Caperea marginata a much smaller whale of the Southern Hemisphere was until recently considered a member of the Family Balaenidae However they are not right whales at all and their taxonomy is presently in doubt Most recent authors place this species into the monotypic Family Neobalaenidae 22 but a 2012 study suggests that it is instead the last living member of the Family Cetotheriidae a family previously considered extinct 23 Yet another species of right whale was proposed by Emanuel Swedenborg in the 18th century the so called Swedenborg whale The description of this species was based on a collection of fossil bones unearthed at Norra Vanga Sweden in 1705 and believed to be those of giants The bones were examined by Swedenborg who realized they belong to a species of whale The existence of this species has been debated and further evidence for this species was discovered during the construction of a motorway in Stromstad Sweden in 2009 24 To date however scientific consensus still considers Hunterius swedenborgii to be a North Atlantic right whale 25 According to a DNA analysis conducted it was later confirmed that the fossil bones are actually from a bowhead whale 26 Characteristics editSee also Whale anatomy nbsp Right whale eye nbsp An example of baleen plates there are about 50 plates in this photo Adult right whales are typically 13 16 m 43 52 ft long They have extremely thick bodies with a girth as much as 60 of total body length in some cases They have large broad and blunt pectoral flippers and the deeply notched smoothly tipped tail flukes make up to 40 of their body length The North Pacific species is on average the largest of the three species weigh 100 short tons 91 t 89 long tons The upper jaw of a right whale is a bit arched and the lower lip is strongly curved On each side of the upper jaw are 200 270 baleen plates These are narrow and approximately 2 2 8 m 6 6 9 2 ft long and are covered in very thin hairs 27 Right whales have a distinctive wide V shaped blow caused by the widely spaced blowholes on the top of the head The blow rises 5 m 16 ft above the surface 28 The skin is generally black with occasional white blotches on the body while some individuals have mottled patterns 27 Unlike other whales a right whale has distinctive callosities roughened patches of skin on its head The callosities appear white due to large colonies of cyamids whale lice 12 29 Each individual has a unique callosities pattern In 2016 a competitive effort resulted in the use of facial recognition software to derive a process to uniquely identify right whales with about 87 accuracy based on their callosities 30 The primary role of callosities has been considered to be protection against predators Right whale declines might have also reduced barnacles 31 An unusually large 40 of their body weight is blubber which is of relatively low density Consequently unlike many other species of whale dead right whales tend to float 32 33 Many southern right whales are seen with rolls of fats behind blowholes that northern species often lack and these are regarded as a sign of better health condition due to sufficient nutrition supply and could have contributed in vast differences in recovery status between right whales in the southern and northern hemisphere other than direct impacts by humankind 34 The penis on a right whale can be up to 2 7 m 8 9 ft the testes at up to 2 m 6 6 ft in length 78 cm 2 56 ft in diameter and weighing up to 525 kg 1157 lbs are also by far the largest of any animal on Earth 35 The blue whale may be the largest animal on the planet yet the testicles of the right whale are ten times the size of those of the blue whale They also exceed predictions in terms of relative size being six times larger than would be expected on the basis of body mass Together the testicles make up nearly 1 of the right whale s total body weight This strongly suggests sperm competition is important in mating which correlates to the fact that right whales are highly promiscuous 28 36 Range and habitat edit nbsp The distinctive V shaped blow of a right whaleThe three Eubalaena species inhabit three distinct areas of the globe the North Atlantic in the western Atlantic Ocean the North Pacific in a band from Japan to Alaska and all areas of the Southern Ocean The whales can only cope with the moderate temperatures found between 20 and 60 degrees in latitude The warm equatorial waters form a barrier that prevents mixing between the northern and southern groups with minor exclusions 37 Although the southern species in particular must travel across open ocean to reach its feeding grounds the species is not considered to be pelagic In general they prefer to stay close to peninsulas and bays and on continental shelves as these areas offer greater shelter and an abundance of their preferred foods 20 Because the oceans are so large it is very difficult to accurately gauge whale population sizes Approximate figures 18 400 North Atlantic right whales Eubalaena glacialis live in the North Atlantic 23 North Pacific right whales have been identified in the eastern North Pacific Eubalaena japonica and 15 000 southern right whales Eubalaena australis are spread throughout the southern part of the Southern Hemisphere North Atlantic right whale edit Almost all of the 400 North Atlantic right whales live in the western North Atlantic Ocean In northern spring summer and autumn they feed in areas off the Canadian and northeast U S coasts in a range stretching from New York to Newfoundland Particularly popular feeding areas are the Bay of Fundy and Cape Cod Bay In winter they head south towards Georgia and Florida to give birth 38 There have been a smattering of sightings further east over the past few decades several sightings were made close to Iceland in 2003 These are possibly the remains of a virtually extinct eastern Atlantic stock but examination of old whalers records suggests they are more likely to be strays 18 However a few sightings have happened between Norway Ireland Spain Portugal the Canary Islands and Italy 39 40 at least the Norway individuals come from the Western stock 41 North Pacific right whale edit The North Pacific right whale appears to occur in two populations The population in the eastern North Pacific Bering Sea is extremely low numbering about 30 individuals 42 A larger western population of 100 200 appears to be surviving in the Sea of Okhotsk but very little is known about this population Thus the two northern right whale species are the most endangered of all large whales and two of the most endangered animal species in the world Based on current population density trends both species are predicted to become extinct within 200 years 43 The Pacific species was historically found in summer from the Sea of Okhotsk in the west to the Gulf of Alaska in the east generally north of 50 N Today sightings are very rare and generally occur in the mouth of the Sea of Okhotsk and in the eastern Bering Sea Although this species is very likely to be migratory like the other two species its movement patterns are not known 44 Southern right whale edit The last major population review of southern right whales by the International Whaling Commission was in 1998 Researchers used data about adult female populations from three surveys one in each of Argentina South Africa and Australia and extrapolated to include unsurveyed areas and estimated counts of males and calves using available male female and adult calf ratios giving an estimated 1997 population of 7 500 animals More recent data from 2007 indicate those survey areas have shown evidence of strong recovery with a population approaching twice that of a decade earlier However other breeding populations are still very small and data are insufficient to determine whether they too are recovering 3 The southern right whale spends the summer months in the far Southern Ocean feeding probably close to Antarctica It migrates north in winter for breeding and can be seen around the coasts of Argentina Australia Brazil Chile Mozambique New Zealand South Africa and Uruguay 45 The South American South African and Australasian groups apparently intermix very little if at all because of the strong fidelity of mothers to their feeding and calving grounds The mother passes these instincts to her calves 18 Life history editRight whales swim slowly reaching only 5 kn 9 3 km h at top speed However they are highly acrobatic and frequently breach jump clear of the sea surface tail slap and lobtail 27 Diet and predation edit The right whales diets consist primarily of zooplankton primarily the tiny crustaceans called copepods as well as krill and pteropods although they are occasionally opportunistic feeders As with other baleens they feed by filtering prey from the water They swim with an open mouth filling it with water and prey The whale then expels the water using its baleen plates to retain the prey Prey must occur in sufficient numbers to trigger the whale s interest be large enough that the baleen plates can filter it and be slow enough that it cannot escape The skimming may take place on the surface underwater or even at the seabed indicated by mud occasionally observed on right whales bodies 18 The right whales two known predators are humans and orcas When danger lurks a group of right whales may cluster into a circle and thrash their outwards pointing tails They may also head for shallow water which sometimes proves to be an ineffective defense Aside from the strong tails and massive heads equipped with callosities 31 the sheer size of this animal is its best defense making young calves the most vulnerable to orca and shark attacks 28 Vocalization and hearing edit See also Whale sound Vocalizations made by right whales are not elaborate compared to those made by other whale species The whales make groans pops and belches typically at frequencies around 500 Hz The purpose of the sounds is not known but may be a form of communication between whales within the same group Northern right whales responded to sounds similar to police sirens sounds of much higher frequency than their own On hearing the sounds they moved rapidly to the surface The research was of particular interest because northern rights ignore most sounds including those of approaching boats Researchers speculate this information may be useful in attempts to reduce the number of ship whale collisions or to encourage the whales to surface for ease of harvesting 43 46 Courtship and reproduction edit See also Whale reproduction nbsp A female North Atlantic right whale with her calf During the mating season which can occur at any time in the North Atlantic right whales gather into surface active groups made up of as many as 20 males consorting a single female The female has her belly to the surface while the males stroke her with their flippers or keep her underwater The males do not compete as aggressively against each other as male humpbacks The female may not become pregnant but she is still able to assess the condition of potential mates 18 The mean age of first parturition in North Atlantic right whales is estimated at between 7 5 47 and 9 48 years Females breed every 3 5 years 47 49 the most commonly seen calving intervals are 3 years and may vary from 2 up to 21 years due to multiple factors 50 51 Both reproduction and calving take place during the winter months 52 Calves are approximately 1 short ton 0 91 t 0 89 long tons in weight and 4 6 m 13 20 ft in length at birth following a gestation period of 1 year The right whale grows rapidly in its first year typically doubling in length Weaning occurs after eight months to one year and the growth rate in later years is not well understood it may be highly dependent on whether a calf stays with its mother for a second year 18 Respective congregation areas in the same region may function as for different objectives for whales 53 Lifespan edit Very little is known about the life span of right whales One of the few well documented cases is of a female North Atlantic right whale that was photographed with a baby in 1935 then photographed again in 1959 1980 1985 and 1992 Consistent callosity patterns ensured it was the same animal She was last photographed in 1995 with a seemingly fatal head wound presumably from a ship strike By conservative estimates e g she was a new mother who had just reached sexual maturity in 1935 she was nearly 70 years to more than 100 years of age if not older 54 Research on the closely related bowhead whale exceeding 210 years suggests this lifespan is not uncommon and may even be exceeded 18 55 Relationship to humans editWhaling edit Main article History of whaling nbsp Whaling in small wooden boats with hand harpoons was a hazardous enterprise even when hunting the right whale In the early centuries of shore based whaling before 1712 right whales were virtually the only catchable large whales for three reasons They often swam close to shore where they could be spotted by beach lookouts and hunted from beach based whaleboats They are relatively slow swimmers allowing whalers to catch up to them in their whaleboats Once killed by harpoons they were more likely to float and thus could be retrieved However some did sink when killed 10 30 in the North Pacific and were lost unless they later stranded or surfaced 56 Basque people were the first to hunt right whales commercially beginning as early as the 11th century in the Bay of Biscay They initially sought oil but as meat preservation technology improved the animal was also used for food Basque whalers reached eastern Canada by 1530 18 and the shores of Todos os Santos Bay in Bahia Brazil by 1602 The last Basque voyages were made before the Seven Years War 1756 1763 All attempts to revive the trade after the war failed Basque shore whaling continued sporadically into the 19th century Yankee whalers from the new American colonies replaced the Basques Setting out from Nantucket Massachusetts and Long Island New York they took up to a hundred animals in good years By 1750 the commercial hunt of the North Atlantic right whale was essentially over The Yankee whalers moved into the South Atlantic before the end of the 18th century The southernmost Brazilian whaling station was established in 1796 in Imbituba Over the next hundred years Yankee whaling spread into the Southern and Pacific Oceans where the Americans were joined by fleets from several European nations The beginning of the 20th century saw much greater industrialization of whaling and the harvest grew rapidly According to whalers records by 1937 there had been 38 000 takes in the South Atlantic 39 000 in the South Pacific 1 300 in the Indian Ocean and 15 000 in the North Pacific The incompleteness of these records means the actual take was somewhat higher 57 As it became clear the stocks were nearly depleted the world banned right whaling in 1937 The ban was largely successful although violations continued for several decades Madeira took its last two right whales in 1968 Japan took twenty three Pacific right whales in the 1940s and more under scientific permit in the 1960s Illegal whaling continued off the coast of Brazil for many years and the Imbituba land station processed right whales until 1973 The Soviet Union illegally took at least 3 212 southern right whales during the 1950s and 60s although it reported taking only four 58 nbsp A southern right whale approaches close to whale watchers near Peninsula Valdes in PatagoniaWhale watching edit Main article Whale watching The southern right whale has made Hermanus South Africa one of the world centers for whale watching During the winter months July October southern right whales come so close to the shoreline visitors can watch whales from strategically placed hotels 59 The town employs a whale crier cf town crier to walk through the town announcing where whales have been seen 60 In Brazil Imbituba in Santa Catarina has been recognized as the National Right Whale Capital and holds annual Right Whale Week celebrations in September 61 when mothers and calves are more often seen The old whaling station there has been converted to a museum dedicated to the whales 62 In winter in Argentina Peninsula Valdes in Patagonia hosts the largest breeding population of the species with more than 2 000 animals catalogued by the Whale Conservation Institute and Ocean Alliance 63 Conservation edit nbsp North Atlantic right whale on a Faroese stampBoth the North Atlantic and North Pacific species are listed as a species threatened with extinction which is or may be affected by trade Appendix I by CITES and as endangered by the IUCN Red List In the United States the National Marine Fisheries Service NMFS a subagency of the National Oceanic and Atmospheric Administration NOAA has classified all three species as endangered under the Endangered Species Act Under the Marine Mammal Protection Act they are listed as depleted 64 65 66 The southern right whale is listed as endangered under the Australian Environment Protection and Biodiversity Conservation Act as nationally endangered under the New Zealand Threat Classification System as a natural monument by the Argentine National Congress and as a State Natural Monument under the Brazilian National Endangered Species List 66 The US and Brazil added new protections for right whales in the 2000s to address the two primary hazards While environmental campaigners were as reported in 2001 pleased about the plan s positive effects they attempted to force the US government to do more 67 In particular they advocated 12 knots 22 km h speed limits for ships within 40 km 25 mi of US ports in times of high right whale presence Citing concerns about excessive trade disruption it did not institute greater protections The Defenders of Wildlife the Humane Society of the United States and the Ocean Conservancy sued the NMFS in September 2005 for failing to protect the critically endangered North Atlantic Right Whale which the agency acknowledges is the rarest of all large whale species and which federal agencies are required to protect by both the Marine Mammal Protection Act and the Endangered Species Act demanding emergency protection measures 68 The southern right whale listed as endangered by CITES and lower risk conservation dependent by the IUCN is protected in the jurisdictional waters of all countries with known breeding populations Argentina Australia Brazil Chile New Zealand South Africa and Uruguay In Brazil a federal Environmental Protection Area encompassing some 1 560 km2 600 sq mi and 130 km 81 mi of coastline in Santa Catarina State was established in 2000 to protect the species main breeding grounds in Brazil and promote whale watching 69 On February 6 2006 NOAA proposed its Strategy to Reduce Ship Strikes to North Atlantic Right Whales 70 The proposal opposed by some shipping interests limited ship speeds during calving season The proposal was made official when on December 8 2008 NOAA issued a press release that included the following 71 Effective January 2009 ships 65 feet 20 m or longer are limited to 10 knots 19 km h in waters off New England when whales begin gathering in this area as part of their annual migration The restriction extends to 20 nautical miles 37 km around major mid Atlantic ports The speed restriction applies in waters off New England and the southeastern US where whales gather seasonally Southeastern US from St Augustine Florida to Brunswick Georgia from Nov 15 to April 15 Mid Atlantic U S areas from Rhode Island to Georgia from Nov 1 to April 30 Cape Cod Bay from Jan 1 to May 15 Off Race Point at the northern end of Cape Cod from March 1 to April 30 Great South Channel of New England from April 1 to July 31 Temporary voluntary speed limits in other areas or times when a group of three or more right whales is confirmed Scientists would assess the rule s effectiveness before the rule expires in 2013 In 2020 NOAA published its assessment and found that since the speed rule was adopted the total number of documented deaths from vessel strike decreased but serious and non serious injuries have increased 72 A report by the organization Oceana found that between 2017 and 2020 disobedience of the rule reached close to 90 in mandatory speed zones while in voluntary areas disobedience neared 85 73 Threats edit nbsp The remains of a North Atlantic right whale after it collided with a ship propellerThe leading cause of death among the North Atlantic right whale which migrates through some of the world s busiest shipping lanes while journeying off the east coast of the United States and Canada is being struck by ships note 1 74 At least sixteen ship strike deaths were reported between 1970 and 1999 and probably more remain unreported 18 According to NOAA twenty five of the seventy one right whale deaths reported since 1970 resulted from ship strikes 71 A second major cause of morbidity and mortality in the North Atlantic right whale is entanglement in plastic fishing gear Right whales ingest plankton with wide open mouths risking entanglement in any rope or net fixed in the water column Rope wraps around their upper jaws flippers and tails Some are able to escape but others remain tangled 75 Whales can be successfully disentangled if observed and aided In July 1997 the U S NOAA introduced the Atlantic Large Whale Take Reduction Plan which seeks to minimize whale entanglement in fishing gear and record large whale sightings in an attempt to estimate numbers and distribution 76 In 2012 the U S Navy proposed to create a new undersea naval training range immediately adjacent to northern right whale calving grounds in shallow waters off the Florida Georgia border Legal challenges by leading environmental groups including the Natural Resources Defense Council were denied in federal court allowing the Navy to proceed 77 78 These rulings were made despite the extremely low numbers as low as 313 by some estimates of right whales in existence at this time and a very poor calving season 79 Notes edit The Lloyd s mirror effect results in low frequency propeller sounds not being discernible near the surface where most accidents occur Combined with spreading and acoustic shadowing effects the result is that the animal is unable to hear an approaching vessel before it has been run over or entrapped by the hydrodynamic forces of the vessel s passage References edit Fossilworks a b Mead J G Brownell R L Jr 2005 Order Cetacea In Wilson D E Reeder D M eds Mammal Species of the World A Taxonomic and Geographic Reference 3rd ed Johns Hopkins University Press pp 723 743 ISBN 978 0 8018 8221 0 OCLC 62265494 a b c Cooke J G Zerbini A N 2018 Eubalaena australis IUCN Red List of Threatened Species 2018 e T8153A50354147 doi 10 2305 IUCN UK 2018 1 RLTS T8153A50354147 en Retrieved November 12 2021 Bisconti M Lambert O Bosselaers M 2017 Revision of Balaena belgica reveals a new right whale species the possible ancestry of the northern right whale Eubalaena glacialis and the ages of divergence for the living right whale species PeerJ 5 e3464 https doi org 10 7717 peerj 3464 a b Cooke J G 2020 Eubalaena glacialis IUCN Red List of Threatened Species 2020 e T41712A178589687 doi 10 2305 IUCN UK 2020 2 RLTS T41712A178589687 en Retrieved November 12 2021 a b Cooke J G Clapham P J 2018 Eubalaena japonica IUCN Red List of Threatened Species 2018 e T41711A50380694 doi 10 2305 IUCN UK 2018 1 RLTS T41711A50380694 en Retrieved November 12 2021 Kimura T Narita K 2007 A new species of Eubalaena Cetacea Mysticeti Balaenidae from the Gonda Formation latest Miocene early Pliocene of Japan Bulletin of the Gunma Museum of Natural History 11 15 27 Perrin W F 2012 Eubalaena Gray 1864 World Cetacea Database Retrieved September 29 2012 North Atlantic right whale Basic Facts About Right Whales Defenders of Wildlife Defenders org Retrieved May 10 2012 Dolin Eric Jay 2007 Leviathan The History of Whaling in America W W Norton amp Co p 22 ISBN 978 0 393 06057 7 Retrieved September 30 2012 quoting Boston August 21 Connecticut Courant September 1 1766 also quoting Spears John Randolph 1908 The story of the New England whalers The Macmillan Company pp 80 81 Eubalaena glacialis ScienceDirect Retrieved March 20 2023 a b Muller J 1954 Observations of the orbital region of the skull of the Mystacoceti PDF Zoologische Mededelingen 32 239 90 Rice Dale W 1998 Marine Mammals of the World Systematics and distribution Special Publication Vol No 4 Society of Marine Mammalogy ISBN 1891276034 a b Rosenbaum H C Brownell R L Jr Brown M W Schaeff C Portway V White B N et al 2000 World wide genetic differentiation of Eubalaena Questioning the number of right whale species Molecular Ecology 9 11 1793 802 doi 10 1046 j 1365 294x 2000 01066 x PMID 11091315 S2CID 7166876 a href Template Cite journal html title Template Cite journal cite journal a CS1 maint multiple names authors list link a b Malik S Brown M W Kraus S D amp White B N 2000 Analysis of mitochondrial DNA diversity within and between North and South Atlantic right whales Marine Mammal Science 16 3 545 558 doi 10 1111 j 1748 7692 2000 tb00950 x Brownell R L Jr Clapham P J Miyashita T amp Kasuya T 2001 Conservation status of North Pacific right whales Journal of Cetacean Research and Management 2 269 286 List of marine mammal species and subspecies Committee on Taxonomy marinemammalscience org Society for Marine Mammalogy April 3 2012 Retrieved September 29 2012 a b c d e f g h i j Kenney Robert D February 26 2009 North Atlantic North Pacific and Southern right whales In Perrin William F Wursig Bernd Thewissen J G M Hans eds Encyclopedia of Marine Mammals Academic Press pp 806 813 ISBN 978 0 08 091993 5 Kaliszewska Z A Seger J Barco S G Benegas R Best P B Brown M W et al 2005 Population histories of right whales Cetacea Eubalaena inferred from mitochondrial sequence diversities and divergences of their whale lice Amphipoda Cyamus Molecular Ecology 14 11 3439 3456 doi 10 1111 j 1365 294X 2005 02664 x PMID 16156814 S2CID 803511 a href Template Cite journal html title Template Cite journal cite journal a CS1 maint multiple names authors list link a b Mysticeti Palaeobiology and Biodiversity Research Group University of Bristol Archived from the original on July 12 2011 Ross Alison September 20 2005 Whale riders reveal evolution BBC News Jackson Stephen Groves Colin 2015 Taxonomy of Australian Mammals Csiro Publishing p 315 ISBN 9781486300136 Retrieved February 22 2016 The placement of Caperea within the monotypic family Neobalaenidae is done with the qualification that future work may corroborate the referral of Caperea to the Family Cetotheriidae Fordyce R Ewan Marx Felix G 2013 The pygmy right whale Caperea marginata The last of the cetotheres Proceedings of the Royal Society B Biological Sciences 280 1753 20122645 doi 10 1098 rspb 2012 2645 PMC 3574355 PMID 23256199 Prehistoric whale discovered on the west coast of Sweden Sciencedaily com Press release June 5 2009 Retrieved May 10 2012 Perrin W F 2012 Hunterius swedenborgii Lilljeborg 1867 World Cetacea Database Retrieved September 29 2012 Whale bones found in highway were not from mystery whale ScienceNordic com February 7 2013 Retrieved March 26 2013 a b c Kenney Robert D 2008 Right Whales Eubalaena glacialis E japonica and E australis In Perrin W F Wursig B Thewissen J G M eds Encyclopedia of Marine Mammals Academic Press pp 962 69 ISBN 978 0 12 373553 9 Retrieved May 20 2012 a b c Crane J Scott R 2002 Eubalaena glacialis North Atlantic right whale Information Animal Diversity Web University of Michigan Museum of Zoology Retrieved April 30 2006 Carwardine Mark Camm Martin July 1 2010 Whales Dolphins and Porpoises Dorling Kindersley ISBN 978 1 4053 5794 4 Brueck Hilary January 19 2016 A Surprising Tool for Saving the Whales Facial Recognition Software Fortune Retrieved January 22 2016 a b Fig S3 Stranded Pacific right whale Eubalaena japonica at Izu Woodward Becky L Winn Jeremy P Fish Frank E 2006 Morphological Specializations of Baleen Whales Associated With Hydrodynamic Performance and Ecological Niche PDF Journal of Morphology 267 11 1284 1294 doi 10 1002 jmor 10474 PMID 17051544 S2CID 14231425 Archived from the original PDF on March 5 2016 The true buoyancy of a particular whale is dependent upon its body composition particularly the relative quantities of muscle and blubber tissues Balaenopterid whales have a higher proportion of muscle tissue and tend to be negatively buoyant while the opposite is true for right whales Lockyer 1976 Lockyer C 1976 Body weights of some species of large whales Journal du Conseil International pour l Exploration de la Mer 36 3 259 273 doi 10 1093 icesjms 36 3 259 Oceanus Magazine Images Doing the Right Thing for the Right Whale Woods Hole Oceanographic Institution Retrieved on November 5 2014 Feldhamer George A Thompson Bruce C Chapman Joseph A 2003 Wild mammals of North America biology management and conservation 2nd ed Baltimore Md Johns Hopkins University Press p 432 ISBN 9780801874161 Brownell Robert L Jr Ralls K 1986 Potential for sperm competition in baleen whales Report of the International Whaling Commission 8 8 97 112 Waerebeek V K Santillan L Suazo E The Peruvian Centre for Cetacean Research 2009 ON THE NATIVE STATUS OF THE SOUTHERN RIGHT WHALE EUBALAENA AUSTRALIS IN PERU PDF Boletin del Museo Nacional de Historia Natural Chile 58 75 82 Retrieved August 3 2016 Moore M November 3 2004 Whither the North Atlantic Right Whale Scientists explore many facets of whales lives to help species on the edge of extinction Woods Hole Oceanographic Institution Retrieved August 10 2012 Notarbartolo di Sciara G E Politi A Bayed P C Beaubrun amp A Knowlton 1998 A winter cetacean survey off Southern Morocco with a special emphasis on suitable habitats for wintering right whales Reports of the International Whaling Commission 48 547 550 Martin AR Walker FJ May 16 1996 Sighting of a right whale Eubalaena glacialis with calf off S W Portugal Marine Mammal Science 13 1 139 140 doi 10 1111 j 1748 7692 1997 tb00617 x Archived from the original on August 13 2011 Retrieved October 26 2006 Jacobsen K O Marx M Oien N May 21 2003 Two Way Trans Atlantic Migration of a North Atlantic Right Whale Eubalaena Glacialis Marine Mammal Science 20 1 161 166 doi 10 1111 j 1748 7692 2004 tb01147 x Archived from the original on August 13 2011 Retrieved October 26 2006 Viegas Jennifer June 30 2010 Smallest whale population identified Discovery News Retrieved May 19 2012 a b Rincon Paul December 3 2003 Northern Right Whales respond to emergency sirens BBC News Retrieved May 10 2012 North Pacific Right Whale Alaska Sea Grant Marine Education Retrieved August 9 2012 Southern right whales of Patagonia PDF Whale and Dolphin Conservation Society Archived from the original PDF on July 6 2010 Retrieved August 7 2012 Gaines CA Hare MP Beck SE Rosenbaum HC March 7 2005 Nuclear markers confirm taxonomic status and relationships among highly endangered and closely related right whale species Proceedings of the Royal Society B 272 1562 533 542 doi 10 1098 rspb 2004 2895 PMC 1578701 PMID 15846869 a b A R Knowlton S D Kraus amp R D Kenney 1994 Reproduction in North Atlantic right whales Eubalaena glacialis Canadian Journal of Zoology 72 7 1297 1305 doi 10 1139 z94 173 P K Hamilton A R Knowlton M K Marx amp S D Kraus 1998 Age structure and longevity in North Atlantic right whales Eubalaena glacialis and their relation to reproduction Marine Ecology Progress Series 171 285 292 Bibcode 1998MEPS 171 285H doi 10 3354 meps171285 R Payne V Rowntree J S Perkins J G Cooke amp K Lankester 1990 Population size trends and reproductive parameters of right whales Eubalaena australis off Peninsula Valdez Argentina Report of the International Whaling Commission Special Issue 12 271 278 Leaper R Cooke J Trathan P Reid K Rowntree V Payne R 2005 Global climate drives southern right whale Eubalaena australis population dynamics PDF Biology Letters 2 2 289 292 doi 10 1098 rsbl 2005 0431 PMC 1618902 PMID 17148385 Retrieved August 16 2016 Boesak T 2015 Southern right whales distribution and reproduction The Ocean Blue Retrieved August 16 2016 P B Best 1994 Seasonality of reproduction and the length of gestation in southern right whales Eubalaena australis Journal of Zoology London 32 2 175 189 doi 10 1111 j 1469 7998 1994 tb01567 x Payne R 1986 Long term behavioral studies of the southern right whale Eubalaena australis PDF Scientific Reports at International Whaling Commission 10 161 167 Retrieved August 16 2016 Southern Right Whales New Zealand Whale and Dolphin Trust Retrieved August 16 2016 Katona S K Kraus S D 1999 Efforts to conserve the North Atlantic right whale In Twiss John R Reeves Randall R Montgomery Suzanne eds Conservation and Management of Marine Mammals Smithsonian Press pp 311 331 ISBN 978 1 56098 778 9 Scarff JE 2001 Preliminary estimates of whaling induced mortality in the 19th century Pacific northern right whale Eubalaena glacialis fishery adjusting for struck but lost whales and non American whaling J Cetacean Res Manag Special Issue 2 261 268 Tonnessen Johan Nicolay Johnsen Arne Odd 1982 The History of Modern Whaling University of California Press ISBN 978 0 520 03973 5 National Audubon Society 2002 Guide to Marine Mammals of the World A A Knopf ISBN 978 0 375 41141 0 Whale Watching in Hermanus at the Windsor Hotel Windsor Hotel website Retrieved August 10 2012 Hermanus Whale Crier SA Venues com Retrieved August 7 2012 News From Brazil Cetacean Society International Archived from the original on April 14 2013 Retrieved August 10 2012 Ribeiro P Whale Watching in Santa Catarina Brazil Fascinating Encounters With Southern Right Whales About com Brazil Travel Retrieved August 10 2012 Ocean Alliance website Oceanalliance org Archived from the original on March 4 2015 Retrieved May 10 2012 North Atlantic Right Whales Eubalaena glacialis NOAA Fisheries Office of Protected Resources National Marine Fisheries Service Retrieved September 4 2012 North Pacific Right Whales Eubalaena japonica NOAA Fisheries Office of Protected Resources National Marine Fisheries Service Retrieved September 4 2012 a b Southern Right Whales Eubalaena australis NOAA Fisheries Office of Protected Resources National Marine Fisheries Service Retrieved September 4 2012 Right whales need extra protection BBC News BBC News November 28 2001 Retrieved May 2 2006 The Southeast United States Right Whale Recovery Plan Implementation Team and the Northeast Implementation Team November 2005 NMFS and Coast Guard Inactions Bring Litigation PDF Right Whale News 12 4 Retrieved May 2 2006 Area de Protecao Ambiental in Portuguese Projeto Baleia Franca Archived from the original on July 28 2012 Retrieved September 4 2012 NOAA Proposed Strategy to Reduce Ship Strikes to North Atlantic Right Whales a b NOAA December 8 2008 Press Release on Effective Date of Speed Regulations PDF Retrieved December 21 2008 North Atlantic Right Whale Eubalaena glacialis Vessel Speed Rule Assessment PDF Report National Oceanic and Atmospheric Administration June 2020 Retrieved May 4 2022 Speeding Toward Extinction Vessel Strikes Threaten North Atlantic Right Whales Oceana org Retrieved May 4 2022 Vanderlaan amp Taggart 2007 Vessel collisions with whales the probability of lethal injury based on vessel speed PDF Mar Mam Sci Retrieved May 10 2008 Whales Entangled In Fishing Lines What Can Be Done Science Daily Retrieved August 7 2012 Atlantic Large Whale Take Reduction Plan National Oceanic and Atmospheric Administration 1997 Archived from the original on September 24 2008 Retrieved May 2 2006 Strachan Deshayla September 12 2012 Whale Defenders Lose Navy Training Challenge Courthouse News Service Archived from the original on September 23 2015 Retrieved September 20 2012 Heimer Taryn Kiekow September 10 2012 Right Whales Wronged Judge allows Navy to expand sonar use in Florida calving area Natural Resources Defense Council Retrieved October 9 2012 Poor calving season for right whales savannahnow com Archived from the original on January 7 2015 Further reading editKraus Scott D Rolland Rosalind eds 2010 The Urban Whale North Atlantic Right Whales at the Crossroads Harvard University Press ISBN 978 0 674 03475 4 External links edit nbsp Wikimedia Commons has media related to Eubalaena nbsp Wikispecies has information related to Eubalaena IUCN Red List entry Right Whale Lesson Plan from Smithsonian Education North Atlantic Right Whale on the Smithsonian Ocean Portal National Library for the Environment The Fall and Rise of the Right Whale New York Times March 16 2009 Portals nbsp Cetaceans nbsp Mammals nbsp Animals nbsp Biology nbsp Marine Life Retrieved from https en wikipedia org w index php title Right whale amp oldid 1183912883, wikipedia, wiki, book, books, library,

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