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Papilio helenus

Papilio helenus, the red Helen,[1][2] is a large swallowtail butterfly found in forests of southern India and parts of southeast Asia.[1][2]

Red Helen
P. h. fortunius
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Family: Papilionidae
Genus: Papilio
Species:
P. helenus
Binomial name
Papilio helenus

Range edit

Papilio helenus is rarely found in Sri Lanka, southern and north-east India, Nepal, Bhutan, Bangladesh, Myanmar, Thailand, Laos, Kampuchea, Vietnam, southern China (including Hainan, Guangdong province), Taiwan, southern Japan, South Korea, Ryukyu Islands, peninsular and eastern Malaysia, Brunei, Philippines, and Indonesia (Sumatra, Java, Bangka, Kalimantan, the Lesser Sunda Islands except Tanimbar).[1][2]

In India it occurs along the Western Ghats from Kerala to Gujarat, also Palnis and Shevaroys, in the north from Mussoorie eastwards, to north-east India and onto Myanmar.[1][2][3][4]

Description edit

 
P. h. daksha, Kerala, India
 
Woodcut from The Fauna of British India, Including Ceylon and Burma

Male. Upper-side rich brownish-black ; cilia alternated with white. Fore-wing with slender longitudinal streaks of golden-yellow scales within the cell, and hairy-scaled broad streaks between the outer veins ; lower base of the wing also sparsely speckled with golden-yellow scales. Hind-wing with a cream- white upper discal patch composed of three portions, a short almost quadrate portion in the costal interspace, and two lower large elongate portions in the subcostal interspaces, a few white or grey scales also being present below the lower portion ; at the anal angle is a dark purple-red circlet which generally partly encloses a round black spot, followed by a lower submarginal series of three, sometimes four, more or less apparent purple-red incurved lunules, the first or lower one sometimes forming a circle round a black spot ; in some specimens the lower lunule is obsolescent, in others all the latter are obsolete. Underside paler. Fore-wing with prominent greyish-white scaled slender cell-streaks and bread outer vein-streaks. Hind-wing with the upper basal area speckled with greyish-white scale.*, and three similar scaled slender longitudinal streaks within the cell, the base of the median and sub median vein also being white scaled; upper discal white patch composed of three smaller portions than on the upper-side; anal ocellus and a complete series of submarginal lunules dark red ; an inner sub anal confluent lunles also present in the lower median interspace ; the anal and sub anal lunules being speckled with violet-blue scales; beyond is a marginal row of more or less defined red-and-white scaled lunules.

Female. Upper-side and underside similar to the male, except that the base of the hind-wing is paler.

Body brownish-black; collar, front of head and palpi white-spotted; tegulae white fringed; legs and antenna black.

Status edit

Papilio helenus is generally uncommon and slightly threatened ( in certain places) . It is very commonly found Maharashtra, but also rarely in Gujarat and parts of Kerala. Its history can be traced back to Assam and Western Ghats, where it is currently endemic.[5]

Taxonomy edit

 
Illustration

Species group edit

Papilio helenus is the nominate member of the helenus species group. The members of this clade are:

Subspecies edit

There are up to thirteen different subspecies, two of which occur in India:

  • P. h. daksha Moore - South India. Not rare
  • P. h. helenus Linnaeus - Mussoorie to Myanmar. Common

and one in Taiwan:

  • P. h. fortunius Fruhstorfer, 1908

Life cycle edit

 
Similar species

This butterfly flies throughout the year in southern India.

Eggs edit

The egg is pale apricot-yellow in colour when freshly deposited, spherical in shape and has a slightly roughened exterior which looks like the skin of an orange when seen through a microscope. The diameter of an egg is 1.2 mm.

The eggs are deposited singly on the tips of very young leaves and shoots in shady parts of thick jungle. Before hatching, the eggs appear to be marked by chocolate coloured lines and flecks. The egg hatches in 4 to 7 days.

Caterpillar edit

The freshly emerged larva is about 3 mm long. Throughout the caterpillar stage, if it is agitated it can evert a yellow-to-red osmeterium from the first segment, just behind the head. Each of the other segments bears, on the back, a pair of tufts of stiff hairs, each tuft arising from a small, yellowish conical process. The overall colour is brown, but there is a whitish saddle-like patch about the middle and the tail segments are also whitish in colour.

Like other Papilio species the larva can evert a two-pronged horn-like osmeterium when it is irritated. The osmeterium secretes an repugnatorial|unpleasant-smelling liquid which is believed to repel predators and parasites.

After the first moult the caterpillar has the appearance of a shiny bird dropping. The larva is grass green in colour, mottled black and white and smoky grey. The osmeterium, when everted, is generally yellow to red.

While inactive, mainly during daylight hours, the young larva lies along the midrib of the underside of the leaf. Later on, when it is largely fully grown, it is greener and lies on the centre of the upperside of the leaf, on a stem or a twig. The fifth instar larva is about 5 cm long.

See also edit

References edit

  1. ^ a b c d Varshney, R.K.; Smetacek, Peter (2015). A Synoptic Catalogue of the Butterflies of India. New Delhi: Butterfly Research Centre, Bhimtal & Indinov Publishing, New Delhi. p. 8. doi:10.13140/RG.2.1.3966.2164. ISBN 978-81-929826-4-9.
  2. ^ a b c d Savela, Markku. "Papilio helenus Linnaeus, 1758". Lepidoptera and Some Other Life Forms. Retrieved July 3, 2018.
  3. ^ Bingham, C.T. (1907). The Fauna of British India, Including Ceylon and Burma. Vol. II (1st ed.). London: Taylor and Francis, Ltd. pp. 41–43.
  4. ^ a b   One or more of the preceding sentences incorporates text from this source, which is in the public domain: Moore, Frederic (1901–1903). Lepidoptera Indica. Vol. V. London: Lovell Reeve and Co. pp. 208–210.
  5. ^ A Pictorial Guide Butterflies of Gorumara National Park (1st ed.). Jalpaiguri, West Bengal: Department of Forests Government of West Bengal. March 2013. p. 18.
  • Collins, N. Mark; Morris, Michael G. (1985). Threatened Swallowtail Butterflies of the World: The IUCN Red Data Book. Gland & Cambridge: IUCN. ISBN 978-2-88032-603-6 – via Biodiversity Heritage Library.
  • Evans, W.H. (1932). The Identification of Indian Butterflies (2nd ed.). Mumbai, India: Bombay Natural History Society.
  • Gay, Thomas; Kehimkar, Isaac David; Punetha, Jagdish Chandra (1992). Common Butterflies of India. Nature Guides. Bombay, India: World Wide Fund for Nature-India by Oxford University Press. ISBN 978-0195631647.
  • Haribal, Meena (1992). The Butterflies of Sikkim Himalaya and Their Natural History. Gangtok, Sikkim, India: Sikkim Nature Conservation Foundation.
  • Kunte, Krushnamegh (2000). Butterflies of Peninsular India. India, A Lifescape. Hyderabad, India: Universities Press. ISBN 978-8173713545.
  • Wynter-Blyth, Mark Alexander (1957). Butterflies of the Indian Region. Bombay, India: Bombay Natural History Society. ISBN 978-8170192329.

External links edit

  •   Media related to Papilio helenus at Wikimedia Commons

papilio, helenus, helen, large, swallowtail, butterfly, found, forests, southern, india, parts, southeast, asia, helen, fortunius, scientific, classification, domain, eukaryota, kingdom, animalia, phylum, arthropoda, class, insecta, order, lepidoptera, family,. Papilio helenus the red Helen 1 2 is a large swallowtail butterfly found in forests of southern India and parts of southeast Asia 1 2 Red Helen P h fortunius Scientific classification Domain Eukaryota Kingdom Animalia Phylum Arthropoda Class Insecta Order Lepidoptera Family Papilionidae Genus Papilio Species P helenus Binomial name Papilio helenusLinnaeus 1758 Contents 1 Range 2 Description 3 Status 4 Taxonomy 4 1 Species group 4 2 Subspecies 5 Life cycle 5 1 Eggs 5 2 Caterpillar 6 See also 7 References 8 External linksRange editPapilio helenus is rarely found in Sri Lanka southern and north east India Nepal Bhutan Bangladesh Myanmar Thailand Laos Kampuchea Vietnam southern China including Hainan Guangdong province Taiwan southern Japan South Korea Ryukyu Islands peninsular and eastern Malaysia Brunei Philippines and Indonesia Sumatra Java Bangka Kalimantan the Lesser Sunda Islands except Tanimbar 1 2 In India it occurs along the Western Ghats from Kerala to Gujarat also Palnis and Shevaroys in the north from Mussoorie eastwards to north east India and onto Myanmar 1 2 3 4 Description edit nbsp P h daksha Kerala India nbsp Woodcut from The Fauna of British India Including Ceylon and Burma For a key to the terms used see Glossary of entomology terms Male Upper side rich brownish black cilia alternated with white Fore wing with slender longitudinal streaks of golden yellow scales within the cell and hairy scaled broad streaks between the outer veins lower base of the wing also sparsely speckled with golden yellow scales Hind wing with a cream white upper discal patch composed of three portions a short almost quadrate portion in the costal interspace and two lower large elongate portions in the subcostal interspaces a few white or grey scales also being present below the lower portion at the anal angle is a dark purple red circlet which generally partly encloses a round black spot followed by a lower submarginal series of three sometimes four more or less apparent purple red incurved lunules the first or lower one sometimes forming a circle round a black spot in some specimens the lower lunule is obsolescent in others all the latter are obsolete Underside paler Fore wing with prominent greyish white scaled slender cell streaks and bread outer vein streaks Hind wing with the upper basal area speckled with greyish white scale and three similar scaled slender longitudinal streaks within the cell the base of the median and sub median vein also being white scaled upper discal white patch composed of three smaller portions than on the upper side anal ocellus and a complete series of submarginal lunules dark red an inner sub anal confluent lunles also present in the lower median interspace the anal and sub anal lunules being speckled with violet blue scales beyond is a marginal row of more or less defined red and white scaled lunules Female Upper side and underside similar to the male except that the base of the hind wing is paler Body brownish black collar front of head and palpi white spotted tegulae white fringed legs and antenna black Frederic Moore Lepidoptera Indica Vol V 4 Status editPapilio helenus is generally uncommon and slightly threatened in certain places It is very commonly found Maharashtra but also rarely in Gujarat and parts of Kerala Its history can be traced back to Assam and Western Ghats where it is currently endemic 5 Taxonomy edit nbsp Illustration Species group edit Papilio helenus is the nominate member of the helenus species group The members of this clade are Papilio helenus Linnaeus 1758 Papilio iswara White 1842 Papilio iswaroides Fruhstorfer 1898 Papilio nephelus Boisduval 1836 Papilio nubilus Staudinger 1895 Papilio sataspes C amp R Felder 1865 Subspecies edit There are up to thirteen different subspecies two of which occur in India P h daksha Moore South India Not rare P h helenus Linnaeus Mussoorie to Myanmar Common and one in Taiwan P h fortunius Fruhstorfer 1908Life cycle edit nbsp Similar species This butterfly flies throughout the year in southern India Eggs edit The egg is pale apricot yellow in colour when freshly deposited spherical in shape and has a slightly roughened exterior which looks like the skin of an orange when seen through a microscope The diameter of an egg is 1 2 mm The eggs are deposited singly on the tips of very young leaves and shoots in shady parts of thick jungle Before hatching the eggs appear to be marked by chocolate coloured lines and flecks The egg hatches in 4 to 7 days Caterpillar edit The freshly emerged larva is about 3 mm long Throughout the caterpillar stage if it is agitated it can evert a yellow to red osmeterium from the first segment just behind the head Each of the other segments bears on the back a pair of tufts of stiff hairs each tuft arising from a small yellowish conical process The overall colour is brown but there is a whitish saddle like patch about the middle and the tail segments are also whitish in colour Like other Papilio species the larva can evert a two pronged horn like osmeterium when it is irritated The osmeterium secretes an repugnatorial unpleasant smelling liquid which is believed to repel predators and parasites After the first moult the caterpillar has the appearance of a shiny bird dropping The larva is grass green in colour mottled black and white and smoky grey The osmeterium when everted is generally yellow to red While inactive mainly during daylight hours the young larva lies along the midrib of the underside of the leaf Later on when it is largely fully grown it is greener and lies on the centre of the upperside of the leaf on a stem or a twig The fifth instar larva is about 5 cm long See also editPapilionidae List of butterflies of India List of butterflies of India Papilionidae References edit a b c d Varshney R K Smetacek Peter 2015 A Synoptic Catalogue of the Butterflies of India New Delhi Butterfly Research Centre Bhimtal amp Indinov Publishing New Delhi p 8 doi 10 13140 RG 2 1 3966 2164 ISBN 978 81 929826 4 9 a b c d Savela Markku Papilio helenus Linnaeus 1758 Lepidoptera and Some Other Life Forms Retrieved July 3 2018 Bingham C T 1907 The Fauna of British India Including Ceylon and Burma Vol II 1st ed London Taylor and Francis Ltd pp 41 43 a b nbsp One or more of the preceding sentences incorporates text from this source which is in the public domain Moore Frederic 1901 1903 Lepidoptera Indica Vol V London Lovell Reeve and Co pp 208 210 A Pictorial Guide Butterflies of Gorumara National Park 1st ed Jalpaiguri West Bengal Department of Forests Government of West Bengal March 2013 p 18 Collins N Mark Morris Michael G 1985 Threatened Swallowtail Butterflies of the World The IUCN Red Data Book Gland amp Cambridge IUCN ISBN 978 2 88032 603 6 via Biodiversity Heritage Library Evans W H 1932 The Identification of Indian Butterflies 2nd ed Mumbai India Bombay Natural History Society Gay Thomas Kehimkar Isaac David Punetha Jagdish Chandra 1992 Common Butterflies of India Nature Guides Bombay India World Wide Fund for Nature India by Oxford University Press ISBN 978 0195631647 Haribal Meena 1992 The Butterflies of Sikkim Himalaya and Their Natural History Gangtok Sikkim India Sikkim Nature Conservation Foundation Kunte Krushnamegh 2000 Butterflies of Peninsular India India A Lifescape Hyderabad India Universities Press ISBN 978 8173713545 Wynter Blyth Mark Alexander 1957 Butterflies of the Indian Region Bombay India Bombay Natural History Society ISBN 978 8170192329 External links edit nbsp Media related to Papilio helenus at Wikimedia Commons Retrieved from https en wikipedia org w index php title Papilio helenus amp oldid 1164627074, wikipedia, wiki, book, books, library,

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