Calamus (palm)

Calamus is a genus of flowering plants in the palm family Arecaceae that are among several genera known as rattan palms. There are an estimated 400 species in this genus, all native to tropical and subtropical Asia, Africa, and Australia.^[2][3][4]

Calamus
Calamus gibbsianus
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Monocots
Clade:	Commelinids
Order:	Arecales
Family:	Arecaceae
Subfamily:	Calamoideae
Tribe:	Calameae
Genus:	Calamus L.
Species
Numerous, see text
Synonyms^[1]
Calospatha Becc. Ceratolobus Blume ex Schult. & Schult.f. Cornera Furtado Daemonorops Blume Palmijuncus Rumph. ex Kuntze Pogonotium J.Dransf. Retispatha J.Dransf. Rotang Adans. Rotanga Boehm. Schizospatha Furtado Zalaccella Becc.

Description

They are dioecious,^[5] mostly leaf-climbing lianas with slender, reedy stems. To aid scrambling some species have evolved hooks on the underside of the midrib, or more commonly by modified "pinnae" or tendrils in the form of stout, backward-pointing spines. These stems may grow to lengths of 200 metres.^[6][7]

The fruits of certain species, in particular Calamus draco, produce a red resin known as "Dragon's blood".^[8]

Species

As of April 2022^[update], Plants of the World Online recognises 415 species:^[1]

1. Calamus acamptostachys (Becc.) W.J.Baker
2. Calamus acanthochlamys J.Dransf.
3. Calamus acanthophyllus Becc.
4. Calamus acanthospathus Griff.
5. Calamus acaulis A.J.Hend., N.K.Ban & N.Q.Dung
6. Calamus adspersus Blume
7. Calamus aidae Fernando
8. Calamus albidus L.X.Guo & A.J.Hend.
9. Calamus altiscandens Burret
10. Calamus andamanicus Kurz
11. Calamus anomalus Burret
12. Calamus applanatus (A.J.Hend. & N.Q.Dung) A.J.Hend.
13. Calamus arborescens Griff.
14. Calamus aruensis Becc.
15. Calamus arugda Becc.
16. Calamus ashtonii J.Dransf.
17. Calamus asperrimus Blume
18. Calamus asteracanthus (Becc.) W.J.Baker
19. Calamus ater (J.Dransf.) W.J.Baker
20. Calamus australis Mart.
21. Calamus axillaris Becc.
22. Calamus bacularis Becc.
23. Calamus badius J.Dransf. & W.J.Baker
24. Calamus baiyerensis W.J.Baker & J.Dransf.
25. Calamus balerensis Fernando
26. Calamus balingensis Furtado
27. Calamus banggiensis (J.Dransf.) W.J.Baker
28. Calamus bankae W.J.Baker & J.Dransf.
29. Calamus baratangensis Renuka & Vijayak.
30. Calamus barbatus Zipp. ex Blume
31. Calamus barfodii W.J.Baker & J.Dransf.
32. Calamus barisanensis A.J.Hend.
33. Calamus batanensis (Becc.) Baja-Lapis
34. Calamus beccarii A.J.Hend.
35. Calamus bicolor Becc.
36. Calamus billitonensis Becc. ex K.Heyne
37. Calamus bimanifer T.Evans, Sengdala, Viengkham, Thamm. & J.Dransf.
38. Calamus boniensis Becc. ex K.Heyne
39. Calamus bousigonii Pierre ex Becc.
40. Calamus brandisii Becc.
41. Calamus brevicaulis (A.J.Hend. & N.Q.Dung) W.J.Baker
42. Calamus brevissimus A.J.Hend.
43. Calamus brunneus A.J.Hend., Pitopang & Moh.Iqbal
44. Calamus bulubabi W.J.Baker & J.Dransf.
45. Calamus burckianus Becc.
46. Calamus burkillianus Becc. ex Ridl.
47. Calamus caesius Blume
48. Calamus calapparius Mart.
49. Calamus calciphilus A.J.Hend.
50. Calamus calicarpus Griff.
51. Calamus calospathus (Ridl.) W.J.Baker & J.Dransf.
52. Calamus capillosus W.J.Baker & J.Dransf.
53. Calamus carsicola Adorador & Fernando
54. Calamus caryotoides A.Cunn. ex Mart.
55. Calamus castaneus Griff.
56. Calamus caurinus A.J.Hend. & Rustiami
57. Calamus centralis A.J.Hend., N.K.Ban & N.Q.Dung
58. Calamus ceratophorus Conrard
59. Calamus cheirophyllus J.Dransf. & W.J.Baker
60. Calamus ciliaris Blume
61. Calamus cinereus A.J.Hend. & N.Q.Dung
62. Calamus clivorum A.J.Hend. & N.Q.Dung
63. Calamus cockburnii J.Dransf.
64. Calamus compsostachys Burret
65. Calamus comptus J.Dransf.
66. Calamus concinnus Mart.
67. Calamus concolor (Blume) W.J.Baker
68. Calamus confusus (Furtado) W.J.Baker
69. Calamus conirostris Becc.
70. Calamus conjugatus Furtado
71. Calamus conspectus W.J.Baker
72. Calamus convallium J.Dransf.
73. Calamus crassifolius J.Dransf.
74. Calamus crinitus (Blume) Miq.
75. Calamus cristatus (Becc.) W.J.Baker
76. Calamus croftii J.Dransf. & W.J.Baker
77. Calamus cumingianus Becc.
78. Calamus cuthbertsonii Becc.
79. Calamus dasyacanthus W.J.Baker, Bayton, J.Dransf. & Maturb.
80. Calamus deerratus G.Mann & H.Wendl.
81. Calamus delicatulus Thwaites
82. Calamus densifloropsis A.J.Hend.
83. Calamus densiflorus Becc.
84. Calamus depauperatus Ridl.
85. Calamus dianbaiensis C.F.Wei
86. Calamus didymocarpus Warb. ex Becc.
87. Calamus diepenhorstii Miq.
88. Calamus digitatus Becc.
89. Calamus dilaceratus Becc.
90. Calamus dioicus Lour.
91. Calamus discolor Mart.
92. Calamus disjunctus A.J.Hend.
93. Calamus distentus Burret
94. Calamus divaricatus Becc.
95. Calamus divergens A.J.Hend., Pitopang & Moh.Iqbal
96. Calamus dongnaiensis Pierre ex Becc.
97. Calamus dracunculus (Ridl.) W.J.Baker
98. Calamus dumetosus (J.Dransf.) A.J.Hend. & Floda
99. Calamus egregius Burret
100. Calamus elegans Becc. ex Ridl.
101. Calamus erectus Roxb.
102. Calamus erinaceus (Becc.) Becc.
103. Calamus erioacanthus Becc.
104. Calamus erythrocarpus W.J.Baker & J.Dransf.
105. Calamus essigii W.J.Baker
106. Calamus eugenei W.J.Baker
107. Calamus evansii A.J.Hend.
108. Calamus exiguus A.J.Hend.
109. Calamus exilis Griff.
110. Calamus fertilis Becc.
111. Calamus filipendulus Becc.
112. Calamus filispadix Becc.
113. Calamus fissilis A.J.Hend., N.K.Ban & N.Q.Dung
114. Calamus flabellatus Becc.
115. Calamus flagellum Griff. ex Walp.
116. Calamus floribundus Griff.
117. Calamus formicarius (Becc.) W.J.Baker
118. Calamus formosanus Becc.
119. Calamus furvus A.J.Hend.
120. Calamus gaharuensis A.J.Hend.
121. Calamus gajoensis A.J.Hend. & Rustiami
122. Calamus gamblei Becc.
123. Calamus geniculatus Griff.
124. Calamus gibbsianus Becc.
125. Calamus glaucescens (Blume) D.Dietr.
126. Calamus godefroyi Becc.
127. Calamus gonospermus Becc.
128. Calamus goramensis A.J.Hend.
129. Calamus gracilipes Miq.
130. Calamus gracilis Roxb.
131. Calamus griseus J.Dransf.
132. Calamus guruba Buch.-Ham. ex Mart.
133. Calamus hallierianus (Becc. ex K.Heyne) W.J.Baker
134. Calamus harmandii Pierre ex Becc.
135. Calamus heatubunii W.J.Baker & J.Dransf.
136. Calamus helferianus Kurz
137. Calamus henryanus Becc.
138. Calamus heteracanthopsis A.J.Hend.
139. Calamus heteracanthus Zipp. ex Blume
140. Calamus heteroideus Blume
141. Calamus hirsutus (Blume) Miq.
142. Calamus holttumii Furtado
143. Calamus hookerianus Becc.
144. Calamus horrens Blume
145. Calamus hosensis A.J.Hend.
146. Calamus hukaungensis A.J.Hend.
147. Calamus impressus A.J.Hend., Pitopang & Moh.Iqbal
148. Calamus inermis T.Anderson
149. Calamus ingens (J.Dransf.) W.J.Baker
150. Calamus inops Becc. ex K.Heyne
151. Calamus insignis Griff.
152. Calamus insolitus A.J.Hend.
153. Calamus insularis A.J.Hend.
154. Calamus interruptus Becc.
155. Calamus jacobsii W.J.Baker & J.Dransf.
156. Calamus javensis Blume
157. Calamus johanis A.J.Hend.
158. Calamus johndransfieldii W.J.Baker
159. Calamus johnsii W.J.Baker & J.Dransf.
160. Calamus kampucheaensis A.J.Hend. & Hourt
161. Calamus kandariensis Becc.
162. Calamus karuensis Ridl.
163. Calamus katikii W.J.Baker & J.Dransf.
164. Calamus kebariensis Maturb., J.Dransf. & W.J.Baker
165. Calamus kinabaluensis A.J.Hend.
166. Calamus kingianus Becc.
167. Calamus kjellbergii Furtado
168. Calamus klossii Ridl.
169. Calamus kontumensis A.J.Hend., N.K.Ban & N.Q.Dung
170. Calamus koordersianus Becc.
171. Calamus kostermansii W.J.Baker & J.Dransf.
172. Calamus kubahensis A.J.Hend.
173. Calamus kunstleri (Becc.) W.J.Baker
174. Calamus lakshmanae Renuka
175. Calamus lambirensis J.Dransf.
176. Calamus lamprolepis (Becc.) W.J.Baker
177. Calamus laoensis T.Evans, Sengdala, Viengkham, Thamm. & J.Dransf.
178. Calamus lateralis A.J.Hend., N.K.Ban & N.Q.Dung
179. Calamus latifolius Roxb.
180. Calamus latus A.J.Hend.
181. Calamus lauterbachii Becc.
182. Calamus laxissimus Ridl.
183. Calamus leiocaulis Becc. ex K.Heyne
184. Calamus leloi J.Dransf.
185. Calamus lengguanii A.J.Hend.
186. Calamus leptopus Griff.
187. Calamus leptospadix Griff.
188. Calamus leptostachys Becc. ex K.Heyne
189. Calamus lobatus A.J.Hend., Pitopang & Moh.Iqbal
190. Calamus lobbianus Becc.
191. Calamus loherianus (Becc.) W.J.Baker
192. Calamus longibracteatus W.J.Baker
193. Calamus longipes Griff.
194. Calamus longipinna K.Schum. & Lauterb.
195. Calamus longisetus Griff.
196. Calamus longispatha Ridl.
197. Calamus longiusculus A.J.Hend.
198. Calamus lucysmithiae W.J.Baker & J.Dransf.
199. Calamus macrochlamys Becc.
200. Calamus macropterus Miq.
201. Calamus macrorhynchus Burret
202. Calamus macrosphaerion Becc.
203. Calamus maculatus (J.Dransf.) W.J.Baker
204. Calamus mahanandensis S.Mondal, S.K.Basu & M.Chowdhury
205. Calamus maiadum J.Dransf.
206. Calamus malawaliensis J.Dransf.
207. Calamus manan Miq.
208. Calamus manglaensis A.J.Hend. & N.Q.Dung
209. Calamus manillensis (Mart.) H.Wendl.
210. Calamus marginatus (Blume) Mart. ex Walp.
211. Calamus maturbongsii W.J.Baker & J.Dransf.
212. Calamus megaphyllus Becc.
213. Calamus meghalayensis A.J.Hend.
214. Calamus melanacanthos Mart.
215. Calamus melanochaetes (Blume) Miq.
216. Calamus melanochrous Burret
217. Calamus melanoloma Mart.
218. Calamus melanorhynchus Becc.
219. Calamus metzianus Schltdl.
220. Calamus micracanthus Griff.
221. Calamus micranthus Blume
222. Calamus microsphaerion Becc.
223. Calamus minahassae Warb. ex Becc.
224. Calamus minor A.J.Hend.
225. Calamus minutus J.Dransf.
226. Calamus mirabilis Mart.
227. Calamus mitis Becc.
228. Calamus modestus T.Evans & T.P.Anh
229. Calamus mogeae J.Dransf.
230. Calamus mollispinus (J.Dransf.) W.J.Baker
231. Calamus moorei (J.Dransf.) W.J.Baker
232. Calamus moorhousei Furtado
233. Calamus moseleyanus Becc.
234. Calamus moszkowskianus Becc.
235. Calamus moti F.M.Bailey
236. Calamus muelleri H.Wendl.
237. Calamus multispicatus Burret
238. Calamus muricatus Becc.
239. Calamus myriacanthus Becc.
240. Calamus myrianthus Becc.
241. Calamus nagbettai R.R.Fernandez & Dey
242. Calamus nambariensis Becc.
243. Calamus nanduensis W.J.Baker & J.Dransf.
244. Calamus nannostachys Burret
245. Calamus nanodendron J.Dransf.
246. Calamus nematospadix Becc.
247. Calamus nicobaricus Becc.
248. Calamus nielsenii J.Dransf.
249. Calamus nitidus Mart.
250. Calamus notabilis A.J.Hend.
251. Calamus novae-georgii W.J.Baker & J.Dransf.
252. Calamus nudus W.J.Baker & S.Venter
253. Calamus nuichuaensis A.J.Hend., N.K.Ban & N.Q.Dung
254. Calamus nuralievii A.J.Hend. & N.Q.Dung
255. Calamus obiensis A.J.Hend.
256. Calamus oblatus (J.Dransf.) W.J.Baker
257. Calamus oblongus Reinw. ex Blume
258. Calamus occidentalis Witono & J.Dransf.
259. Calamus ochrolepis (Becc.) W.J.Baker
260. Calamus ocreatus (A.J.Hend. & N.Q.Dung) W.J.Baker
261. Calamus oligostachys T.Evans, Sengdala, Viengkham, Thamm. & J.Dransf.
262. Calamus optimus Becc.
263. Calamus oresbiopsis A.J.Hend.
264. Calamus oresbius W.J.Baker & J.Dransf.
265. Calamus ornatus Blume
266. Calamus ovoideus Thwaites ex Trimen
267. Calamus oxleyanus Teijsm. & Binn. ex Miq.
268. Calamus oxleyoides A.J.Hend.
269. Calamus oxycarpus Becc.
270. Calamus oxycoccus W.J.Baker
271. Calamus pachystemonus Thwaites
272. Calamus padangensis Furtado
273. Calamus pahangensis A.J.Hend.
274. Calamus pandanosmus Furtado
275. Calamus papuanus Becc.
276. Calamus parutan Fernando
277. Calamus parvulus A.J.Hend. & N.Q.Dung
278. Calamus paspalanthus Becc.
279. Calamus paulii J.Dransf.
280. Calamus pedicellaris (Becc.) W.J.Baker
281. Calamus pedicellatus Becc. ex K.Heyne
282. Calamus penicillatus Roxb.
283. Calamus perakensis Becc.
284. Calamus peregrinus Furtado
285. Calamus periacanthus (Miq.) Miq.
286. Calamus pertenuis A.J.Hend. & Rustiami
287. Calamus pholidostachys J.Dransf. & W.J.Baker
288. Calamus pilosellus Becc.
289. Calamus pilosissimus Becc.
290. Calamus platyspathus Mart. ex Kunth
291. Calamus plicatus Blume
292. Calamus poensis Becc.
293. Calamus pogonacanthus Becc. ex H.J.P.Winkl.
294. Calamus pogonotium W.J.Baker
295. Calamus poilanei Conrard
296. Calamus politus (Fernando) W.J.Baker
297. Calamus polycladus Burret
298. Calamus posoanus Rustiami
299. Calamus powlingii A.J.Hend.
300. Calamus praetermissus J.Dransf.
301. Calamus propinquus (Becc.) A.J.Hend.
302. Calamus pseudoconcolor (J.Dransf.) W.J.Baker
303. Calamus pseudomollis Becc.
304. Calamus pseudotenuis Becc.
305. Calamus pseudozebrinus Burret
306. Calamus psilocladus J.Dransf.
307. Calamus pycnocarpus (Furtado) J.Dransf.
308. Calamus pygmaeus Becc.
309. Calamus quangngaiensis A.J.Hend. & N.Q.Dung
310. Calamus radiatus Thwaites
311. Calamus radicalis H.Wendl. & Drude
312. Calamus radulosus Becc.
313. Calamus reinwardtii Mart.
314. Calamus reticulatus Burret
315. Calamus retroflexus J.Dransf. & W.J.Baker
316. Calamus rhabdocladus Burret
317. Calamus rheedei Griff.
318. Calamus rhomboideus Blume
319. Calamus rhytidomus Becc.
320. Calamus ridleyanus Becc.
321. Calamus robinsonianus Becc.
322. Calamus rotang L.
323. Calamus ruber Reinw. ex Mart.
324. Calamus rudentum Lour.
325. Calamus rugosus Becc.
326. Calamus rumphii Blume
327. Calamus ruptilis (Becc.) W.J.Baker
328. Calamus ruptiloides A.J.Hend.
329. Calamus ruvidus Becc.
330. Calamus sabalensis J.Dransf.
331. Calamus salicifolius Becc.
332. Calamus saltuensis A.J.Hend.
333. Calamus samian Becc.
334. Calamus sandsii Rustiami
335. Calamus sarawakensis Becc.
336. Calamus sashae J.Dransf. & W.J.Baker
337. Calamus scabridulus Becc.
338. Calamus scabrispathus Becc.
339. Calamus scapiger (Becc.) W.J.Baker
340. Calamus schistoacanthus Blume
341. Calamus schlechteri (Burret) W.J.Baker
342. Calamus schlechterianus Becc.
343. Calamus scipionum Lour.
344. Calamus scleracanthus Becc. ex K.Heyne
345. Calamus sedens J.Dransf.
346. Calamus senalingensis J.Dransf.
347. Calamus septimus A.J.Hend. & Rustiami
348. Calamus seriatus A.J.Hend. & N.Q.Dung
349. Calamus seropakensis A.J.Hend.
350. Calamus serpentinus (J.Dransf.) W.J.Baker
351. Calamus serrulatus Becc.
352. Calamus sessiliflorus A.J.Hend. & Rustiami
353. Calamus simplex Becc.
354. Calamus siphonospathus Mart. ex Walp.
355. Calamus smitinandii (J.Dransf.) A.J.Hend.
356. Calamus sordidus J.Dransf.
357. Calamus spanostachys W.J.Baker & J.Dransf.
358. Calamus sparsiflorus (Becc.) W.J.Baker
359. Calamus spectatissimus Furtado
360. Calamus spiculifer J.Dransf. & W.J.Baker
361. Calamus spinosus A.J.Hend., Pitopang & Moh.Iqbal
362. Calamus spinulinervis Becc.
363. Calamus spiralis A.J.Hend., N.K.Ban & N.Q.Dung
364. Calamus suaveolens W.J.Baker & J.Dransf.
365. Calamus subangulatus Miq.
366. Calamus subsolanus A.J.Hend. & Rustiami
367. Calamus sulawesiensis A.J.Hend.
368. Calamus sumatranus (Becc.) A.J.Hend.
369. Calamus superciliatus W.J.Baker & J.Dransf.
370. Calamus susukensis A.J.Hend. & Rustiami
371. Calamus symphysipus Mart. ex Walp.
372. Calamus tadulakoensis A.J.Hend., Moh.Iqbal, Rusydi & Pitopang
373. Calamus tambingensis A.J.Hend., Pitopang & Moh.Iqbal
374. Calamus tapanensis A.J.Hend.
375. Calamus temii T.Evans
376. Calamus tenompokensis Furtado
377. Calamus tenuis Roxb.
378. Calamus tetradactyloides Burret
379. Calamus tetradactylus Hance
380. Calamus thwaitesii Becc.
381. Calamus thysanolepis Hance
382. Calamus timorensis Becc.
383. Calamus trachycoleus Becc.
384. Calamus trigynus A.J.Hend., Pitopang & Moh.Iqbal
385. Calamus tumidus Furtado
386. Calamus unifarius H.Wendl.
387. Calamus unijugus (J.Dransf.) W.J.Baker
388. Calamus ursinus (Becc.) W.J.Baker
389. Calamus usitatus Blanco
390. Calamus validus W.J.Baker
391. Calamus vattayila Renuka
392. Calamus velutinus A.J.Hend. & N.Q.Dung
393. Calamus verticillaris Griff.
394. Calamus vestitus Becc.
395. Calamus vidalianus Becc.
396. Calamus viminalis Willd.
397. Calamus vinaceus A.J.Hend., Pitopang & Moh.Iqbal
398. Calamus vinosus Becc.
399. Calamus viridis A.J.Hend., Pitopang & Moh.Iqbal
400. Calamus vitiensis Warb. ex Becc.
401. Calamus walkeri Hance
402. Calamus wanggaii W.J.Baker & J.Dransf.
403. Calamus warayanus Adorador & Fernando
404. Calamus warburgii K.Schum.
405. Calamus wedaensis A.J.Hend.
406. Calamus whitmorei J.Dransf.
407. Calamus wightii Griff.
408. Calamus womersleyi J.Dransf. & W.J.Baker
409. Calamus wuliangshanensis San Y.Chen, K.L.Wang & S.J.Pei
410. Calamus yentuensis A.J.Hend. & N.Q.Dung
411. Calamus zebrinus Becc.
412. Calamus zeylanicus Becc.
413. Calamus zieckii Fernando
414. Calamus zollingeri Becc.
415. Calamus zonatus Becc.

Uses

The stems may be harvested for their cores, which are used for everything from canes to furniture.

References

1. "Calamus L." Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 9 April 2022.
2. "World Checklist of Selected Plant Families: Royal Botanic Gardens, Kew". apps.kew.org. Retrieved 14 March 2018.
3. Govaerts, Rafaël; Dransfield, John (2005). World Checklist of Palms. Royal Botanic Gardens Kew. ISBN 1842460935.
4. "Calamus". Flora of China. Retrieved 9 April 2022.
5. Kurian, Binoy; Hemanthakumar, A.S.; Jacob, Joemon; Noushad, Muhammad Ali; Sabu, K. K. (2018). "Rapidly evolving sex-specific sequences in Calamus travancoricus Bedd. ex. Becc. and Calamus nagbettai R.R.Fernald & Dey". Tree Genetics & Genomes. 14. doi:10.1007/s11295-017-1220-2. S2CID 3299464.
6. Dictionary of Flowering Plants and Ferns - JC Willis (1960)
7. Dransfield 1978.
8. Gibbs A; Green C; Doctor VM. (1983). "Isolation and anticoagulant properties of polysaccharides of Typha Augustata and Daemonorops species". Thromb. Res. 32 (2): 97–108. doi:10.1016/0049-3848(83)90021-X. PMID 6658717.

Bibliography

• Tomlinson, P. B.; Zimmerman, Martin, eds. (1978). Tropical Trees as Living Systems (Proceedings of the fourth Cabot Symposium held at Harvard Forest, Petersham Massachusetts on April 26-30, 1976). Cambridge University Press. ISBN 978-0-521-14247-2.
• Dransfield, John (10 June 2010). Growth forms of rain forest palms. pp. 247–268. ISBN 9780521142472., in Tomlinson & Zimmerman (1978)

External links

•   Media related to Calamus (Arecaceae) at Wikimedia Commons
• Calamus deerratus in West African plants – A Photo Guide.