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Ornithocheiroidea

Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with fossil remains found all over the world except Antarctica.

Ornithocheiroids
Temporal range: Early - Late Cretaceous,
~145–66 Ma Possible Late Jurassic record[1]
Restored skeleton of Thalassodromeus sethi, with Anhanguera behind
Skeletal cast of Tropeognathus mesembrinus in the National Museum of Brazil
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Infraorder: Eupterodactyloidea
Clade: Ornithocheiroidea
Seeley, 1870
Subgroups

Ornithocheiroids were the most advanced group of pterosaurs, as the group includes the clade Azhdarchoidea, of which its members lived until the Maastrichtian stage of the Late Cretaceous, around 66 million years ago. Notable pterosaurs from this group include the pteranodontians Pteranodon and Nyctosaurus, the ornithocheirid Ornithocheirus, the anhanguerid Tropeognathus, as well as the azhdarchids Hatzegopteryx and Quetzalcoatlus.

Classification edit

The name Ornithocheiroidea was originally defined as an apomorphy-based taxon by Christopher Bennett in 1994. It was given a relationship-based definition in 2003 by Alexander Kellner, who defined it as the least inclusive clade containing Anhanguera blittersdorffi, Pteranodon longiceps, Dsungaripterus weii, and Quetzalcoatlus northropi.[2] Later that year, David Unwin suggested a more restrictive definition, in which the clade only contains Pteranodon longiceps, Istiodactylus latidens, and their descentants.[3] Brian Andres (2008, 2010, 2014) in his analyses, defined Ornithocheiroidea using the definition of Kellner (2003) to avoid confusion with similarly-defined groups, like Pteranodontoidea.[4][5]

Below is a cladogram showing the results of a phylogenetic analysis presented by Longrich and colleagues in 2018. They found Ornithocheiroidea to consist of the clades Pteranodontoidea and Azhdarchoidea, as well as the genus Piksi.[6]

Eupterodactyloidea

Haopterus gracilis

Ornithocheiroidea

Piksi barbarulna

Pteranodontoidea
Azhdarchoidea
Tapejaromorpha
Neoazhdarchia

In 2019, a phylogenetic analysis conducted by Kellner and colleagues had recovered Ornithocheiroidea as the sister taxon of the Archaeopterodactyloidea, and consisting of the clades Tapejaroidea and Pteranodontoidea.[7] Several recent studies have followed this or a similar concept.[8][9][10] The cladogram of the analysis by Kellner and colleagues is presented below:

References edit

  1. ^ Unwin, David M.; Heinrich, Wolf-Dieter (1999). "On a pterosaur jaw from the Upper Jurassic of Tendaguru (Tanzania)". Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaftliche Reihe. 2: 121–134.
  2. ^ Kellner, A. W. A., (2003): Pterosaur phylogeny and comments on the evolutionary history of the group. pp. 105-137. — in Buffetaut, E. & Mazin, J.-M., (eds.): Evolution and Palaeobiology of Pterosaurs. Geological Society of London, Special Publications 217, London, 1-347
  3. ^ Unwin, D. M., (2003): On the phylogeny and evolutionary history of pterosaurs. pp. 139-190. — in Buffetaut, E. & Mazin, J.-M., (eds.): Evolution and Palaeobiology of Pterosaurs. Geological Society of London, Special Publications 217, London, 1-347 doi:10.1144/GSL.SP.2003.217.01.11
  4. ^ Andres, Brian Blake (2014). Systematics of the Pterosauria. Yale University. p. 366. A preview that shows the cladogram without clade names
  5. ^ Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24 (9): 1011–6. doi:10.1016/j.cub.2014.03.030. PMID 24768054.
  6. ^ Longrich, N.R., Martill, D.M., and Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS Biology, 16(3): e2001663. doi:10.1371/journal.pbio.2001663
  7. ^ Kellner, Alexander W. A.; Weinschütz, Luiz C.; Holgado, Borja; Bantim, Renan A. M.; Sayão, Juliana M. (August 19, 2019). "A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert". Anais da Academia Brasileira de Ciências. 91 (suppl 2): e20190768. doi:10.1590/0001-3765201920190768. ISSN 0001-3765. PMID 31432888.
  8. ^ Kellner, Alexander W. A.; Caldwell, Michael W.; Holgado, Borja; Vecchia, Fabio M. Dalla; Nohra, Roy; Sayão, Juliana M.; Currie, Philip J. (2019). "First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity". Scientific Reports. 9 (1): 17875. Bibcode:2019NatSR...917875K. doi:10.1038/s41598-019-54042-z. PMC 6884559. PMID 31784545.
  9. ^ Borja Holgado, Rodrigo V. Pêgas, José Ignacio Canudo, Josep Fortuny, Taissa Rodrigues, Julio Company & Alexander W.A. Kellner, 2019, "On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria", Scientific Reports 9: 4940 doi:10.1038/s41598-019-41280-4
  10. ^ Jiang, Shun-Xing; Zhang, Xin-Jun; Cheng, Xin; Wang, Xiao-Lin (2020). "A new pteranodontoid pterosaur forelimb from the upper Yixian Formation, with a revision of Yixianopterus jingangshanensis" (PDF). Vertebrata PalAsiatica. doi:10.19615/j.cnki.1000-3118.201124.

ornithocheiroidea, ornithocheiroids, group, pterosaurs, within, extinct, suborder, pterodactyloidea, they, were, typically, large, pterosaurs, that, lived, from, early, late, cretaceous, periods, valanginian, maastrichtian, stages, with, fossil, remains, found. Ornithocheiroidea or ornithocheiroids is a group of pterosaurs within the extinct suborder Pterodactyloidea They were typically large pterosaurs that lived from the Early to Late Cretaceous periods Valanginian to Maastrichtian stages with fossil remains found all over the world except Antarctica OrnithocheiroidsTemporal range Early Late Cretaceous 145 66 Ma PreꞒ Ꞓ O S D C P T J K Pg N Possible Late Jurassic record 1 Restored skeleton of Thalassodromeus sethi with Anhanguera behind Skeletal cast of Tropeognathus mesembrinus in the National Museum of Brazil Scientific classification Domain Eukaryota Kingdom Animalia Phylum Chordata Order Pterosauria Suborder Pterodactyloidea Infraorder Eupterodactyloidea Clade OrnithocheiroideaSeeley 1870 Subgroups Piksi Pteranodontoidea Tapejaroidea Ornithocheiroids were the most advanced group of pterosaurs as the group includes the clade Azhdarchoidea of which its members lived until the Maastrichtian stage of the Late Cretaceous around 66 million years ago Notable pterosaurs from this group include the pteranodontians Pteranodon and Nyctosaurus the ornithocheirid Ornithocheirus the anhanguerid Tropeognathus as well as the azhdarchids Hatzegopteryx and Quetzalcoatlus Classification editThe name Ornithocheiroidea was originally defined as an apomorphy based taxon by Christopher Bennett in 1994 It was given a relationship based definition in 2003 by Alexander Kellner who defined it as the least inclusive clade containing Anhanguera blittersdorffi Pteranodon longiceps Dsungaripterus weii and Quetzalcoatlus northropi 2 Later that year David Unwin suggested a more restrictive definition in which the clade only contains Pteranodon longiceps Istiodactylus latidens and their descentants 3 Brian Andres 2008 2010 2014 in his analyses defined Ornithocheiroidea using the definition of Kellner 2003 to avoid confusion with similarly defined groups like Pteranodontoidea 4 5 Below is a cladogram showing the results of a phylogenetic analysis presented by Longrich and colleagues in 2018 They found Ornithocheiroidea to consist of the clades Pteranodontoidea and Azhdarchoidea as well as the genus Piksi 6 Eupterodactyloidea Haopterus gracilis Ornithocheiroidea Piksi barbarulna Pteranodontoidea Pteranodontia Pteranodontidae Tethydraco regalis Pteranodon longiceps nbsp Pteranodon sternbergi Nyctosauridae Alamodactylus byrdi Volgadraco bogolubovi Cretornis hlavaci Alcione elainus Simurghia robusta Muzquizopteryx coahuilensis Barbaridactylus grandis Nyctosaurus lamegoi Nyctosaurus nanus Nyctosaurus gracilis nbsp Ornithocheiromorpha Hongshanopterus lacustris Lonchodectidae Lonchodraco giganteus Lonchodectes compressirostris Boreopteridae Boreopterus cuiae Zhenyuanopterus longirostris Lanceodontia Istiodactylidae Nurhachius ignaciobritoi Liaoxipterus brachyognathus Istiodactylus sinensis Istiodactylus latidens Aetodactylus halli Cimoliopterus dunni Cimoliopterus cuvieri Anhangueria Guidraco Ludodactylus nbsp Cearadactylus Brasileodactylus Ornithocheirae Anhangueridae nbsp Ornithocheiridae Azhdarchoidea Tapejaromorpha Bennettazhia oregonensis Tapejaridae Tapejarinae Tapejara wellnhoferi Europejara olcadesorum Vectidraco daisymorrisae Caiuajara dobruskii Tupandactylus navigans Tupandactylus imperator nbsp Bakonydraco galaczi nbsp Huaxiapterus benxiensis Huaxiapterus corollatus Eopteranodon lii Huaxiapterus jii Sinopterus dongi Neoazhdarchia Dsungaripteromorpha Dsungaripteridae Dsungaripterinae Dsungaripterus weii Domeykodactylus ceciliae nbsp Noripterinae Noripterus parvus Noripterus complicidens Thalassodromidae Tupuxuara longicristatus nbsp Tupuxuara leonardii Thalassodromeus sethi Alanqa saharica Aerotitan sudamericanus nbsp Neopterodactyloidea Chaoyangopteridae Microtuban altivolans Chaoyangopterinae Shenzhoupterus Chaoyangopterus Jidapterus Eoazhdarcho liaoxiensis Radiodactylus langstoni Azhdarchidae Montanazhdarcho minor Azhdarcho lancicollis Phosphatodraco mauritanicus nbsp Aralazhdarcho bostobensis Eurazhdarcho langendorfensis aff Quetzalcoatlus Zhejiangopterus linhaiensis Arambourgiania philadelphiae Hatzegopteryx thambema nbsp Quetzalcoatlus spp nbsp In 2019 a phylogenetic analysis conducted by Kellner and colleagues had recovered Ornithocheiroidea as the sister taxon of the Archaeopterodactyloidea and consisting of the clades Tapejaroidea and Pteranodontoidea 7 Several recent studies have followed this or a similar concept 8 9 10 The cladogram of the analysis by Kellner and colleagues is presented below Pterodactyloidea Archaeopterodactyloidea nbsp Ornithocheiroidea Pteranodontoidea Pteranodontia Pteranodontidae Pteranodon longiceps nbsp Tethydraco regalis Nyctosauridae Nyctosaurus gracilis nbsp Muzquizopteryx coahuilensis Lanceodontia Hongshanopterus lacustris Istiodactylidae Nurhachius ignaciobritoi Istiodactylinae Istiodactylus latidens Istiodactylus sinensis Haopterus gracilis Lonchodraco giganteus Ikrandraco avatar Ornithocheirae Ornithocheirus simus nbsp Cimoliopterus cuvieri Anhangueria Camposipterus sedgwickii Hamipteridae Hamipterus tianshanensis Iberodactylus andreui Anhangueridae Tropeognathus mesembrinus Coloborhynchinae Coloborhynchus clavirostris Uktenadactylus wadleighi Anhanguerinae Caulkicephalus trimicrodon Guidraco venator Ludodactylus sibbicki nbsp Liaoningopterus gui nbsp Cearadactylus atrox Maaradactylus kellneri AMNH 22555 Anhanguera blittersdorffi Anhanguera piscator Tapejaroidea Dsungaripteridae Dsungaripterus weii Noripterus parvus Azhdarchoidea Azhdarchidae Azhdarcho lancicollis Quetzalcoatlus sp nbsp Zhejiangopterus linhaiensis Chaoyangopteridae Chaoyangopterus zhangi Jidapterus edentus Shenzhoupterus chaoyangensis Tapejaromorpha Keresdrakon vilsoni Tapejaridae Thalassodrominae Thalassodromeus sethi Tupuxuara leonardii Tapejarinae Caupedactylus ybaka Aymberedactylus cearensis Eopteranodon lii Huaxiapterus benxiensis Huaxiapterus corollatus Sinopterus dongi Tapejarini Europejara olcadesorum Caiuajara dobruskii Tapejara wellnhoferi Tupandactylus imperator nbsp References edit Unwin David M Heinrich Wolf Dieter 1999 On a pterosaur jaw from the Upper Jurassic of Tendaguru Tanzania Mitteilungen aus dem Museum fur Naturkunde in Berlin Geowissenschaftliche Reihe 2 121 134 Kellner A W A 2003 Pterosaur phylogeny and comments on the evolutionary history of the group pp 105 137 in Buffetaut E amp Mazin J M eds Evolution and Palaeobiology of Pterosaurs Geological Society of London Special Publications 217 London 1 347 Unwin D M 2003 On the phylogeny and evolutionary history of pterosaurs pp 139 190 in Buffetaut E amp Mazin J M eds Evolution and Palaeobiology of Pterosaurs Geological Society of London Special Publications 217 London 1 347 doi 10 1144 GSL SP 2003 217 01 11 Andres Brian Blake 2014 Systematics of the Pterosauria Yale University p 366 A preview that shows the cladogram without clade names Andres B Clark J Xu X 2014 The Earliest Pterodactyloid and the Origin of the Group Current Biology 24 9 1011 6 doi 10 1016 j cub 2014 03 030 PMID 24768054 Longrich N R Martill D M and Andres B 2018 Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous Paleogene boundary PLoS Biology 16 3 e2001663 doi 10 1371 journal pbio 2001663 Kellner Alexander W A Weinschutz Luiz C Holgado Borja Bantim Renan A M Sayao Juliana M August 19 2019 A new toothless pterosaur Pterodactyloidea from Southern Brazil with insights into the paleoecology of a Cretaceous desert Anais da Academia Brasileira de Ciencias 91 suppl 2 e20190768 doi 10 1590 0001 3765201920190768 ISSN 0001 3765 PMID 31432888 Kellner Alexander W A Caldwell Michael W Holgado Borja Vecchia Fabio M Dalla Nohra Roy Sayao Juliana M Currie Philip J 2019 First complete pterosaur from the Afro Arabian continent insight into pterodactyloid diversity Scientific Reports 9 1 17875 Bibcode 2019NatSR 917875K doi 10 1038 s41598 019 54042 z PMC 6884559 PMID 31784545 Borja Holgado Rodrigo V Pegas Jose Ignacio Canudo Josep Fortuny Taissa Rodrigues Julio Company amp Alexander W A Kellner 2019 On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria Scientific Reports 9 4940 doi 10 1038 s41598 019 41280 4 Jiang Shun Xing Zhang Xin Jun Cheng Xin Wang Xiao Lin 2020 A new pteranodontoid pterosaur forelimb from the upper Yixian Formation with a revision of Yixianopterus jingangshanensis PDF Vertebrata PalAsiatica doi 10 19615 j cnki 1000 3118 201124 Retrieved from https en wikipedia org w index php title Ornithocheiroidea amp oldid 1179672077, wikipedia, wiki, book, books, library,

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