The Hydroptilidae are a large family of caddisflies (Trichoptera) with a worldwide distribution. They are commonly known as microcaddisflies or purse-case caddisflies, in reference to two characteristic traits of this family: Hydroptilidae are much smaller than other caddisflies, rarely exceeding 5 mm (0.20 in) in length. Their larvae do not build a protective case until the final instar of their growth. At that time however, they build a typically Purse-shaped case, either portable or stuck to the substrate, in which the larva finishes growth and pupates.[1]
Their systematic placement among the caddisflies is still disputed. They are traditionally placed in the suborderSpicipalpia, which do not seem to be a natural monophyletic group, but rather an evolutionary grade of moderately advanced caddisflies. Some authors downrank the Spicipalpia to a superfamily of the more basalAnnulipalpia and call them Rhyacophiloidea (which otherwise refers to a subfamily of suborder Spicipalpia), but recent studies generally rejected this view.[1]
More often, the Hydroptilidae are placed in a monotypic superfamily Hydroptiloidea, either in the Spicipalpia or – probably more appropriately considering the present state of caddisfly phylogeny – incertae sedis in the Trichoptera. It may be that the Glossosomatoidea are particularly closely related to the Hydroptilidae; together they might even be closer to the most advanced caddisflies (the tube case caddisflies, Integripalpia) than any other living caddisfly. But this view is almost as disputed as including the Spicipalpia in the Annulipalpia. In any case, were Glossosomatoidea and Hydroptiloidea to be merged into a single superfamily, the older name Hydroptiloidea would apply for the combined group.[1]
The peculiarly apomorphicgeneraPalaeagapetus and Ptilocolepus have been separated as subfamily Ptilocolepinae, while all other genera form the Hydroptilinae family with its multiple tribes. It is far from certain that the Ptilocolepinae are the living fossils such an arrangement would imply them to be.
Generaedit
Around 70 genera with at least 1,700 species have been described from this family:[2]
†Novajerseya Botosaneanu, Johnson & P. R. Dillon, 1998
Orphninotrichia Mosely, 1934
Referencesedit
^ abcRalph W. Holzenthal; Roger J. Blahnik; Aysha Prather; Karl Kjer (January 12, 2010). "Trichoptera". Tree of Life Web Project. Retrieved April 4, 2010.
^. Trichoptera World Checklist. Clemson University. Archived from the original on June 10, 2010. Retrieved April 4, 2010.
External linksedit
January 01, 1970
hydroptilidae, large, family, caddisflies, trichoptera, with, worldwide, distribution, they, commonly, known, microcaddisflies, purse, case, caddisflies, reference, characteristic, traits, this, family, much, smaller, than, other, caddisflies, rarely, exceedin. The Hydroptilidae are a large family of caddisflies Trichoptera with a worldwide distribution They are commonly known as microcaddisflies or purse case caddisflies in reference to two characteristic traits of this family Hydroptilidae are much smaller than other caddisflies rarely exceeding 5 mm 0 20 in in length Their larvae do not build a protective case until the final instar of their growth At that time however they build a typically Purse shaped case either portable or stuck to the substrate in which the larva finishes growth and pupates 1 Hydroptilidae Microcaddisfly from Pryor Creek Oklahoma Scientific classification Domain Eukaryota Kingdom Animalia Phylum Arthropoda Class Insecta Order Trichoptera Suborder Spicipalpia Superfamily Hydroptiloidea Family HydroptilidaeStephens 1836 Subfamilies Hydroptilinae Stephens 1836 Leucotrichiinae Flint 1970 Neotrichiinae Ross 1956 Ochrotrichiinae Marshall 1979 Orthotrichiinae Nielsen 1948 Stactobiinae Botosaneanu 1956 Hydroptila larva Contents 1 Systematics and taxonomy 2 Genera 3 References 4 External linksSystematics and taxonomy editTheir systematic placement among the caddisflies is still disputed They are traditionally placed in the suborder Spicipalpia which do not seem to be a natural monophyletic group but rather an evolutionary grade of moderately advanced caddisflies Some authors downrank the Spicipalpia to a superfamily of the more basal Annulipalpia and call them Rhyacophiloidea which otherwise refers to a subfamily of suborder Spicipalpia but recent studies generally rejected this view 1 More often the Hydroptilidae are placed in a monotypic superfamily Hydroptiloidea either in the Spicipalpia or probably more appropriately considering the present state of caddisfly phylogeny incertae sedis in the Trichoptera It may be that the Glossosomatoidea are particularly closely related to the Hydroptilidae together they might even be closer to the most advanced caddisflies the tube case caddisflies Integripalpia than any other living caddisfly But this view is almost as disputed as including the Spicipalpia in the Annulipalpia In any case were Glossosomatoidea and Hydroptiloidea to be merged into a single superfamily the older name Hydroptiloidea would apply for the combined group 1 The peculiarly apomorphic genera Palaeagapetus and Ptilocolepus have been separated as subfamily Ptilocolepinae while all other genera form the Hydroptilinae family with its multiple tribes It is far from certain that the Ptilocolepinae are the living fossils such an arrangement would imply them to be Genera editAround 70 genera with at least 1 700 species have been described from this family 2 Hydroptilinae Stephens 1836 Acanthotrichia Wells 1982 Acritoptila Wells 1982 Agraylea Curtis 1834 Allotrichia McLachlan 1880 Austratrichia Wells 1982 Cyclopsiella Kjaerandsen 1997 Dhatrichia Mosely 1948 Hellyethira Neboiss 1977 Hydroptila Dalman 1819 Hydroptilina Martynov 1934 Jabitrichia Wells 1990 Microptila Ris 1897 Missitrichia Wells 1991 Mulgravia Wells 1982 Oxyethira Eaton 1873 Paroxyethira Mosely 1924 Paucicalcaria Mathis amp Bowles 1989 Tangatrichia Wells amp Andersen 1995 Tricholeiochiton Kloet amp Hincks 1944 Ugandatrichia Mosely 1939 Vietrichia Olah 1989 Wlitrichia Kjaerandsen 1997 Xuthotrichia Mosely 1934 Leucotrichiinae Flint 1970 Abtrichia Mosely 1939 Acostatrichia Mosely 1939 Alisotrichia Flint 1964 Anchitrichia Flint 1970 Ascotrichia Flint 1983 Betrichia Mosely 1939 Celaenotrichia Mosely 1934 Cerasmatrichia Flint Harris amp Botosaneanu 1994 Ceratotrichia Flint 1992 Costatrichia Mosely 1937 Eutonella Mueller 1921 Leucotrichia Mosely 1934 Mejicanotrichia Harris amp Holzenthal 1997 Peltopsyche Mueller 1879 Scelobotrichia Harris amp Bueno Soria 1993 Zumatrichia Mosely 1937 Neotrichiinae Ross 1956 Kumanskiella Harris amp Flint 1992 Mayatrichia Mosely 1937 Neotrichia Morton 1905 Taraxitrichia Flint amp Harris 1991 Ochrotrichiinae Marshall 1979 Angrisanoia Ozdikmen 2008 Caledonotrichia Sykora 1967 Maydenoptila Neboiss 1977 Metrichia Ross 1938 Ochrotrichia Mosely 1934 Nothotrichia Flint 1967 Ragatrichia Olah amp Johanson 2011 Rhyacopsyche Mueller 1879 Orthotrichiinae Nielsen 1948 Ithytrichia Eaton 1873 Orthotrichia Eaton 1873 Stactobiinae Botosaneanu 1956 Bredinia Flint 1968 Byrsopteryx Flint 1981 Catoxyethira Ulmer 1912 Chrysotrichia Schmid 1958 Flintiella Angrisano 1995 Niuginitrichia Wells 1990 Orinocotrichia Harris Flint amp Holzenthal 2002 Parastactobia Schmid 1958 Plethus Hagen 1887 Scelotrichia Ulmer 1951 Stactobia McLachlan 1880 Stactobiella Martynov 1924 Tizatetrichia Harris Flint amp Holzenthal 2002 incertae sedis Burminoptila Botosaneanu 1981 Dibusa Ross 1939 Dicaminus Mueller 1879 Electrotrichia Ulmer 1912 Macrostactobia Schmid 1958 Novajerseya Botosaneanu Johnson amp P R Dillon 1998 Orphninotrichia Mosely 1934References edit a b c Ralph W Holzenthal Roger J Blahnik Aysha Prather Karl Kjer January 12 2010 Trichoptera Tree of Life Web Project Retrieved April 4 2010 Classification Hierarchy Trichoptera World Checklist Clemson University Archived from the original on June 10 2010 Retrieved April 4 2010 External links editTrichoptera World Checklist Retrieved from https en wikipedia org w index php title Hydroptilidae amp oldid 1180305417, wikipedia, wiki, book, books, library,
