fbpx
Wikipedia

Dinosaur size

Size is an important aspect of dinosaur paleontology, of interest to both the general public and professional scientists. Dinosaurs show some of the most extreme variations in size of any land animal group, ranging from tiny hummingbirds, which can weigh as little as two grams, to the extinct titanosaurs, which could weigh as much as 50–100 t (55–110 short tons).

Scale diagram comparing a human and the longest-known dinosaurs of five major clades
An adult male bee hummingbird, the smallest known and the smallest living dinosaur

The latest evidence suggests that dinosaurs' average size varied through the Triassic, early Jurassic, late Jurassic and Cretaceous periods, and dinosaurs probably only became widespread during the early or mid Jurassic.[1] Predatory theropod dinosaurs, which occupied most terrestrial carnivore niches during the Mesozoic, most often fall into the 100–1,000 kg (220–2,200 lb) category when sorted by estimated weight into categories based on order of magnitude, whereas recent predatory carnivoran mammals peak in the range of 10–100 kg (22–220 lb).[2] The mode of Mesozoic dinosaur body masses is between one and ten metric tonnes.[3] This contrasts sharply with the size of Cenozoic mammals, estimated by the National Museum of Natural History as about 2 to 5 kg (4.4 to 11.0 lb).[4]

Size estimation

Scientists will probably never be certain of the largest and smallest dinosaurs. This is because only a small fraction of animals ever fossilize, and most of these remains will either never be uncovered, or will be unintentionally destroyed as a result of human activity. Of the specimens that are recovered, few are even relatively complete skeletons, and impressions of skin and other soft tissues are rarely discovered. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species is an inexact art (though governed by some established allometric trends), and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork, and never perfect.[5] Mass estimates for dinosaurs are much more variable than length estimates given the lack of soft tissue preservation in the fossilization process. Modern mass estimation is often done with the laser scan skeleton technique that puts a "virtual" skin over the known or implied skeleton, but the limitations inherent in previous mass estimation techniques remain.[6]

Sauropodomorphs

 
size comparison of selected giant sauropod dinosaurs

Sauropodomorph size is difficult to estimate given their usually fragmentary state of preservation. Sauropods are often preserved without their tails, so the margin of error in overall length estimates is high. Mass is calculated using the cube of the length, so for species in which the length is particularly uncertain, the weight is even more so. Estimates that are particularly uncertain (due to very fragmentary or lost material) are preceded by a question mark. Each number represents the highest estimate of a given research paper. One large sauropod, Maraapunisaurus fragillimus, was based on particularly scant remains that have been lost since their description by paleontologists in 1878. Analysis of the illustrations included in the original report suggested that M. fragillimus may have been the largest land animal of all time, possibly weighing 100–150 t (110–170 short tons) and measuring between 40–60 m (130–200 ft) long.[7][8] One later analysis of the surviving evidence, and the biological plausibility of such a large land animal, suggested that the enormous size of this animal was an over-estimate due partly to typographical errors in the original report.[9] This would later be challenged by a different study, which argued Cope's measurements were genuine and that there was no basis for assuming typographical errors. The study, however, also reclassified the species and correspondingly gave a much lower length estimate of 30.3 metres (99 ft) and a mass of 78.5 t (86.5 short tons).[10] This in itself would later be disputed as being too small for an animal of such size, with some believing it to be even larger at around 35–40 metres (115–131 ft) and weighing around 80–120 t (88–132 short tons).[11]

Another large but even more controversial sauropod is Bruhathkayosaurus which had a calculated weight ranging between 126–220 t (139–243 short tons) and a length of 44.1 m (145 ft)[12][13][14] Although the existence of this sauropod had long been dismissed as a potential fake or a misidentification of a petrified tree trunk, recent photographic evidence emerged, confirming its existence.[15] More recent and reliable estimates in 2023 have rescaled Bruhathkayosaurus to weigh around 110–130 t (120–140 short tons) with its most liberal estimate being 240 t (260 short tons), making it incredibly massive for such an animal.[16] If the upper unlikely size estimates were to be taken at face value, Bruhathkayosaurus would not only be the largest dinosaur to have ever lived, but also the largest animal to have lived, exceeding even the largest blue whale recorded. According to Gregory S. Paul, 'super-sauropods' or 'land-whales' such as Maraapunisaurus, Bruhathkayosaurus and the Broome Titanosaur footprints as he calls them, should not be surprising as sauropods were more heat tolerant and grew rapidly, which allowed them to reach truly titanic sizes that rivalled the largest whales in mass despite the prevalence of air sacs.[16] Other potential factors for such extreme sauropod sizes include increasing bone robustness and load-distributing cartilaginous features to better redistribute and support such massive weights.[16]

Generally, the giant sauropods can be divided into two categories: the shorter but stockier and more massive forms (mainly titanosaurs and some brachiosaurids), and the longer but slenderer and more light-weight forms (mainly diplodocids).

Because different methods of estimation sometimes give conflicting results, mass estimates for sauropods can vary widely causing disagreement among scientists over the accurate number. For example, the titanosaur Dreadnoughtus was originally estimated to weigh 59.3 tonnes by the allometric scaling of limb-bone proportions, whereas more recent estimates, based on three-dimensional reconstructions, yield a much smaller figure of 22.1–38.2 tonnes.[17]

 
Reconstructed skeleton of the titanosaur Argentinosaurus huinculensis, often considered the largest-known dinosaur

The sauropods were the longest and heaviest dinosaurs. For much of the dinosaur era, the smallest sauropods were larger than almost anything else in their habitat, and the largest were an order of magnitude more massive than anything else known to have walked the Earth since. Giant prehistoric mammals such as Paraceratherium and Palaeoloxodon (the largest land mammals ever discovered[18]) were dwarfed by the giant sauropods, and only modern whales approach or surpass them in weight, though they live in the oceans.[19] There are several proposed advantages for the large size of sauropods, including protection from predation, reduction of energy use, and longevity, but it may be that the most important advantage was dietary. Large animals are more efficient at digestion than small animals, because food spends more time in their digestive systems. This also permits them to subsist on food with lower nutritive value than smaller animals. Sauropod remains are mostly found in rock formations interpreted as dry or seasonally dry, and the ability to eat large quantities of low-nutrient browse would have been advantageous in such environments.[20]

One of the tallest and heaviest dinosaurs known from good skeletons is Giraffatitan brancai (previously classified as a species of Brachiosaurus). Its remains were discovered in Tanzania between 1907 and 1912. Bones from several similar-sized individuals were incorporated into the skeleton now mounted and on display at the Museum für Naturkunde Berlin;[21] this mount is 12–13.27 metres (39.4–43.5 ft) tall and 21.8–22.5 metres (72–74 ft) long,[22][23][24] and would have belonged to an animal that weighed between 30,000 to 60,000 kilograms (66,000 to 132,000 lb). One of the longest complete dinosaurs is the 27-metre-long (89 ft) Diplodocus, which was discovered in Wyoming in the United States and displayed in Pittsburgh's Carnegie Natural History Museum in 1907.[25]

There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest herbivorous specimens on record were discovered in the 1970s or later, and include the massive titanosaur Argentinosaurus huinculensis, which is the largest dinosaur known from uncontroversial and relatively substantial evidence, estimated to have been 70–80 t (77–88 short tons) and 36 m (118 ft) long.[26][7] Some of the longest sauropods were those with exceptionally long, whip-like tails, such as the 29–30 m (95–98 ft) Diplodocus hallorum[7][26] (formerly Seismosaurus) and the 39 m Supersaurus.[27]

In 2014, the fossilized remains of a previously unknown species of sauropod were discovered in Argentina.[28] The titanosaur, named Patagotitan mayorum, was estimated to have been around 40 m (130 ft) long weighing around 77 t (85 short tons), larger than any other previously found sauropod. The specimens found were remarkably complete, significantly more so than previous titanosaurs. It since been suggested that Patagotitan was not necessarily larger than Argentinosaurus and Puertasaurus.[29] In 2019, Patagotitan was estimated to have been 31 metres (102 ft) long and about 55 tonnes (121,000 lb) massive.[30]

The largest of non-sauropod sauropodomorphs was the unnamed 16 metres (52 ft) long 10 tonnes (22,000 lb) unnamed Elliot giant.[26] Another large sauropodomorph was Euskelosaurus. It reached 12.2 m (40 ft) in length and 2 t (2.2 short tons) in weight.[31] Yunnanosaurus youngi also reached a length of 13 m (43 ft).[32]

Theropods

 
Size comparison of selected giant theropod dinosaurs

Tyrannosaurus was for many decades the largest and best-known theropod to the general public. Since its discovery, however, a number of other giant carnivorous dinosaurs have been described, including Spinosaurus, Carcharodontosaurus, and Giganotosaurus.[33] These large theropod dinosaurs are estimated to rival or even exceeded Tyrannosaurus rex in size, though more recent studies and reconstructions show that Tyrannosaurus, although shorter, was the bulkier animal overall. Specimens such as Sue and Scotty are both estimated to be the most massive theropods known to science. There is still no clear explanation for exactly why these animals grew so bulky and heavy compared to the land predators that came before and after them.

 
Skeleton of Giganotosaurus, one of the largest theropods known.

The largest extant theropod is the common ostrich, up to 2.74 metres (9 ft 0 in) tall and weighs between 63.5 and 145.15 kilograms (140.0 and 320.0 lb).[34]

The smallest non-avialan theropod known from adult specimens may be Anchiornis huxleyi, at 110 grams (3.9 ounces) in weight and 34 centimetres (13 in) in length,[35] although later study discovered larger specimen reaching 62 centimetres (24 in).[36] However, some studies suggest that Anchiornis was actually an avialan.[37] The smallest dinosaur known from adult specimens which is definitely not an avialan is Parvicursor remotus, at 162 grams (5.7 oz) and measuring 39 centimetres (15 in) long.[38] However, in 2022 its holotype was recognized as a juvenile individual.[39] Among living dinosaurs, the bee hummingbird (Mellisuga helenae) is smallest at 1.9 g (0.067 oz) and 5.5 cm (2.2 in) long.[40] The smallest theropod overall (including avians) is the currently extant Bee Hummingbird at 6.12 cm long and 2.6g for females, and 5.51 cm long and 3.25g for the males.[41]

In the theropod lineage leading to birds, body size shrank continuously over a period of 50 million years, from an average of 163 kilograms (359 lb) down to 0.8 kg (1.8 lb). This was the only dinosaur lineage to get continuously smaller over such an extended time period, and their skeletons developed adaptations at about four times the average rate for dinosaurs.[42][43]

See also

References

  1. ^ Sereno PC (1999). "The evolution of dinosaurs". Science. 284 (5423): 2137–2147. doi:10.1126/science.284.5423.2137. PMID 10381873.
  2. ^ Farlow JA (1993). "On the rareness of big, fierce animals: speculations about the body sizes, population densities, and geographic ranges of predatory mammals and large, carnivorous dinosaurs". In Dodson, Peter; Gingerich, Philip (eds.). Functional Morphology and Evolution. American Journal of Science, Special Volume. Vol. 293-A. pp. 167–199.
  3. ^ Peczkis, J. (1994). "Implications of body-mass estimates for dinosaurs". Journal of Vertebrate Paleontology. 14 (4): 520–33. doi:10.1080/02724634.1995.10011575.
  4. ^ "Anatomy and evolution". National Museum of Natural History. from the original on 2007-11-11. Retrieved 2007-11-21.
  5. ^ Paul, Gregory S. (2010). Princeton Field Guide to Dinosaurs. Princeton University Press. ISBN 978-0-691-13720-9.
  6. ^ Strauss, Bob."Why Were Dinosaurs So Big? The Facts and Theories Behind Dinosaur Gigantism". About Education 2014-10-06 at the Wayback Machine
  7. ^ a b c Paul, Gregory S. (2016). The Princeton Field Guide to Dinosaurs: 2nd Edition. United States of America: Princeton University Press. ISBN 978-0-691-16766-4.
  8. ^ Paul, G.S. (1997). "Dinosaur models: the good, the bad, and using them to estimate the mass of dinosaurs" (PDF). In Wolberg, D.L.; Stump, E.; Rosenberg, G.D. (eds.). DinoFest International Proceedings. Dinofest International. The Academy of Natural Sciences. pp. 129–154. (PDF) from the original on March 4, 2016.
  9. ^ Woodruff, C; Foster, JR (2015). "The fragile legacy of Amphicoelias fragillimus (Dinosauria: Sauropoda; Morrison Formation - Latest Jurassic)". PeerJ PrePrints. doi:10.7287/peerj.preprints.838v1.
  10. ^ Carpenter, Kenneth (2018-10-19). "Maraapunisaurus fragillimus, N.G. (formerly Amphicoelias fragillimus), a basal Rebbachisaurid from the Morrison Formation (Upper Jurassic) of Colorado". Geology of the Intermountain West. 5: 227–244. doi:10.31711/giw.v5.pp227-244. ISSN 2380-7601.
  11. ^ Paul, Gregory S. (2019). "Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals" (PDF). Annals of the Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. S2CID 210840060.
  12. ^ Mortimer, M. (2001), "Re: Bruhathkayosaurus", discussion group, The Dinosaur Mailing List, 19 June 2001. Accessed 23 May 2008.
  13. ^ Mortimer, M. (2004), "Re: Largest Dinosaurs" 2019-09-13 at the Wayback Machine, discussion group, The Dinosaur Mailing List, 7 September 2004. Accessed 23 May 2008.
  14. ^ Mortimer, M. (2001), "Titanosaurs too large?" 2016-03-03 at the Wayback Machine, discussion group, The Dinosaur Mailing List, 12 September 2001. Accessed 23 May 2008.
  15. ^ Pal, Saurabh; Ayyasami, Krishnan (27 June 2022). "The lost titan of Cauvery". Geology Today. 38 (3): 112–116. doi:10.1111/gto.12390. ISSN 0266-6979. S2CID 250056201.
  16. ^ a b c Paul, Gregory S.; Larramendi, Asier (11 April 2023). "Body mass estimate of Bruhathkayosaurus and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales". Lethaia. 56 (2): 1–11. doi:10.18261/let.56.2.5. ISSN 0024-1164. S2CID 259782734.
  17. ^ Bates, Karl T.; Falkingham, Peter L.; Macaulay, Sophie; Brassey, Charlotte; Maidment, Susannah C.R. (2015). "Downsizing a giant: re-evaluating Dreadnoughtus body mass". Biol Lett. 11 (6): 20150215. doi:10.1098/rsbl.2015.0215. PMC 4528471. PMID 26063751.
  18. ^ Larramendi, A. (2016). "Shoulder height, body mass and shape of proboscideans" (PDF). Acta Palaeontologica Polonica. 61. doi:10.4202/app.00136.2014. S2CID 2092950. (PDF) from the original on 2016-08-24.
  19. ^ Sander, P. Martin; Christian, Andreas; Clauss, Marcus; Fechner, Regina; Gee, Carole T.; Griebeler, Eva-Maria; Gunga, Hanns-Christian; Hummel, Jürgen; Mallison, Heinrich; et al. (2011). "Biology of the sauropod dinosaurs: the evolution of gigantism". Biological Reviews. 86 (1): 117–155. doi:10.1111/j.1469-185X.2010.00137.x. PMC 3045712. PMID 21251189.
  20. ^ Carpenter, K. (2006). "Biggest of the big: a critical re-evaluation of the mega-sauropod Amphicoelias fragillimus." In Foster, J.R. and Lucas, S.G., eds., 2006, Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36: 131–138.
  21. ^ Colbert, Edwin Harris (1971). Men and dinosaurs: the search in field and laboratory. Harmondsworth [Eng.]: Penguin. ISBN 978-0-14-021288-4.
  22. ^ Mazzetta, G.V.; et al. (2004). "Giants and Bizarres: Body Size of Some Southern South American Cretaceous Dinosaurs". Historical Biology. 16 (2–4): 1–13. CiteSeerX 10.1.1.694.1650. doi:10.1080/08912960410001715132. S2CID 56028251.
  23. ^ Janensch, W. (1950). "The Skeleton Reconstruction of Brachiosaurus brancai": 97–103. {{cite journal}}: Cite journal requires |journal= (help)
  24. ^ "The World of Dinosaurs". Museum für Naturkunde. from the original on 2018-11-16. Retrieved 2018-11-16.
  25. ^ Lucas, H.; Hecket, H. (2004). "Reappraisal of Seismosaurus, a Late Jurassic Sauropod". Proceeding, Annual Meeting of the Society of Paleontology. 36 (5): 422.
  26. ^ a b c Molina-Pérez, Rubén; Larramendi, Asier (2020). Dinosaur Facts and Figures The Sauropods and Other Sauropodomorphs. Translated by Donaghey, Joan. Illustrated by Andrey Atuchin and Sante Mazzei. Princeton: Princeton University Press. ISBN 978-0-691-20297-6. OCLC 1157079384.
  27. ^ "SVP 2021 Symposium". Society of Vertebrate Paleontology. Retrieved 2023-12-16.
  28. ^ Morgan, James (2014-05-17). "'Biggest dinosaur ever' discovered". BBC News. from the original on 2017-02-18. Retrieved 2017-03-21.
  29. ^ "Don't believe the hype: Patagotitan was not bigger than Argentinosaurus". Sauropod Vertebra Picture of the Week. 2017-08-09. from the original on 2019-04-14. Retrieved 2019-02-19.
  30. ^ Paul, Gregory (December 2019). "Determining the Largest Known Land Animal: A Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals" (PDF). Annals of Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. ISSN 0097-4463. S2CID 210840060.
  31. ^ Patricia Vickers Rich, Thomas Hewitt Rich, Mildred Adams Fenton, Carroll Lane (January 15, 2020). The Fossil Book: A Record of Prehistoric Life. Dover Publications. p. 444. ISBN 9780486838557. Retrieved 2022-08-25.{{cite book}}: CS1 maint: multiple names: authors list (link)
  32. ^ Lu, J., Li, T., Zhong, S., Azuma, Y., Fujita, M., Dong, Z., and Ji, Q. (2007). "New yunnanosaurid dinosaur (Dinosauria, Prosauropoda) from the Middle Jurassic Zhanghe Formation of Yuanmou, Yunnan Province of China". Memoir of the Fukui Prefectural Dinosaur Museum, 6: 1-15.
  33. ^ Therrien, F.; Henderson, D. M. (2007). "My theropod is bigger than yours...or not: estimating body size from skull length in theropods". Journal of Vertebrate Paleontology. 27 (1): 108–115. doi:10.1671/0272-4634(2007)27[108:MTIBTY]2.0.CO;2. S2CID 86025320.
  34. ^ "See what African Wildlife Foundation is doing to protect these iconic flightless birds". 2013-02-25. from the original on 2015-04-29. Retrieved 2015-04-18.
  35. ^ Xu, X., Zhao, Q., Norell, M., Sullivan, C., Hone, D., Erickson, G., Wang, X., Han, F. and Guo, Y. (2009). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin." Chinese Science Bulletin, 6 pages, accepted November 15, 2008.
  36. ^ Pei, Rui; Li, Quanguo; Meng, Qingjin; Norell, Mark A.; Gao, Ke-Qin (2017). "New Specimens ofAnchiornis huxleyi(Theropoda: Paraves) from the Late Jurassic of Northeastern China". Bulletin of the American Museum of Natural History. 411: 1–67. doi:10.1206/0003-0090-411.1.1. ISSN 0003-0090. S2CID 90650697.
  37. ^ Pascal Godefroit; Andrea Cau; Hu Dong-Yu; François Escuillié; Wu Wenhao; Gareth Dyke (2013). "A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds". Nature. 498 (7454): 359–62. Bibcode:2013Natur.498..359G. doi:10.1038/nature12168. PMID 23719374. S2CID 4364892.
  38. ^ Which was the smallest dinosaur? 2011-07-06 at the Wayback Machine Royal Tyrrell Museum. Last accessed 22 September 2022.
  39. ^ Averianov AO, Lopatin AV (2022). "A re-appraisal of Parvicursor remotus from the Late Cretaceous of Mongolia: implications for the phylogeny and taxonomy of alvarezsaurid theropod dinosaurs". Journal of Systematic Palaeontology. 19 (16): 1097–1128. doi:10.1080/14772019.2021.2013965. S2CID 247222017.
  40. ^ Conservation International (Content Partner); Mark McGinley (Topic Editor). 2008. "Biological diversity in the Caribbean Islands." In: Encyclopedia of Earth. Eds. Cutler J. Cleveland (Washington, D.C.: Environmental Information Coalition, National Council for Science and the Environment). [First published in the Encyclopedia of Earth May 3, 2007; Last revised August 22, 2008; Retrieved November 9, 2009]. 2013-05-23 at the Wayback Machine>
  41. ^ Glick, Adrienne Animal Diversity Web, Retrieved 04 October 2023
  42. ^ Borenstein, Seth (July 31, 2014). . AP News. Archived from the original on August 8, 2014. Retrieved August 3, 2014.
  43. ^ Zoe Gough (31 July 2014). "Dinosaurs 'shrank' regularly to become birds". BBC. from the original on 20 February 2015. Retrieved 20 April 2015.

External links

  •   Media related to Dinosauria size comparisons at Wikimedia Commons
  • The Biggest Carnivore: Dinosaur History Rewritten
  • Holtz, Thomas R. Jr.; Rey, Luis V. (2007). Dinosaurs: the most complete, up-to-date encyclopedia for dinosaur lovers of all ages. New York: Random House. ISBN 978-0-375-82419-7. (Dinosaur size#References)
  • "Dinosaur records", Czech article by Vladimír Socha; DinosaurusBlog.com, August 1, 2016

dinosaur, size, this, article, require, cleanup, meet, wikipedia, quality, standards, specific, problem, outdated, information, unreliable, overestimates, please, help, improve, this, article, august, 2022, learn, when, remove, this, template, message, size, i. This article may require cleanup to meet Wikipedia s quality standards The specific problem is Outdated information and unreliable overestimates Please help improve this article if you can August 2022 Learn how and when to remove this template message Size is an important aspect of dinosaur paleontology of interest to both the general public and professional scientists Dinosaurs show some of the most extreme variations in size of any land animal group ranging from tiny hummingbirds which can weigh as little as two grams to the extinct titanosaurs which could weigh as much as 50 100 t 55 110 short tons Scale diagram comparing a human and the longest known dinosaurs of five major cladesAn adult male bee hummingbird the smallest known and the smallest living dinosaurThe latest evidence suggests that dinosaurs average size varied through the Triassic early Jurassic late Jurassic and Cretaceous periods and dinosaurs probably only became widespread during the early or mid Jurassic 1 Predatory theropod dinosaurs which occupied most terrestrial carnivore niches during the Mesozoic most often fall into the 100 1 000 kg 220 2 200 lb category when sorted by estimated weight into categories based on order of magnitude whereas recent predatory carnivoran mammals peak in the range of 10 100 kg 22 220 lb 2 The mode of Mesozoic dinosaur body masses is between one and ten metric tonnes 3 This contrasts sharply with the size of Cenozoic mammals estimated by the National Museum of Natural History as about 2 to 5 kg 4 4 to 11 0 lb 4 Contents 1 Size estimation 2 Sauropodomorphs 3 Theropods 4 See also 5 References 6 External linksSize estimationScientists will probably never be certain of the largest and smallest dinosaurs This is because only a small fraction of animals ever fossilize and most of these remains will either never be uncovered or will be unintentionally destroyed as a result of human activity Of the specimens that are recovered few are even relatively complete skeletons and impressions of skin and other soft tissues are rarely discovered Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar better known species is an inexact art though governed by some established allometric trends and reconstructing the muscles and other organs of the living animal is at best a process of educated guesswork and never perfect 5 Mass estimates for dinosaurs are much more variable than length estimates given the lack of soft tissue preservation in the fossilization process Modern mass estimation is often done with the laser scan skeleton technique that puts a virtual skin over the known or implied skeleton but the limitations inherent in previous mass estimation techniques remain 6 SauropodomorphsMain article Sauropodomorpha nbsp size comparison of selected giant sauropod dinosaursSauropodomorph size is difficult to estimate given their usually fragmentary state of preservation Sauropods are often preserved without their tails so the margin of error in overall length estimates is high Mass is calculated using the cube of the length so for species in which the length is particularly uncertain the weight is even more so Estimates that are particularly uncertain due to very fragmentary or lost material are preceded by a question mark Each number represents the highest estimate of a given research paper One large sauropod Maraapunisaurus fragillimus was based on particularly scant remains that have been lost since their description by paleontologists in 1878 Analysis of the illustrations included in the original report suggested that M fragillimus may have been the largest land animal of all time possibly weighing 100 150 t 110 170 short tons and measuring between 40 60 m 130 200 ft long 7 8 One later analysis of the surviving evidence and the biological plausibility of such a large land animal suggested that the enormous size of this animal was an over estimate due partly to typographical errors in the original report 9 This would later be challenged by a different study which argued Cope s measurements were genuine and that there was no basis for assuming typographical errors The study however also reclassified the species and correspondingly gave a much lower length estimate of 30 3 metres 99 ft and a mass of 78 5 t 86 5 short tons 10 This in itself would later be disputed as being too small for an animal of such size with some believing it to be even larger at around 35 40 metres 115 131 ft and weighing around 80 120 t 88 132 short tons 11 Another large but even more controversial sauropod is Bruhathkayosaurus which had a calculated weight ranging between 126 220 t 139 243 short tons and a length of 44 1 m 145 ft 12 13 14 Although the existence of this sauropod had long been dismissed as a potential fake or a misidentification of a petrified tree trunk recent photographic evidence emerged confirming its existence 15 More recent and reliable estimates in 2023 have rescaled Bruhathkayosaurus to weigh around 110 130 t 120 140 short tons with its most liberal estimate being 240 t 260 short tons making it incredibly massive for such an animal 16 If the upper unlikely size estimates were to be taken at face value Bruhathkayosaurus would not only be the largest dinosaur to have ever lived but also the largest animal to have lived exceeding even the largest blue whale recorded According to Gregory S Paul super sauropods or land whales such as Maraapunisaurus Bruhathkayosaurus and the Broome Titanosaur footprints as he calls them should not be surprising as sauropods were more heat tolerant and grew rapidly which allowed them to reach truly titanic sizes that rivalled the largest whales in mass despite the prevalence of air sacs 16 Other potential factors for such extreme sauropod sizes include increasing bone robustness and load distributing cartilaginous features to better redistribute and support such massive weights 16 Generally the giant sauropods can be divided into two categories the shorter but stockier and more massive forms mainly titanosaurs and some brachiosaurids and the longer but slenderer and more light weight forms mainly diplodocids Because different methods of estimation sometimes give conflicting results mass estimates for sauropods can vary widely causing disagreement among scientists over the accurate number For example the titanosaur Dreadnoughtus was originally estimated to weigh 59 3 tonnes by the allometric scaling of limb bone proportions whereas more recent estimates based on three dimensional reconstructions yield a much smaller figure of 22 1 38 2 tonnes 17 nbsp Reconstructed skeleton of the titanosaur Argentinosaurus huinculensis often considered the largest known dinosaurThe sauropods were the longest and heaviest dinosaurs For much of the dinosaur era the smallest sauropods were larger than almost anything else in their habitat and the largest were an order of magnitude more massive than anything else known to have walked the Earth since Giant prehistoric mammals such as Paraceratherium and Palaeoloxodon the largest land mammals ever discovered 18 were dwarfed by the giant sauropods and only modern whales approach or surpass them in weight though they live in the oceans 19 There are several proposed advantages for the large size of sauropods including protection from predation reduction of energy use and longevity but it may be that the most important advantage was dietary Large animals are more efficient at digestion than small animals because food spends more time in their digestive systems This also permits them to subsist on food with lower nutritive value than smaller animals Sauropod remains are mostly found in rock formations interpreted as dry or seasonally dry and the ability to eat large quantities of low nutrient browse would have been advantageous in such environments 20 One of the tallest and heaviest dinosaurs known from good skeletons is Giraffatitan brancai previously classified as a species of Brachiosaurus Its remains were discovered in Tanzania between 1907 and 1912 Bones from several similar sized individuals were incorporated into the skeleton now mounted and on display at the Museum fur Naturkunde Berlin 21 this mount is 12 13 27 metres 39 4 43 5 ft tall and 21 8 22 5 metres 72 74 ft long 22 23 24 and would have belonged to an animal that weighed between 30 000 to 60 000 kilograms 66 000 to 132 000 lb One of the longest complete dinosaurs is the 27 metre long 89 ft Diplodocus which was discovered in Wyoming in the United States and displayed in Pittsburgh s Carnegie Natural History Museum in 1907 25 There were larger dinosaurs but knowledge of them is based entirely on a small number of fragmentary fossils Most of the largest herbivorous specimens on record were discovered in the 1970s or later and include the massive titanosaur Argentinosaurus huinculensis which is the largest dinosaur known from uncontroversial and relatively substantial evidence estimated to have been 70 80 t 77 88 short tons and 36 m 118 ft long 26 7 Some of the longest sauropods were those with exceptionally long whip like tails such as the 29 30 m 95 98 ft Diplodocus hallorum 7 26 formerly Seismosaurus and the 39 m Supersaurus 27 In 2014 the fossilized remains of a previously unknown species of sauropod were discovered in Argentina 28 The titanosaur named Patagotitan mayorum was estimated to have been around 40 m 130 ft long weighing around 77 t 85 short tons larger than any other previously found sauropod The specimens found were remarkably complete significantly more so than previous titanosaurs It since been suggested that Patagotitan was not necessarily larger than Argentinosaurus and Puertasaurus 29 In 2019 Patagotitan was estimated to have been 31 metres 102 ft long and about 55 tonnes 121 000 lb massive 30 The largest of non sauropod sauropodomorphs was the unnamed 16 metres 52 ft long 10 tonnes 22 000 lb unnamed Elliot giant 26 Another large sauropodomorph was Euskelosaurus It reached 12 2 m 40 ft in length and 2 t 2 2 short tons in weight 31 Yunnanosaurus youngi also reached a length of 13 m 43 ft 32 Theropods nbsp Size comparison of selected giant theropod dinosaursMain article Theropoda Tyrannosaurus was for many decades the largest and best known theropod to the general public Since its discovery however a number of other giant carnivorous dinosaurs have been described including Spinosaurus Carcharodontosaurus and Giganotosaurus 33 These large theropod dinosaurs are estimated to rival or even exceeded Tyrannosaurus rex in size though more recent studies and reconstructions show that Tyrannosaurus although shorter was the bulkier animal overall Specimens such as Sue and Scotty are both estimated to be the most massive theropods known to science There is still no clear explanation for exactly why these animals grew so bulky and heavy compared to the land predators that came before and after them nbsp Skeleton of Giganotosaurus one of the largest theropods known The largest extant theropod is the common ostrich up to 2 74 metres 9 ft 0 in tall and weighs between 63 5 and 145 15 kilograms 140 0 and 320 0 lb 34 The smallest non avialan theropod known from adult specimens may be Anchiornis huxleyi at 110 grams 3 9 ounces in weight and 34 centimetres 13 in in length 35 although later study discovered larger specimen reaching 62 centimetres 24 in 36 However some studies suggest that Anchiornis was actually an avialan 37 The smallest dinosaur known from adult specimens which is definitely not an avialan is Parvicursor remotus at 162 grams 5 7 oz and measuring 39 centimetres 15 in long 38 However in 2022 its holotype was recognized as a juvenile individual 39 Among living dinosaurs the bee hummingbird Mellisuga helenae is smallest at 1 9 g 0 067 oz and 5 5 cm 2 2 in long 40 The smallest theropod overall including avians is the currently extant Bee Hummingbird at 6 12 cm long and 2 6g for females and 5 51 cm long and 3 25g for the males 41 In the theropod lineage leading to birds body size shrank continuously over a period of 50 million years from an average of 163 kilograms 359 lb down to 0 8 kg 1 8 lb This was the only dinosaur lineage to get continuously smaller over such an extended time period and their skeletons developed adaptations at about four times the average rate for dinosaurs 42 43 See also nbsp Dinosaurs portalLargest prehistoric animals List of largest birds Megafauna Pterosaur sizeReferences Sereno PC 1999 The evolution of dinosaurs Science 284 5423 2137 2147 doi 10 1126 science 284 5423 2137 PMID 10381873 Farlow JA 1993 On the rareness of big fierce animals speculations about the body sizes population densities and geographic ranges of predatory mammals and large carnivorous dinosaurs In Dodson Peter Gingerich Philip eds Functional Morphology and Evolution American Journal of Science Special Volume Vol 293 A pp 167 199 Peczkis J 1994 Implications of body mass estimates for dinosaurs Journal of Vertebrate Paleontology 14 4 520 33 doi 10 1080 02724634 1995 10011575 Anatomy and evolution National Museum of Natural History Archived from the original on 2007 11 11 Retrieved 2007 11 21 Paul Gregory S 2010 Princeton Field Guide to Dinosaurs Princeton University Press ISBN 978 0 691 13720 9 Strauss Bob Why Were Dinosaurs So Big The Facts and Theories Behind Dinosaur Gigantism About EducationArchived 2014 10 06 at the Wayback Machine a b c Paul Gregory S 2016 The Princeton Field Guide to Dinosaurs 2nd Edition United States of America Princeton University Press ISBN 978 0 691 16766 4 Paul G S 1997 Dinosaur models the good the bad and using them to estimate the mass of dinosaurs PDF In Wolberg D L Stump E Rosenberg G D eds DinoFest International Proceedings Dinofest International The Academy of Natural Sciences pp 129 154 Archived PDF from the original on March 4 2016 Woodruff C Foster JR 2015 The fragile legacy of Amphicoelias fragillimus Dinosauria Sauropoda Morrison Formation Latest Jurassic PeerJ PrePrints doi 10 7287 peerj preprints 838v1 Carpenter Kenneth 2018 10 19 Maraapunisaurus fragillimus N G formerly Amphicoelias fragillimus a basal Rebbachisaurid from the Morrison Formation Upper Jurassic of Colorado Geology of the Intermountain West 5 227 244 doi 10 31711 giw v5 pp227 244 ISSN 2380 7601 Paul Gregory S 2019 Determining the largest known land animal A critical comparison of differing methods for restoring the volume and mass of extinct animals PDF Annals of the Carnegie Museum 85 4 335 358 doi 10 2992 007 085 0403 S2CID 210840060 Mortimer M 2001 Re Bruhathkayosaurus discussion group The Dinosaur Mailing List 19 June 2001 Accessed 23 May 2008 Mortimer M 2004 Re Largest Dinosaurs Archived 2019 09 13 at the Wayback Machine discussion group The Dinosaur Mailing List 7 September 2004 Accessed 23 May 2008 Mortimer M 2001 Titanosaurs too large Archived 2016 03 03 at the Wayback Machine discussion group The Dinosaur Mailing List 12 September 2001 Accessed 23 May 2008 Pal Saurabh Ayyasami Krishnan 27 June 2022 The lost titan of Cauvery Geology Today 38 3 112 116 doi 10 1111 gto 12390 ISSN 0266 6979 S2CID 250056201 a b c Paul Gregory S Larramendi Asier 11 April 2023 Body mass estimate of Bruhathkayosaurus and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales Lethaia 56 2 1 11 doi 10 18261 let 56 2 5 ISSN 0024 1164 S2CID 259782734 Bates Karl T Falkingham Peter L Macaulay Sophie Brassey Charlotte Maidment Susannah C R 2015 Downsizing a giant re evaluating Dreadnoughtus body mass Biol Lett 11 6 20150215 doi 10 1098 rsbl 2015 0215 PMC 4528471 PMID 26063751 Larramendi A 2016 Shoulder height body mass and shape of proboscideans PDF Acta Palaeontologica Polonica 61 doi 10 4202 app 00136 2014 S2CID 2092950 Archived PDF from the original on 2016 08 24 Sander P Martin Christian Andreas Clauss Marcus Fechner Regina Gee Carole T Griebeler Eva Maria Gunga Hanns Christian Hummel Jurgen Mallison Heinrich et al 2011 Biology of the sauropod dinosaurs the evolution of gigantism Biological Reviews 86 1 117 155 doi 10 1111 j 1469 185X 2010 00137 x PMC 3045712 PMID 21251189 Carpenter K 2006 Biggest of the big a critical re evaluation of the mega sauropod Amphicoelias fragillimus In Foster J R and Lucas S G eds 2006 Paleontology and Geology of the Upper Jurassic Morrison Formation New Mexico Museum of Natural History and Science Bulletin 36 131 138 Colbert Edwin Harris 1971 Men and dinosaurs the search in field and laboratory Harmondsworth Eng Penguin ISBN 978 0 14 021288 4 Mazzetta G V et al 2004 Giants and Bizarres Body Size of Some Southern South American Cretaceous Dinosaurs Historical Biology 16 2 4 1 13 CiteSeerX 10 1 1 694 1650 doi 10 1080 08912960410001715132 S2CID 56028251 Janensch W 1950 The Skeleton Reconstruction of Brachiosaurus brancai 97 103 a href Template Cite journal html title Template Cite journal cite journal a Cite journal requires journal help The World of Dinosaurs Museum fur Naturkunde Archived from the original on 2018 11 16 Retrieved 2018 11 16 Lucas H Hecket H 2004 Reappraisal of Seismosaurus a Late Jurassic Sauropod Proceeding Annual Meeting of the Society of Paleontology 36 5 422 a b c Molina Perez Ruben Larramendi Asier 2020 Dinosaur Facts and Figures The Sauropods and Other Sauropodomorphs Translated by Donaghey Joan Illustrated by Andrey Atuchin and Sante Mazzei Princeton Princeton University Press ISBN 978 0 691 20297 6 OCLC 1157079384 SVP 2021 Symposium Society of Vertebrate Paleontology Retrieved 2023 12 16 Morgan James 2014 05 17 Biggest dinosaur ever discovered BBC News Archived from the original on 2017 02 18 Retrieved 2017 03 21 Don t believe the hype Patagotitan was not bigger than Argentinosaurus Sauropod Vertebra Picture of the Week 2017 08 09 Archived from the original on 2019 04 14 Retrieved 2019 02 19 Paul Gregory December 2019 Determining the Largest Known Land Animal A Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals PDF Annals of Carnegie Museum 85 4 335 358 doi 10 2992 007 085 0403 ISSN 0097 4463 S2CID 210840060 Patricia Vickers Rich Thomas Hewitt Rich Mildred Adams Fenton Carroll Lane January 15 2020 The Fossil Book A Record of Prehistoric Life Dover Publications p 444 ISBN 9780486838557 Retrieved 2022 08 25 a href Template Cite book html title Template Cite book cite book a CS1 maint multiple names authors list link Lu J Li T Zhong S Azuma Y Fujita M Dong Z and Ji Q 2007 New yunnanosaurid dinosaur Dinosauria Prosauropoda from the Middle Jurassic Zhanghe Formation of Yuanmou Yunnan Province of China Memoir of the Fukui Prefectural Dinosaur Museum 6 1 15 Therrien F Henderson D M 2007 My theropod is bigger than yours or not estimating body size from skull length in theropods Journal of Vertebrate Paleontology 27 1 108 115 doi 10 1671 0272 4634 2007 27 108 MTIBTY 2 0 CO 2 S2CID 86025320 See what African Wildlife Foundation is doing to protect these iconic flightless birds 2013 02 25 Archived from the original on 2015 04 29 Retrieved 2015 04 18 Xu X Zhao Q Norell M Sullivan C Hone D Erickson G Wang X Han F and Guo Y 2009 A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin Chinese Science Bulletin 6 pages accepted November 15 2008 Pei Rui Li Quanguo Meng Qingjin Norell Mark A Gao Ke Qin 2017 New Specimens ofAnchiornis huxleyi Theropoda Paraves from the Late Jurassic of Northeastern China Bulletin of the American Museum of Natural History 411 1 67 doi 10 1206 0003 0090 411 1 1 ISSN 0003 0090 S2CID 90650697 Pascal Godefroit Andrea Cau Hu Dong Yu Francois Escuillie Wu Wenhao Gareth Dyke 2013 A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds Nature 498 7454 359 62 Bibcode 2013Natur 498 359G doi 10 1038 nature12168 PMID 23719374 S2CID 4364892 Which was the smallest dinosaur Archived 2011 07 06 at the Wayback Machine Royal Tyrrell Museum Last accessed 22 September 2022 Averianov AO Lopatin AV 2022 A re appraisal of Parvicursor remotus from the Late Cretaceous of Mongolia implications for the phylogeny and taxonomy of alvarezsaurid theropod dinosaurs Journal of Systematic Palaeontology 19 16 1097 1128 doi 10 1080 14772019 2021 2013965 S2CID 247222017 Conservation International Content Partner Mark McGinley Topic Editor 2008 Biological diversity in the Caribbean Islands In Encyclopedia of Earth Eds Cutler J Cleveland Washington D C Environmental Information Coalition National Council for Science and the Environment First published in the Encyclopedia of Earth May 3 2007 Last revised August 22 2008 Retrieved November 9 2009 Archived 2013 05 23 at the Wayback Machine gt Glick Adrienne Mellisuga helenae Animal Diversity Web Retrieved 04 October 2023 Borenstein Seth July 31 2014 Study traces dinosaur evolution into early birds AP News Archived from the original on August 8 2014 Retrieved August 3 2014 Zoe Gough 31 July 2014 Dinosaurs shrank regularly to become birds BBC Archived from the original on 20 February 2015 Retrieved 20 April 2015 External links nbsp Media related to Dinosauria size comparisons at Wikimedia Commons The Biggest Carnivore Dinosaur History Rewritten Holtz Thomas R Jr Rey Luis V 2007 Dinosaurs the most complete up to date encyclopedia for dinosaur lovers of all ages New York Random House ISBN 978 0 375 82419 7 Dinosaur size References Dinosaur records Czech article by Vladimir Socha DinosaurusBlog com August 1 2016 Retrieved from https en wikipedia org w index php title Dinosaur size amp oldid 1194420308, wikipedia, wiki, book, books, library,

article

, read, download, free, free download, mp3, video, mp4, 3gp, jpg, jpeg, gif, png, picture, music, song, movie, book, game, games.