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Diabloceratops

Diabloceratops (/dˌæblˈsɛrətɒps/ dy-AB-loh-SERR-ə-tops) is an extinct genus of centrosaurine ceratopsian dinosaur that lived approximately 81.4-81 million years ago during the latter part of the Cretaceous Period in what is now Utah, in the United States.[1] Diabloceratops was a medium-sized, moderately built, ground-dwelling, quadrupedal herbivore, that could grow up to an estimated 4.5 metres (15 ft) in length and 1.3 metric tons (1.4 short tons) in body mass. At the time of its discovery, it was the oldest-known ceratopsid, and first centrosaurine known from latitudes south of the U.S. state of Montana. The generic name Diabloceratops means "devil-horned face," coming from Diablo, Spanish for "devil," and ceratops, Latinized Greek for "horned face." The specific name honors Jeffrey Eaton, a paleontologist at Weber State University and long time friend of the lead author Jim Kirkland. Eaton had a big role in establishing the Grand Staircase-Escalante National Monument where the specimen was found. The type species, Diabloceratops eatoni, was named and described in 2010 by James Ian Kirkland and Donald DeBlieux.

Diabloceratops
Temporal range: Late Cretaceous, 81.4–81 Ma
Restored skull
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Ceratopsia
Family: Ceratopsidae
Subfamily: Centrosaurinae
Genus: Diabloceratops
Kirkland and DeBlieux, 2010
Species:
D. eatoni
Binomial name
Diabloceratops eatoni
Kirkland and DeBlieux, 2010

Discovery edit

 
Preserved fossil material of specimen UMNH VP 16704
 
Holotype skull of Diabloceratops

The only specimen of Diabloceratops eatoni was recovered at the Last Chance Creek Member of the Wahweap Formation, in Kane County, Utah. Type specimen UMNH VP 16699 was collected by Don DeBlieux in 2002, at the Last Chance Creek locality of this formation, in intraclastic sandstone that was deposited during the Campanian stage of the Cretaceous period, approximately 82.2 to 77.3 million years ago.[2] It consists of a partial skull with a piece of the lower jaw, with the right side being intact and part of the left side, which has been weathered. Another specimen UMNH VP 16704 was discovered years earlier in 1998 by Joshua A. Smith of the same formation, but was not described until 2010, when it was assigned to Diabloceratops. However, this specimen may not belong to Diabloceratops, because it shares features found only in more derived centrosaurines.[3] Some researchers suggest that the UMNH VP 16704 specimen is more similar to Machairoceratops, Yehuecauhceratops and Menefeeceratops than to Diabloceratops due to the fan-shaped shape end of the squamosal.[4][5] These specimens are housed in the collection of the Natural History Museum of Utah.

Description edit

 
Life restoration

Diabloceratops was a medium-sized ceratopsian, growing up to 4.5 metres (15 ft) in length and 1.3 metric tons (1.4 short tons) in body mass.[6] It was built like a typical ceratopsian in that it had a large neck frill made of bone. It had a small horn on the nose, perhaps a second horn in front of that, and a pair of relatively small horns above the eyes. The skull is deeper and shorter than that of any other centrosaurines.[7] Upon the frill it also had a pair of very long spikes as in Einiosaurus and Styracosaurus. It being one of the earliest centrosaurine ceratopsids, Kirkland noted a character Diabloceratops shared with the more "primitive" protoceratopsid forms. Both possess an accessory opening in the skull that would become much reduced or disappear in later, more advanced ceratopsids. Kirkland saw this as an indication that the earlier species were not together included in some single natural group but instead presented a gradual sequence of ever more derived forms, increasingly closer related to the Ceratopsidae. Additionally, Morschauser et al. found Diabloceratops to not be one of the most basal Centrosaurines, but to lie just outside of the Centrosaurine-Chasmosaurine split, finding it to be, by the definition of Ceratopsidae, its sister.[8]

Classification edit

 
Restored skulls of Nasutoceratops (left) and Diabloceratops, Natural History Museum of Utah

A phylogenetic tree after a recent phylogenetic analysis by Chiba et al. (2017):[9]

However, a 2019 analysis suggested this genus to be outside of the ceratopsid family.[8]

Paleoecology edit

 
Diabloceratops in environment

Habitat edit

The Wahweap Formation has been radiometrically dated as being between 82.2 and 77.3 million years old. The precise age of Diabloceratops has been estimated to be 81.27 Ma, with a range of uncertainty between 81.45-80.99 Ma.[2] During the time that Diabloceratops lived, the Western Interior Seaway was at its widest extent, almost completely isolating southern Laramidia from the rest of North America. The area where dinosaurs lived included lakes, floodplains, and east-flowing rivers. The Wahweap Formation is part of the Grand Staircase region, an immense sequence of sedimentary rock layers that stretch south from Bryce Canyon National Park through Zion National Park and into the Grand Canyon. The presence of rapid sedimentation and other evidence suggests a wet, seasonal climate.[10]

Paleofauna edit

Diabloceratops shared its paleoenvironment with other dinosaurs, such as the hadrosaur Acristavus gagslarsoni,[11][12] and the lambeosaur Adelolophus hutchisoni,[13] unnamed ankylosaurs and pachycephalosaurs, and the theropod Lythronax argestes, which was likely the apex predator in its ecosystem.[14][15][16] Vertebrates present in the Wahweap Formation at the time of Diabloceratops included freshwater fish, bowfins, abundant rays and sharks, turtles like Compsemys, crocodilians,[17] and lungfish.[18] A fair number of mammals lived in this region, which included several genera of multituberculates, cladotherians, marsupials, and placental insectivores.[19] The mammals are more primitive than those that lived in the area that is now the Kaiparowits Formation. Trace fossils are relatively abundant in the Wahweap Formation, and suggest the presence of crocodylomorphs, as well as ornithischian and theropod dinosaurs.[20] In 2010 a unique trace fossil was discovered that suggests a predator-prey relationship between dinosaurs and primitive mammals. The trace fossil includes at least two fossilized mammalian den complexes as well as associated digging grooves presumably caused by a maniraptoran dinosaur. The proximity indicates a case of probable active predation of the burrow inhabitants by the animals that made the claw marks.[21] Invertebrate activity in this formation ranged from fossilized insect burrows in petrified logs[22] to various mollusks, large crabs,[23] and a wide diversity of gastropods and ostracods.[24]

See also edit

References edit

  1. ^ Kirkland, J.I. and DeBlieux, D.D. (2010). "New basal centrosaurine ceratopsian skulls from the Wahweap Formation (Middle Campanian), Grand Staircase–Escalante National Monument, southern Utah", In: Ryan, M.J., Chinnery-Allgeier, B.J., and Eberth, D.A. (eds.) New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium. Bloomington, Indiana University Press, pp. 117–140
  2. ^ a b Beveridge, Tegan L.; Roberts, Eric M.; Ramezani, Jahandar; Titus, Alan L.; Eaton, Jeffrey G.; Irmis, Randall B.; Sertich, Joseph J.W. (April 2022). "Refined geochronology and revised stratigraphic nomenclature of the Upper Cretaceous Wahweap Formation, Utah, U.S.A. and the age of early Campanian vertebrates from southern Laramidia". Palaeogeography, Palaeoclimatology, Palaeoecology. 591: 110876. Bibcode:2022PPP...59110876B. doi:10.1016/j.palaeo.2022.110876. ISSN 0031-0182. S2CID 246766015.
  3. ^ Loewen, Mark A.; Farke, Andrew A.; Sampson, Scott D.; Getty, Michael A.; Lund, Eric K.; O'Connor, Patrick M. (2013). "Ceratopsid Dinosaurs from the Grand Staircase of Southern Utah". In Titus, Alan L.; Loewen, Mark A. (eds.). At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah. Bloomington: Indiana University Press. pp. 488–503. ISBN 9780253008961.
  4. ^ Eric K. Lund; Patrick M. O’Connor; Mark A. Loewen; Zubair A. Jinnah (2016). "A New Centrosaurine Ceratopsid, Machairoceratops cronusi gen et sp. nov., from the Upper Sand Member of the Wahweap Formation (Middle Campanian), Southern Utah". PLOS ONE. 11 (5): e0154403. Bibcode:2016PLoSO..1154403L. doi:10.1371/journal.pone.0154403. PMC 4871575. PMID 27192148.
  5. ^ Rivera-Sylva, H.E.; Hendrick, B.P.; Dodson, P. (2016). "A Centrosaurine (Dinosauria: Ceratopsia) from the Aguja Formation (Late Campanian) of Northern Coahuila, Mexico". PLOS ONE. 11 (4): e0150529. Bibcode:2016PLoSO..1150529R. doi:10.1371/journal.pone.0150529. PMC 4830452. PMID 27073969.
  6. ^ Paul, G. S. (2016). The Princeton Field Guide to Dinosaurs (2nd ed.). Princeton, New Jersey: Princeton University Press. p. 286. ISBN 9780691167664.
  7. ^ "DIABLOCERATOPS". Retrieved December 11, 2013.
  8. ^ a b Morschhauser, E.M.; You, H.; Li, D.; Dodson, P. (2019). "Phylogenetic history of Auroraceratops rugosus (Ceratopsia: Ornithischia) from the Lower Cretaceous of Gansu Province, China". Journal of Vertebrate Paleontology. 38 (Supplement): 117–147. doi:10.1080/02724634.2018.1509866. S2CID 202867827.
  9. ^ Kentaro Chiba; Michael J. Ryan; Federico Fanti; Mark A. Loewen; David C. Evans (2018). "New material and systematic re-evaluation of Medusaceratops lokii (Dinosauria, Ceratopsidae) from the Judith River Formation (Campanian, Montana)". Journal of Paleontology. 92 (2): 272–288. Bibcode:2018JPal...92..272C. doi:10.1017/jpa.2017.62. S2CID 134031275.
  10. ^ Zubair A. Jinnah, #30088 (2009)Sequence Stratigraphic Control from Alluvial Architecture of Upper Cretaceous Fluvial System - Wahweap Formation, Southern Utah, U.S.A. Search and Discovery Article #30088. Posted June 16, 2009.
  11. ^ Gates, T. A.; Horner, J. R.; Hanna, R. R.; Nelson, C. R. (2011). "New unadorned hadrosaurine hadrosaurid (Dinosauria, Ornithopoda) from the Campanian of North America". Journal of Vertebrate Paleontology. 31 (4): 798. Bibcode:2011JVPal..31..798G. doi:10.1080/02724634.2011.577854. S2CID 8878474.
  12. ^ (PDF). Utah Geology. 2007. Archived from the original (PDF) on August 27, 2013. Retrieved December 11, 2013.
  13. ^ Terry A. Gates; Zubair Jinnah; Carolyn Levitt; Michael A. Getty (2014). "New hadrosaurid (Dinosauria, Ornithopoda) specimens from the lower-middle Campanian Wahweap Formation of southern Utah". In David A. Eberth; David C. Evans (eds.). Hadrosaurs: Proceedings of the International Hadrosaur Symposium. Indiana University Press. pp. 156–173. ISBN 978-0-253-01385-9.
  14. ^ John Wesley Powell Memorial Museum display, visited April 30th, 2009
  15. ^ . Natural History Museum of Utah. May 14, 2012. Archived from the original on November 13, 2013. Retrieved November 16, 2013.
  16. ^ http://geology.utah.gov/surveynotes/articles/pdf/horned_dinos_39-3.pdf August 27, 2013, at the Wayback Machine [bare URL PDF]
  17. ^ Thompson, Cameron R. "A preliminary report on biostratigraphy of Cretaceous freshwater rays, Wahweap Formation and John Henry Member of the Straight Cliffs Formation, southern Utah." Abstracts with Programs - Geological Society of America, vol.36, no.4, pp.91, Apr 2004
  18. ^ Orsulak, Megan et al. "A lungfish burrow in late Cretaceous upper capping sandstone member of the Wahweap Formation Cockscomb area, Grand Staircase-Escalante National Monument, Utah." Abstracts with Programs - Geological Society of America, vol. 39, no. 5, pp.43, May 2007
  19. ^ Eaton, Jeffrey G; Cifelli, Richard L. "Review of Cretaceous mammalian paleontology; Grand Staircase-Escalante National Monument, Utah. Abstracts with Programs - Geological Society of America, vol.37, no.7, pp.115, Oct 2005
  20. ^ Tester, Edward et al. Isolated vertebrate tracks from the Upper Cretaceous capping sandstone member of the Wahweap Formation; Grand Staircase-Escalante National Monument, UtahAbstracts with Programs - Geological Society of America, vol. 39, no. 5, pp.42, May 2007
  21. ^ Simpson, Edward L.; Hilbert-Wolf, Hannah L.; Wizevich, Michael C.; Tindall, Sarah E.; Fasinski, Ben R.; Storm, Lauren P.; Needle, Mattathias D. (2010). "Predatory digging behavior by dinosaurs". Geology. 38 (8): 699–702. Bibcode:2010Geo....38..699S. doi:10.1130/G31019.1.
  22. ^ De Blieux, Donald D. "Analysis of Jim's hadrosaur site; a dinosaur site in the middle Campanian (Cretaceous) Wahweap Formation of Grand Staircase-Escalante National Monument (GSENM), southern Utah." Abstracts with Programs - Geological Society of America, vol. 39, no. 5, pp.6, May 2007
  23. ^ Kirkland, James Ian. "An inventory of paleontological resources in the lower Wahweap Formation (lower Campanian), southern Kaiparowits Plateau, Grand Staircase-Escalante National Monument, Utah." Abstracts with Programs - Geological Society of America, vol.37, no.7, pp.114, Oct 2005.
  24. ^ Williams, Jessica A J; Lohrengel, C Frederick. Preliminary study of freshwater gastropods in the Wahweap Formation, Bryce Canyon National Park, Utah. Abstracts with Programs - Geological Society of America, vol. 39, no. 5, pp.43, May 2007

External links edit

  • archosaurmusings - diabloceratops-eatoni

diabloceratops, serr, tops, extinct, genus, centrosaurine, ceratopsian, dinosaur, that, lived, approximately, million, years, during, latter, part, cretaceous, period, what, utah, united, states, medium, sized, moderately, built, ground, dwelling, quadrupedal,. Diabloceratops d aɪ ˌ ae b l oʊ ˈ s ɛr e t ɒ p s dy AB loh SERR e tops is an extinct genus of centrosaurine ceratopsian dinosaur that lived approximately 81 4 81 million years ago during the latter part of the Cretaceous Period in what is now Utah in the United States 1 Diabloceratops was a medium sized moderately built ground dwelling quadrupedal herbivore that could grow up to an estimated 4 5 metres 15 ft in length and 1 3 metric tons 1 4 short tons in body mass At the time of its discovery it was the oldest known ceratopsid and first centrosaurine known from latitudes south of the U S state of Montana The generic name Diabloceratops means devil horned face coming from Diablo Spanish for devil and ceratops Latinized Greek for horned face The specific name honors Jeffrey Eaton a paleontologist at Weber State University and long time friend of the lead author Jim Kirkland Eaton had a big role in establishing the Grand Staircase Escalante National Monument where the specimen was found The type species Diabloceratops eatoni was named and described in 2010 by James Ian Kirkland and Donald DeBlieux DiabloceratopsTemporal range Late Cretaceous 81 4 81 Ma PreꞒ Ꞓ O S D C P T J K Pg N Restored skull Scientific classification Domain Eukaryota Kingdom Animalia Phylum Chordata Clade Dinosauria Clade Ornithischia Clade Ceratopsia Family Ceratopsidae Subfamily Centrosaurinae Genus DiabloceratopsKirkland and DeBlieux 2010 Species D eatoni Binomial name Diabloceratops eatoniKirkland and DeBlieux 2010 Contents 1 Discovery 2 Description 3 Classification 4 Paleoecology 4 1 Habitat 4 2 Paleofauna 5 See also 6 References 7 External linksDiscovery edit nbsp Preserved fossil material of specimen UMNH VP 16704 nbsp Holotype skull of Diabloceratops The only specimen of Diabloceratops eatoni was recovered at the Last Chance Creek Member of the Wahweap Formation in Kane County Utah Type specimen UMNH VP 16699 was collected by Don DeBlieux in 2002 at the Last Chance Creek locality of this formation in intraclastic sandstone that was deposited during the Campanian stage of the Cretaceous period approximately 82 2 to 77 3 million years ago 2 It consists of a partial skull with a piece of the lower jaw with the right side being intact and part of the left side which has been weathered Another specimen UMNH VP 16704 was discovered years earlier in 1998 by Joshua A Smith of the same formation but was not described until 2010 when it was assigned to Diabloceratops However this specimen may not belong to Diabloceratops because it shares features found only in more derived centrosaurines 3 Some researchers suggest that the UMNH VP 16704 specimen is more similar to Machairoceratops Yehuecauhceratops and Menefeeceratops than to Diabloceratops due to the fan shaped shape end of the squamosal 4 5 These specimens are housed in the collection of the Natural History Museum of Utah Description edit nbsp Life restoration Diabloceratops was a medium sized ceratopsian growing up to 4 5 metres 15 ft in length and 1 3 metric tons 1 4 short tons in body mass 6 It was built like a typical ceratopsian in that it had a large neck frill made of bone It had a small horn on the nose perhaps a second horn in front of that and a pair of relatively small horns above the eyes The skull is deeper and shorter than that of any other centrosaurines 7 Upon the frill it also had a pair of very long spikes as in Einiosaurus and Styracosaurus It being one of the earliest centrosaurine ceratopsids Kirkland noted a character Diabloceratops shared with the more primitive protoceratopsid forms Both possess an accessory opening in the skull that would become much reduced or disappear in later more advanced ceratopsids Kirkland saw this as an indication that the earlier species were not together included in some single natural group but instead presented a gradual sequence of ever more derived forms increasingly closer related to the Ceratopsidae Additionally Morschauser et al found Diabloceratops to not be one of the most basal Centrosaurines but to lie just outside of the Centrosaurine Chasmosaurine split finding it to be by the definition of Ceratopsidae its sister 8 Classification edit nbsp Restored skulls of Nasutoceratops left and Diabloceratops Natural History Museum of Utah A phylogenetic tree after a recent phylogenetic analysis by Chiba et al 2017 9 Centrosaurinae Diabloceratops eatoni Machairoceratops cronusi Nasutoceratopsini Avaceratops lammersi ANSP 15800 MOR 692 CMN 8804 Nasutoceratops titusi Malta new taxon Xenoceratops foremostensis Sinoceratops zhuchengensis Wendiceratops pinhornensis Albertaceratops nesmoi Medusaceratops lokii Eucentrosaura Centrosaurini Rubeosaurus ovatus Styracosaurus albertensis Coronosaurus brinkmani Centrosaurus apertus Spinops sternbergorum Pachyrhinosaurini Einiosaurus procurvicornis Pachyrostra Achelousaurus horneri Pachyrhinosaurus canadensis Pachyrhinosaurus lakustai Pachyrhinosaurus perotorum However a 2019 analysis suggested this genus to be outside of the ceratopsid family 8 Paleoecology edit nbsp Diabloceratops in environment Habitat edit The Wahweap Formation has been radiometrically dated as being between 82 2 and 77 3 million years old The precise age of Diabloceratops has been estimated to be 81 27 Ma with a range of uncertainty between 81 45 80 99 Ma 2 During the time that Diabloceratops lived the Western Interior Seaway was at its widest extent almost completely isolating southern Laramidia from the rest of North America The area where dinosaurs lived included lakes floodplains and east flowing rivers The Wahweap Formation is part of the Grand Staircase region an immense sequence of sedimentary rock layers that stretch south from Bryce Canyon National Park through Zion National Park and into the Grand Canyon The presence of rapid sedimentation and other evidence suggests a wet seasonal climate 10 Paleofauna edit Diabloceratops shared its paleoenvironment with other dinosaurs such as the hadrosaur Acristavus gagslarsoni 11 12 and the lambeosaur Adelolophus hutchisoni 13 unnamed ankylosaurs and pachycephalosaurs and the theropod Lythronax argestes which was likely the apex predator in its ecosystem 14 15 16 Vertebrates present in the Wahweap Formation at the time of Diabloceratops included freshwater fish bowfins abundant rays and sharks turtles like Compsemys crocodilians 17 and lungfish 18 A fair number of mammals lived in this region which included several genera of multituberculates cladotherians marsupials and placental insectivores 19 The mammals are more primitive than those that lived in the area that is now the Kaiparowits Formation Trace fossils are relatively abundant in the Wahweap Formation and suggest the presence of crocodylomorphs as well as ornithischian and theropod dinosaurs 20 In 2010 a unique trace fossil was discovered that suggests a predator prey relationship between dinosaurs and primitive mammals The trace fossil includes at least two fossilized mammalian den complexes as well as associated digging grooves presumably caused by a maniraptoran dinosaur The proximity indicates a case of probable active predation of the burrow inhabitants by the animals that made the claw marks 21 Invertebrate activity in this formation ranged from fossilized insect burrows in petrified logs 22 to various mollusks large crabs 23 and a wide diversity of gastropods and ostracods 24 See also edit nbsp dinosaurs portal Timeline of ceratopsian researchReferences edit Kirkland J I and DeBlieux D D 2010 New basal centrosaurine ceratopsian skulls from the Wahweap Formation Middle Campanian Grand Staircase Escalante National Monument southern Utah In Ryan M J Chinnery Allgeier B J and Eberth D A eds New Perspectives on Horned Dinosaurs The Royal Tyrrell Museum Ceratopsian Symposium Bloomington Indiana University Press pp 117 140 a b Beveridge Tegan L Roberts Eric M Ramezani Jahandar Titus Alan L Eaton Jeffrey G Irmis Randall B Sertich Joseph J W April 2022 Refined geochronology and revised stratigraphic nomenclature of the Upper Cretaceous Wahweap Formation Utah U S A and the age of early Campanian vertebrates from southern Laramidia Palaeogeography Palaeoclimatology Palaeoecology 591 110876 Bibcode 2022PPP 59110876B doi 10 1016 j palaeo 2022 110876 ISSN 0031 0182 S2CID 246766015 Loewen Mark A Farke Andrew A Sampson Scott D Getty Michael A Lund Eric K O Connor Patrick M 2013 Ceratopsid Dinosaurs from the Grand Staircase of Southern Utah In Titus Alan L Loewen Mark A eds At the Top of the Grand Staircase The Late Cretaceous of Southern Utah Bloomington Indiana University Press pp 488 503 ISBN 9780253008961 Eric K Lund Patrick M O Connor Mark A Loewen Zubair A Jinnah 2016 A New Centrosaurine Ceratopsid Machairoceratops cronusi gen et sp nov from the Upper Sand Member of the Wahweap Formation Middle Campanian Southern Utah PLOS ONE 11 5 e0154403 Bibcode 2016PLoSO 1154403L doi 10 1371 journal pone 0154403 PMC 4871575 PMID 27192148 Rivera Sylva H E Hendrick B P Dodson P 2016 A Centrosaurine Dinosauria Ceratopsia from the Aguja Formation Late Campanian of Northern Coahuila Mexico PLOS ONE 11 4 e0150529 Bibcode 2016PLoSO 1150529R doi 10 1371 journal pone 0150529 PMC 4830452 PMID 27073969 Paul G S 2016 The Princeton Field Guide to Dinosaurs 2nd ed Princeton New Jersey Princeton University Press p 286 ISBN 9780691167664 DIABLOCERATOPS Retrieved December 11 2013 a b Morschhauser E M You H Li D Dodson P 2019 Phylogenetic history of Auroraceratops rugosus Ceratopsia Ornithischia from the Lower Cretaceous of Gansu Province China Journal of Vertebrate Paleontology 38 Supplement 117 147 doi 10 1080 02724634 2018 1509866 S2CID 202867827 Kentaro Chiba Michael J Ryan Federico Fanti Mark A Loewen David C Evans 2018 New material and systematic re evaluation of Medusaceratops lokii Dinosauria Ceratopsidae from the Judith River Formation Campanian Montana Journal of Paleontology 92 2 272 288 Bibcode 2018JPal 92 272C doi 10 1017 jpa 2017 62 S2CID 134031275 Zubair A Jinnah 30088 2009 Sequence Stratigraphic Control from Alluvial Architecture of Upper Cretaceous Fluvial System Wahweap Formation Southern Utah U S A Search and Discovery Article 30088 Posted June 16 2009 Gates T A Horner J R Hanna R R Nelson C R 2011 New unadorned hadrosaurine hadrosaurid Dinosauria Ornithopoda from the Campanian of North America Journal of Vertebrate Paleontology 31 4 798 Bibcode 2011JVPal 31 798G doi 10 1080 02724634 2011 577854 S2CID 8878474 New Horned Dinosaurs from the Wahweap Formation PDF Utah Geology 2007 Archived from the original PDF on August 27 2013 Retrieved December 11 2013 Terry A Gates Zubair Jinnah Carolyn Levitt Michael A Getty 2014 New hadrosaurid Dinosauria Ornithopoda specimens from the lower middle Campanian Wahweap Formation of southern Utah In David A Eberth David C Evans eds Hadrosaurs Proceedings of the International Hadrosaur Symposium Indiana University Press pp 156 173 ISBN 978 0 253 01385 9 John Wesley Powell Memorial Museum display visited April 30th 2009 Diabloceratops eatoni Natural History Museum of Utah May 14 2012 Archived from the original on November 13 2013 Retrieved November 16 2013 http geology utah gov surveynotes articles pdf horned dinos 39 3 pdf Archived August 27 2013 at the Wayback Machine bare URL PDF Thompson Cameron R A preliminary report on biostratigraphy of Cretaceous freshwater rays Wahweap Formation and John Henry Member of the Straight Cliffs Formation southern Utah Abstracts with Programs Geological Society of America vol 36 no 4 pp 91 Apr 2004 Orsulak Megan et al A lungfish burrow in late Cretaceous upper capping sandstone member of the Wahweap Formation Cockscomb area Grand Staircase Escalante National Monument Utah Abstracts with Programs Geological Society of America vol 39 no 5 pp 43 May 2007 Eaton Jeffrey G Cifelli Richard L Review of Cretaceous mammalian paleontology Grand Staircase Escalante National Monument Utah Abstracts with Programs Geological Society of America vol 37 no 7 pp 115 Oct 2005 Tester Edward et al Isolated vertebrate tracks from the Upper Cretaceous capping sandstone member of the Wahweap Formation Grand Staircase Escalante National Monument UtahAbstracts with Programs Geological Society of America vol 39 no 5 pp 42 May 2007 Simpson Edward L Hilbert Wolf Hannah L Wizevich Michael C Tindall Sarah E Fasinski Ben R Storm Lauren P Needle Mattathias D 2010 Predatory digging behavior by dinosaurs Geology 38 8 699 702 Bibcode 2010Geo 38 699S doi 10 1130 G31019 1 De Blieux Donald D Analysis of Jim s hadrosaur site a dinosaur site in the middle Campanian Cretaceous Wahweap Formation of Grand Staircase Escalante National Monument GSENM southern Utah Abstracts with Programs Geological Society of America vol 39 no 5 pp 6 May 2007 Kirkland James Ian An inventory of paleontological resources in the lower Wahweap Formation lower Campanian southern Kaiparowits Plateau Grand Staircase Escalante National Monument Utah Abstracts with Programs Geological Society of America vol 37 no 7 pp 114 Oct 2005 Williams Jessica A J Lohrengel C Frederick Preliminary study of freshwater gastropods in the Wahweap Formation Bryce Canyon National Park Utah Abstracts with Programs Geological Society of America vol 39 no 5 pp 43 May 2007External links editarchosaurmusings diabloceratops eatoni Retrieved from https en wikipedia org w index php title Diabloceratops amp oldid 1214159736, wikipedia, wiki, book, books, library,

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