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Fukuiraptor

Fukuiraptor ("thief of Fukui") was a medium-sized megaraptoran theropod dinosaur of the Early Cretaceous epoch (either Barremian or Aptian) that lived in what is now Japan.[1] Fukuiraptor is known from the Kitadani Formation[2] and possibly also the Sebayashi Formation.[3]

Fukuiraptor
Temporal range: Early Cretaceous, 127–115 Ma
Reconstructed skeleton
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Megaraptora
Genus: Fukuiraptor
Azuma & Currie, 2000
Type species
Fukuiraptor kitadaniensis
Azuma & Currie, 2000

History edit

 
Reconstructed skeleton and holotype fossils on the lower right

The type specimen is a partial skeleton (designated FPDM-V97122) discovered in the Kitadani quarry near Katsuyama in the Fukui prefecture. It is thought that this specimen was not mature and an adult may have been larger. The remains of many other individuals have been found in the quarry, with numerous humeri, femurs, and teeth being assigned to this species.[2] However, the other individuals recovered from the same locality are mostly juveniles that were smaller than the holotype (Currie & Azuma, 2006), in the smallest case less than a quarter of the holotype's size. A tooth (NDC-P0001) discovered in a block of conglomerate from the Sebayashi Formation has been referred to Fukuiraptor as well.[4]

Description edit

 
Size comparison

As indicated by its slender phalanges, Fukuiraptor was a relatively lightly built animal, regardless of its maturity.[1] In 2010 Gregory Paul gave a length of 5 meters (16 ft) and a weight of 300 kg (660 lbs).[5] Molina-Pérez and Larramendi estimated a length of 4.3 meters (14.1 ft) and a weight of 590 kg (1,300 lbs) in 2016.[6]

The distinctive teeth of Fukuiraptor show similarities with both carcharodontosaurids (being very compressed and blade-like, as well as having wrinkled enamel) and tyrannosaurids (having oblique blood grooves near the serrations).[2] The holotype also had very large and flat manual unguals (hand claws), which played a role in its initial classification as a dromaeosaurid (as the hand claws were mistaken for foot claws) as well as its current classification as a megaraptoran.[7]

Classification edit

Initially considered a member of the Dromaeosauridae when first discovered, its initial describers considered it a carnosaur, related to Allosaurus. More recent studies consider it a megaraptoran, an enigmatic group which may have been part of the family Neovenatoridae.[7] However, more recently, another analysis has proposed that all megaraptorans are actually tyrannosauroids, which would reclassify Fukuiraptor as a tyrannosauroid coelurosaur.[8] Recent cladistic analysis of the theropod Gualicho has suggested that Fukuiraptor and other megaraptorans are either allosauroids, or non-tyrannosauroid basal coelurosaurs.[9]

It has been suggested that Fukuiraptor is a close relative to the Australian megaraptoran known as Australovenator,[10] however a subsequent study has placed Australovenator as a megaraptorid megaraptoran alongside other derived South American taxa, while Fukuiraptor remains a megaraptoran outside of Megaraptoridae.[11]

Below is a cladogram reconstructing the position of Fukuiraptor in the Megaraptora as per Delcourt and Grillo, 2018.[11]

Megaraptora

Fukuiraptor  

Megaraptoridae

Palaeoecology edit

 
Stratigraphic positions of the non-avialan dinosaurs of the Kitadani Formation, Japan.

In the Kitadani Formation, where fossils of Fukuiraptor have been recovered,[1] fossils of other vertebrates have been preserved. Among the species that have been properly described and named are Fukuisaurus tetoriensis,[12] a styracostern ornithischian, Fukuititan nipponensis,[13] a titanosauriform sauropod, Koshisaurus katsuyama,[14] a hadrosauroid ornithischian, Fukuivenator paradoxus,[15] a therizinosaurian theropod, and Fukuipteryx prima,[16] an avialan as defined by the authors.

External links edit

  • Image of the holotype skeleton, published by the twitter account of Thomas Holtz
  • . Fukui Prefectural Dinosaur Museum. Archived from the original on 29 August 2006.

References edit

  1. ^ a b c Azuma, Yoichi; Philip J., Currie (2000). "A new carnosaur (Dinosauria: Theropoda) from the Lower Cretaceous of Japan" (PDF). Canadian Journal of Earth Sciences. 37 (12): 1735–1753. Bibcode:2000CaJES..37.1735A. doi:10.1139/e00-064.
  2. ^ a b c Currie, P.J.; Azuma, Y. (2006). "New specimens, including a growth series, of Fukuiraptor (Dinosauria, Theropoda) from the Lower Cretaceous Kitadani Quarry of Japan". J. Paleont. Soc. Korea. 22 (1): 173–193 – via ResearchGate.
  3. ^ "A tooth of Fukuiraptor aff. F. kitadaniensis from the Lower Cretaceous Sebayashi Formation, Sanchu Cretaceous, Japan: 東京学芸大学リポジトリ". ir.u-gakugei.ac.jp. Retrieved 9 October 2020.
  4. ^ Molnar, Ralph E.; Obata, Ikuwo; Tanimoto, Masahiro; Matsukawa, Masaki (2009). "A tooth of Fukuiraptor aff. F. kitadaniensis from the Lower Cretaceous Sebayashi Formation, Sanchu Cretaceous, Japan". Bulletin of Tokyo Gakugei University, Division of Natural Sciences. 61: 105–117 – via ResearchGate.
  5. ^ Paul, Gregory S. (2010). The Princeton Field Guide to Dinosaurs. New Jersey: Princeton University Press. p. 98.
  6. ^ Molina-Pérez & Larramendi (2016). Récords y curiosidades de los dinosaurios Terópodos y otros dinosauromorfos. Barcelona, Spain: Larousse. p. 263.
  7. ^ a b Benson, R. B. J.; Carrano, M. T.; Brusatte, S. L. (2010). "A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic". Naturwissenschaften. 97 (1): 71–78. Bibcode:2010NW.....97...71B. doi:10.1007/s00114-009-0614-x. PMID 19826771. S2CID 22646156.
  8. ^ Novas, F. E.; Agnolín, F. L.; Ezcurra, M. D.; Canale, J. I.; Porfiri, J. D. (2012). "Megaraptorans as members of an unexpected evolutionary radiation of tyrant-reptiles in Gondwana". Ameghiniana. 49 (4(R): Suplemento Resúmenes [Abstracts]): R33.
  9. ^ Apesteguía, Sebastián; Smith, Nathan D.; Valieri, Rubén Juárez; Makovicky, Peter J. (13 July 2016). "An Unusual New Theropod with a Didactyl Manus from the Upper Cretaceous of Patagonia, Argentina". PLOS ONE. 11 (7): e0157793. Bibcode:2016PLoSO..1157793A. doi:10.1371/journal.pone.0157793. ISSN 1932-6203. PMC 4943716. PMID 27410683.
  10. ^ Currie, Phillip J.; Coria, Rodolfo A. (20 July 2016). "A New Megaraptoran Dinosaur (Dinosauria, Theropoda, Megaraptoridae) from the Late Cretaceous of Patagonia". PLoS One.
  11. ^ a b Delcourt, Rafael; Grillo, Orlando Nelson (15 December 2018). "Tyrannosauroids from the Southern Hemisphere: Implications for biogeography, evolution, and taxonomy". Palaeogeography, Palaeoclimatology, Palaeoecology. 511: 379–387. doi:10.1016/j.palaeo.2018.09.003. ISSN 0031-0182. S2CID 133830150.
  12. ^ Kobayashi, Yoshitsugu; Azuma, Yoichi (11 April 2003). "A new iguanodontian (Dinosauria: Ornithopoda) from the Lower Cretaceous Kitadani Formation of Fukui Prefecture, Japan". Journal of Vertebrate Paleontology. 23 (1): 166–175. doi:10.1671/0272-4634(2003)23[166:anidof]2.0.co;2. ISSN 0272-4634. S2CID 131386181.
  13. ^ Azuma, Yoichi; Shibata, Masateru (2010). "Fukuititan nipponensis, A New Titanosauriform Sauropod from the Early Cretaceous Tetori Group of Fukui Prefecture, Japan". Acta Geologica Sinica - English Edition. 84 (3): 454–462. doi:10.1111/j.1755-6724.2010.00268.x. ISSN 1755-6724. S2CID 128897110.
  14. ^ "New basal hadrosauroid (Dinosauria: Ornithopoda) from the Lower Cretaceous Kitadani Formation, Fukui, central Japan" (PDF). Zootaxa.
  15. ^ Azuma, Yoichi; Xu, Xing; Shibata, Masateru; Kawabe, Soichiro; Miyata, Kazunori; Imai, Takuya (23 February 2016). "A bizarre theropod from the Early Cretaceous of Japan highlighting mosaic evolution among coelurosaurians". Scientific Reports. 6 (1): 20478. doi:10.1038/srep20478. ISSN 2045-2322. PMC 4763874. PMID 26908367.
  16. ^ Imai, Takuya; Azuma, Yoichi; Kawabe, Soichiro; Shibata, Masateru; Miyata, Kazunori; Wang, Min; Zhou, Zhonghe (14 November 2019). "An unusual bird (Theropoda, Avialae) from the Early Cretaceous of Japan suggests complex evolutionary history of basal birds". Communications Biology. 2 (1): 399. doi:10.1038/s42003-019-0639-4. ISSN 2399-3642. PMC 6856171. PMID 31754639.

fukuiraptor, thief, fukui, medium, sized, megaraptoran, theropod, dinosaur, early, cretaceous, epoch, either, barremian, aptian, that, lived, what, japan, known, from, kitadani, formation, possibly, also, sebayashi, formation, temporal, range, early, cretaceou. Fukuiraptor thief of Fukui was a medium sized megaraptoran theropod dinosaur of the Early Cretaceous epoch either Barremian or Aptian that lived in what is now Japan 1 Fukuiraptor is known from the Kitadani Formation 2 and possibly also the Sebayashi Formation 3 FukuiraptorTemporal range Early Cretaceous 127 115 Ma PreꞒ Ꞓ O S D C P T J K Pg N Reconstructed skeleton Scientific classification Domain Eukaryota Kingdom Animalia Phylum Chordata Clade Dinosauria Clade Saurischia Clade Theropoda Clade Megaraptora Genus FukuiraptorAzuma amp Currie 2000 Type species Fukuiraptor kitadaniensisAzuma amp Currie 2000 Not to be confused with Fukuivenator Contents 1 History 2 Description 3 Classification 4 Palaeoecology 5 External links 6 ReferencesHistory edit nbsp Reconstructed skeleton and holotype fossils on the lower right The type specimen is a partial skeleton designated FPDM V97122 discovered in the Kitadani quarry near Katsuyama in the Fukui prefecture It is thought that this specimen was not mature and an adult may have been larger The remains of many other individuals have been found in the quarry with numerous humeri femurs and teeth being assigned to this species 2 However the other individuals recovered from the same locality are mostly juveniles that were smaller than the holotype Currie amp Azuma 2006 in the smallest case less than a quarter of the holotype s size A tooth NDC P0001 discovered in a block of conglomerate from the Sebayashi Formation has been referred to Fukuiraptor as well 4 Description edit nbsp Size comparison As indicated by its slender phalanges Fukuiraptor was a relatively lightly built animal regardless of its maturity 1 In 2010 Gregory Paul gave a length of 5 meters 16 ft and a weight of 300 kg 660 lbs 5 Molina Perez and Larramendi estimated a length of 4 3 meters 14 1 ft and a weight of 590 kg 1 300 lbs in 2016 6 The distinctive teeth of Fukuiraptor show similarities with both carcharodontosaurids being very compressed and blade like as well as having wrinkled enamel and tyrannosaurids having oblique blood grooves near the serrations 2 The holotype also had very large and flat manual unguals hand claws which played a role in its initial classification as a dromaeosaurid as the hand claws were mistaken for foot claws as well as its current classification as a megaraptoran 7 Classification editInitially considered a member of the Dromaeosauridae when first discovered its initial describers considered it a carnosaur related to Allosaurus More recent studies consider it a megaraptoran an enigmatic group which may have been part of the family Neovenatoridae 7 However more recently another analysis has proposed that all megaraptorans are actually tyrannosauroids which would reclassify Fukuiraptor as a tyrannosauroid coelurosaur 8 Recent cladistic analysis of the theropod Gualicho has suggested that Fukuiraptor and other megaraptorans are either allosauroids or non tyrannosauroid basal coelurosaurs 9 It has been suggested that Fukuiraptor is a close relative to the Australian megaraptoran known as Australovenator 10 however a subsequent study has placed Australovenator as a megaraptorid megaraptoran alongside other derived South American taxa while Fukuiraptor remains a megaraptoran outside of Megaraptoridae 11 Below is a cladogram reconstructing the position of Fukuiraptor in the Megaraptora as per Delcourt and Grillo 2018 11 Megaraptora Fukuiraptor nbsp Megaraptoridae Murusraptor nbsp Tratayenia Megaraptor nbsp Orkoraptor nbsp Aerosteon nbsp Australovenator nbsp Palaeoecology edit nbsp Stratigraphic positions of the non avialan dinosaurs of the Kitadani Formation Japan In the Kitadani Formation where fossils of Fukuiraptor have been recovered 1 fossils of other vertebrates have been preserved Among the species that have been properly described and named are Fukuisaurus tetoriensis 12 a styracostern ornithischian Fukuititan nipponensis 13 a titanosauriform sauropod Koshisaurus katsuyama 14 a hadrosauroid ornithischian Fukuivenator paradoxus 15 a therizinosaurian theropod and Fukuipteryx prima 16 an avialan as defined by the authors External links edit nbsp Dinosaurs portal Image of the holotype skeleton published by the twitter account of Thomas Holtz Fukuiraptor kitadaniensis Fukui Prefectural Dinosaur Museum Archived from the original on 29 August 2006 References edit a b c Azuma Yoichi Philip J Currie 2000 A new carnosaur Dinosauria Theropoda from the Lower Cretaceous of Japan PDF Canadian Journal of Earth Sciences 37 12 1735 1753 Bibcode 2000CaJES 37 1735A doi 10 1139 e00 064 a b c Currie P J Azuma Y 2006 New specimens including a growth series of Fukuiraptor Dinosauria Theropoda from the Lower Cretaceous Kitadani Quarry of Japan J Paleont Soc Korea 22 1 173 193 via ResearchGate A tooth of Fukuiraptor aff F kitadaniensis from the Lower Cretaceous Sebayashi Formation Sanchu Cretaceous Japan 東京学芸大学リポジトリ ir u gakugei ac jp Retrieved 9 October 2020 Molnar Ralph E Obata Ikuwo Tanimoto Masahiro Matsukawa Masaki 2009 A tooth of Fukuiraptor aff F kitadaniensis from the Lower Cretaceous Sebayashi Formation Sanchu Cretaceous Japan Bulletin of Tokyo Gakugei University Division of Natural Sciences 61 105 117 via ResearchGate Paul Gregory S 2010 The Princeton Field Guide to Dinosaurs New Jersey Princeton University Press p 98 Molina Perez amp Larramendi 2016 Records y curiosidades de los dinosaurios Teropodos y otros dinosauromorfos Barcelona Spain Larousse p 263 a b Benson R B J Carrano M T Brusatte S L 2010 A new clade of archaic large bodied predatory dinosaurs Theropoda Allosauroidea that survived to the latest Mesozoic Naturwissenschaften 97 1 71 78 Bibcode 2010NW 97 71B doi 10 1007 s00114 009 0614 x PMID 19826771 S2CID 22646156 Novas F E Agnolin F L Ezcurra M D Canale J I Porfiri J D 2012 Megaraptorans as members of an unexpected evolutionary radiation of tyrant reptiles in Gondwana Ameghiniana 49 4 R Suplemento Resumenes Abstracts R33 Apesteguia Sebastian Smith Nathan D Valieri Ruben Juarez Makovicky Peter J 13 July 2016 An Unusual New Theropod with a Didactyl Manus from the Upper Cretaceous of Patagonia Argentina PLOS ONE 11 7 e0157793 Bibcode 2016PLoSO 1157793A doi 10 1371 journal pone 0157793 ISSN 1932 6203 PMC 4943716 PMID 27410683 Currie Phillip J Coria Rodolfo A 20 July 2016 A New Megaraptoran Dinosaur Dinosauria Theropoda Megaraptoridae from the Late Cretaceous of Patagonia PLoS One a b Delcourt Rafael Grillo Orlando Nelson 15 December 2018 Tyrannosauroids from the Southern Hemisphere Implications for biogeography evolution and taxonomy Palaeogeography Palaeoclimatology Palaeoecology 511 379 387 doi 10 1016 j palaeo 2018 09 003 ISSN 0031 0182 S2CID 133830150 Kobayashi Yoshitsugu Azuma Yoichi 11 April 2003 A new iguanodontian Dinosauria Ornithopoda from the Lower Cretaceous Kitadani Formation of Fukui Prefecture Japan Journal of Vertebrate Paleontology 23 1 166 175 doi 10 1671 0272 4634 2003 23 166 anidof 2 0 co 2 ISSN 0272 4634 S2CID 131386181 Azuma Yoichi Shibata Masateru 2010 Fukuititan nipponensis A New Titanosauriform Sauropod from the Early Cretaceous Tetori Group of Fukui Prefecture Japan Acta Geologica Sinica English Edition 84 3 454 462 doi 10 1111 j 1755 6724 2010 00268 x ISSN 1755 6724 S2CID 128897110 New basal hadrosauroid Dinosauria Ornithopoda from the Lower Cretaceous Kitadani Formation Fukui central Japan PDF Zootaxa Azuma Yoichi Xu Xing Shibata Masateru Kawabe Soichiro Miyata Kazunori Imai Takuya 23 February 2016 A bizarre theropod from the Early Cretaceous of Japan highlighting mosaic evolution among coelurosaurians Scientific Reports 6 1 20478 doi 10 1038 srep20478 ISSN 2045 2322 PMC 4763874 PMID 26908367 Imai Takuya Azuma Yoichi Kawabe Soichiro Shibata Masateru Miyata Kazunori Wang Min Zhou Zhonghe 14 November 2019 An unusual bird Theropoda Avialae from the Early Cretaceous of Japan suggests complex evolutionary history of basal birds Communications Biology 2 1 399 doi 10 1038 s42003 019 0639 4 ISSN 2399 3642 PMC 6856171 PMID 31754639 Retrieved from https en wikipedia org w index php title Fukuiraptor amp oldid 1208983687, wikipedia, wiki, book, books, library,

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