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Clymeniida

January 01, 1970

Clymeniida is an order of ammonoid cephalopods from the Upper Devonian characterized by having an unusual dorsal siphuncle. They measured about 4 cm (1.6 in) in diameter and are most common in Europe, North Africa, and South China but are known from North America (New York, Ohio, Oklahoma, and Utah) and Australia (New South Wales) as well.[1]

Clymeniida
Temporal range: Upper Devonian
Clymenia laevigata
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: †Ammonoidea
Order: †Clymeniida
Hyatt, 1884
Suborders

Morphologic characteristics edit

Clymeniids produced a variety of shells ranging from smooth to ribbed and spinose, from evolute with all whorls exposed to strongly involute with the last whorl covering the previous. Some were even triangular in lateral view. With the exception of the first few chambers, all have a siphuncle that runs along the dorsal margin, i.e., along the inner edge of each whorl, rather than the outer edge as in most ammonoids.

Developmentally, the siphuncle in clymeniids starts off ventrally, like that in other ammonoids, but after the first few septa it migrates to a dorsal position indefinitely. Septal necks are retrosiphonate, characteristic of their nautiloid ancestors, and are commonly very long, forming an almost continuous siphuncular tube. The septa, unusually simplified in shape, are convex toward the front as is characteristic of ammonoids.

Classification edit

Miller, Furnish, and Schindewolf, 1957, in Part L of the Treatise on Invertebrate Paleontology[2] included three superfamilies in Clymeniida: Gonioclymeniaceae, Clymeniaceae, and Parawocklumeriaceae. Gonoclymeniaceae unites five families, Clymeniaceae three. Parawocklumeraceae was established for the single family Parawocklumeriidae.

In 1999, Saunders, Work, and Nikolaeva [3] divided Clymeniida into two suborders, Gonioclymeniina and Clymeniina. Gonioclymeniina includes Sellaclymeniaceae, with 7 families and 20 genera; Gonioclyemniaceae, containing a single family with six genera; and Parawocklumeraceae, also containing a single family but with only three genera. Clymeniina contains a single superfamily, Clymeniaceae, which unites five families with a total of 36 genera.

Per the GONIAT website,[4] two suborders fall into Clymeniida. They are Clymeniina with four superfamilies and Gonioclymeniina, established for a single superfamily. In contrast, Dieter Korn in 2006 divided Clymeniida into Clymeniina and Cyrtoclymeniina, named by him in 2002.[5] Shevyrev, on the other hand, in 2006 divided Clymeniida into Clymeniina and Gonioclymeniina, in line with GONIAT and with the work of Saunders, Work, and Nikolaeva.[6] The Taxonomy section below presents Korn's (2006) classification.

Taxonomy edit

Clymeniida

  • Suborder Clymeniina Hyatt, 1884
    • Superfamily Clymeniaceae Edwards, 1849
      • Family Clymeniidae Edwards, 1849
      • Family Kosmoclymeniidae Korn and Price, 1987
        • Subfamily Kosmoclymeniinae Korn and Price, 1987
          • Genus Kosmoclymenia Schindewolf, 1949
          • Genus Linguaclymenia Korn and Price, 1987
          • Genus Lissoclymenia Korn and Price, 1987
          • Genus Muessenbiaergia Korn and Price, 1987
        • Subfamily Rodeckiinae Korn 2002
          • Genus Franconiclymenia Korn and Price, 1987
          • Genus Protoxyclymenia Schindewolf, 1923
          • Genus Rodeckia Korn, 2002
    • Superfamily Gonioclymeniaceae Hyatt, 1884
      • Family Costaclymeniidae Ruzhencev, 1957
        • Genus Costaclymenia Schindewolf, 1920
        • Genus Endosiphonites Ansted, 1838
      • Family Gonioclymeniidae Hyatt, 1884
        • Genus Finiclymenia Price and Korn 1989
        • Genus Gonioclymenia Sepkoski, Jr., 2002
        • Genus Kalloclymenia Wedekind, 1914
        • Genus Leviclymenia Korn, 2002
        • Genus Mesoclymenia Bogoslovsky, 1981
      • Family Sellaclymeniidae Schindewolf, 1923
        • Genus Sellaclymenia Sepkoski, Jr., 2002
      • Family Sphenoclymeniidae Korn, 1992
        • Genus Medioclymenia Korn, 2002
        • Genus Sphenoclymenia Schindewolf, 1920
    • Superfamily Platyclymeniaceae Wedekind [de], 1914
      • Family Glatziellidae Schindewolf, 1928
        • Genus Glatziella Renz, 1914
        • Genus Liroclymenia Czarnocki, 1989
        • Genus Postglatziella Schindewolf, 1937
        • Genus Soliclymenia Schindewolf, 1937
      • Family Piriclymeniidae Korn, 1992
        • Genus Ornatoclymenia Bogoslovsky, 1979
        • Genus Piriclymenia Schindewolf, 1937
        • Genus Sulcoclymenia Schindewolf, 1923
      • Family Platyclymeniidae Wedekind, 1914
        • Subfamily Nodosoclymeniinae Korn, 2002
          • Genus Czarnoclymenia Korn, 1999
          • Genus Nodosoclymenia Czarnocki, 1989
          • Genus Stenoclymenia Lange, 1929
        • Subfamily Platyclymeniinae Wedekind, 1914
          • Genus Fasciclymenia Korn and Price, 1987
          • Genus Platyclymenia Sepkoski, Jr., 2002
          • Genus Progonioclymenia Schindewolf, 1937
          • Genus Spinoclymenia Bogoslovsky, 1962
          • Genus Trigonoclymenia Schindewolf, 1934
          • Genus Varioclymenia Wedekind, 1908
        • Subfamily Pleuroclymeniinae Korn, 2002
          • Genus Borisiclymenia Korn, 2002
          • Genus Nanoclymenia Korn, 2002
          • Genus Pleuroclymenia Schindewolf, 1934
          • Genus Trochoclymenia Schindewolf, 1926
    • Superfamily Wocklumeriaceae Schindewolf, 1937
      • Family Parawocklumeriidae Schindewolf, 1937
        • Genus Kamptoclymenia Schindewolf, 1937
        • Genus Parawocklumeria Schindewolf, 1926
        • Genus Tardewocklumeria Becker, 2000
        • Genus Triaclymenia Schindewolf, 1937
      • Family Wocklumeriidae Schindewolf, 1937
        • Genus Epiwocklumeria Schindewolf, 1937
        • Genus Kielcensia Czarnocki, 1989
        • Genus Synwocklumeria Librovitch, 1957
        • Genus Wocklumeria Wedekind, 1918
  • Suborder Cyrtoclymeniina Korn, 2002
    • Superfamily Biloclymeniaceae Bogoslovsky, 1955
    • Superfamily Cyrtoclymeniaceae Hyatt, 1884
      • Family Carinoclymeniidae Bogoslovsky, 1975
      • Family Cymaclymeniidae Hyatt 1884
        • Subfamily Cymaclymeniinae Hyatt, 1884
          • Genus Cymaclymenia Sepkoski, Jr., 2002
          • Genus Laganoclymenia Bogoslovsky, 1979
          • Genus Procymaclymenia Korn, 2002
          • Genus Rodachia Korn, 2002
        • Subfamily Genuclymeniinae Korn, 2002
          • Genus Flexiclymenia Czarnocki, 1989
          • Genus Genuclymenia Wedekind, 1908
          • Genus Siekluckia Czarnocki, 1989
      • Family Cyrtoclymeniidae Hyatt, 1884
        • Genus Cyrtoclymenia Sepkoski, Jr., 2002
        • Genus Hexaclymenia Schindewolf, 1923
        • Genus Praeflexiclymenia Czarnocki, 1989
        • Genus Pricella Korn, 1991
        • Genus Protactoclymenia Wedekind, 1908
      • Family Rectoclymeniidae Schindewolf, 1923
        • Genus Cteroclymenia Bogoslovsky, 1979
        • Genus Falciclymenia Schindewolf, 1923
        • Genus Karadzharia Korn, 2002
        • Genus Rectoclymenia Wedekind, 1908
  • Suborder Incertae sedis
    • Genus Borkinia
    • Genus Gyroclymenia
    • Genus Kazakhoclymenia
    • Genus Miroclymenia
    • Genus Schizoclymenia

References edit

  1. ^ Dieter, Korn; Kenneth, De Baets (2015), Klug, Christian; Korn, Dieter; De Baets, Kenneth; Kruta, Isabelle (eds.), "Biogeography of Paleozoic Ammonoids", Ammonoid Paleobiology: From macroevolution to paleogeography, vol. 44, Dordrecht: Springer Netherlands, pp. 145–161, doi:10.1007/978-94-017-9633-0_6, ISBN 978-94-017-9632-3, retrieved 2023-07-10
  2. ^ Miller, Furnish, and Schindewolf, 1957; Paleozoic Ammonoidea; Treatise on Invertebrate Paleontology, Part-L, Ammonoidea, Geological Society of America & Univ of Kansas.
  3. ^ Evolution of Complexity in Paleozoic Ammonoid Sutures; Supplementary Materia [1]
  4. ^ GONIAT
  5. ^ D. Korn. 2006. Paleozoic ammonoid classification
  6. ^ Classification of A. A. Shevyrev 2006

January 01, 1970
clymeniida, order, ammonoid, cephalopods, from, upper, devonian, characterized, having, unusual, dorsal, siphuncle, they, measured, about, diameter, most, common, europe, north, africa, south, china, known, from, north, america, york, ohio, oklahoma, utah, aus. Clymeniida is an order of ammonoid cephalopods from the Upper Devonian characterized by having an unusual dorsal siphuncle They measured about 4 cm 1 6 in in diameter and are most common in Europe North Africa and South China but are known from North America New York Ohio Oklahoma and Utah and Australia New South Wales as well 1 ClymeniidaTemporal range Upper Devonian PreꞒ Ꞓ O S D C P T J K Pg N Clymenia laevigata Scientific classification Domain Eukaryota Kingdom Animalia Phylum Mollusca Class Cephalopoda Subclass Ammonoidea Order ClymeniidaHyatt 1884 Suborders Clymeniina Cyrtoclymeniina Contents 1 Morphologic characteristics 2 Classification 3 Taxonomy 4 ReferencesMorphologic characteristics editClymeniids produced a variety of shells ranging from smooth to ribbed and spinose from evolute with all whorls exposed to strongly involute with the last whorl covering the previous Some were even triangular in lateral view With the exception of the first few chambers all have a siphuncle that runs along the dorsal margin i e along the inner edge of each whorl rather than the outer edge as in most ammonoids Developmentally the siphuncle in clymeniids starts off ventrally like that in other ammonoids but after the first few septa it migrates to a dorsal position indefinitely Septal necks are retrosiphonate characteristic of their nautiloid ancestors and are commonly very long forming an almost continuous siphuncular tube The septa unusually simplified in shape are convex toward the front as is characteristic of ammonoids Classification editMiller Furnish and Schindewolf 1957 in Part L of the Treatise on Invertebrate Paleontology 2 included three superfamilies in Clymeniida Gonioclymeniaceae Clymeniaceae and Parawocklumeriaceae Gonoclymeniaceae unites five families Clymeniaceae three Parawocklumeraceae was established for the single family Parawocklumeriidae In 1999 Saunders Work and Nikolaeva 3 divided Clymeniida into two suborders Gonioclymeniina and Clymeniina Gonioclymeniina includes Sellaclymeniaceae with 7 families and 20 genera Gonioclyemniaceae containing a single family with six genera and Parawocklumeraceae also containing a single family but with only three genera Clymeniina contains a single superfamily Clymeniaceae which unites five families with a total of 36 genera Per the GONIAT website 4 two suborders fall into Clymeniida They are Clymeniina with four superfamilies and Gonioclymeniina established for a single superfamily In contrast Dieter Korn in 2006 divided Clymeniida into Clymeniina and Cyrtoclymeniina named by him in 2002 5 Shevyrev on the other hand in 2006 divided Clymeniida into Clymeniina and Gonioclymeniina in line with GONIAT and with the work of Saunders Work and Nikolaeva 6 The Taxonomy section below presents Korn s 2006 classification Taxonomy editClymeniida Suborder Clymeniina Hyatt 1884 Superfamily Clymeniaceae Edwards 1849 Family Clymeniidae Edwards 1849 Genus Aktuboclymenia Bogoslovsky 1979 Genus Clymenia Munster 1834 Family Kosmoclymeniidae Korn and Price 1987 Subfamily Kosmoclymeniinae Korn and Price 1987 Genus Kosmoclymenia Schindewolf 1949 Genus Linguaclymenia Korn and Price 1987 Genus Lissoclymenia Korn and Price 1987 Genus Muessenbiaergia Korn and Price 1987 Subfamily Rodeckiinae Korn 2002 Genus Franconiclymenia Korn and Price 1987 Genus Protoxyclymenia Schindewolf 1923 Genus Rodeckia Korn 2002 Superfamily Gonioclymeniaceae Hyatt 1884 Family Costaclymeniidae Ruzhencev 1957 Genus Costaclymenia Schindewolf 1920 Genus Endosiphonites Ansted 1838 Family Gonioclymeniidae Hyatt 1884 Genus Finiclymenia Price and Korn 1989 Genus Gonioclymenia Sepkoski Jr 2002 Genus Kalloclymenia Wedekind 1914 Genus Leviclymenia Korn 2002 Genus Mesoclymenia Bogoslovsky 1981 Family Sellaclymeniidae Schindewolf 1923 Genus Sellaclymenia Sepkoski Jr 2002 Family Sphenoclymeniidae Korn 1992 Genus Medioclymenia Korn 2002 Genus Sphenoclymenia Schindewolf 1920 Superfamily Platyclymeniaceae Wedekind de 1914 Family Glatziellidae Schindewolf 1928 Genus Glatziella Renz 1914 Genus Liroclymenia Czarnocki 1989 Genus Postglatziella Schindewolf 1937 Genus Soliclymenia Schindewolf 1937 Family Piriclymeniidae Korn 1992 Genus Ornatoclymenia Bogoslovsky 1979 Genus Piriclymenia Schindewolf 1937 Genus Sulcoclymenia Schindewolf 1923 Family Platyclymeniidae Wedekind 1914 Subfamily Nodosoclymeniinae Korn 2002 Genus Czarnoclymenia Korn 1999 Genus Nodosoclymenia Czarnocki 1989 Genus Stenoclymenia Lange 1929 Subfamily Platyclymeniinae Wedekind 1914 Genus Fasciclymenia Korn and Price 1987 Genus Platyclymenia Sepkoski Jr 2002 Genus Progonioclymenia Schindewolf 1937 Genus Spinoclymenia Bogoslovsky 1962 Genus Trigonoclymenia Schindewolf 1934 Genus Varioclymenia Wedekind 1908 Subfamily Pleuroclymeniinae Korn 2002 Genus Borisiclymenia Korn 2002 Genus Nanoclymenia Korn 2002 Genus Pleuroclymenia Schindewolf 1934 Genus Trochoclymenia Schindewolf 1926 Superfamily Wocklumeriaceae Schindewolf 1937 Family Parawocklumeriidae Schindewolf 1937 Genus Kamptoclymenia Schindewolf 1937 Genus Parawocklumeria Schindewolf 1926 Genus Tardewocklumeria Becker 2000 Genus Triaclymenia Schindewolf 1937 Family Wocklumeriidae Schindewolf 1937 Genus Epiwocklumeria Schindewolf 1937 Genus Kielcensia Czarnocki 1989 Genus Synwocklumeria Librovitch 1957 Genus Wocklumeria Wedekind 1918 Suborder Cyrtoclymeniina Korn 2002 Superfamily Biloclymeniaceae Bogoslovsky 1955 Family Biloclymeniidae Bogoslovsky 1955 Genus Biloclymenia Schindewolf 1923 Genus Dimeroclymenia Czarnocki 1989 Genus Kiaclymenia Bogoslovsky 1965 Genus Rhiphaeoclymenia Bogoslovsky 1965 Family Pachyclymeniidae Korn 1992 Genus Pachyclymenia Schindewolf 1937 Genus Uraloclymenia Bogoslovsky 1977 Superfamily Cyrtoclymeniaceae Hyatt 1884 Family Carinoclymeniidae Bogoslovsky 1975 Genus Acriclymenia Bogoslovsky 1975 Genus Carinoclymenia Bogoslovsky 1965 Genus Karaclymenia Bogoslovsky 1983 Genus Pinacoclymenia Bogoslovsky 1975 Family Cymaclymeniidae Hyatt 1884 Subfamily Cymaclymeniinae Hyatt 1884 Genus Cymaclymenia Sepkoski Jr 2002 Genus Laganoclymenia Bogoslovsky 1979 Genus Procymaclymenia Korn 2002 Genus Rodachia Korn 2002 Subfamily Genuclymeniinae Korn 2002 Genus Flexiclymenia Czarnocki 1989 Genus Genuclymenia Wedekind 1908 Genus Siekluckia Czarnocki 1989 Family Cyrtoclymeniidae Hyatt 1884 Genus Cyrtoclymenia Sepkoski Jr 2002 Genus Hexaclymenia Schindewolf 1923 Genus Praeflexiclymenia Czarnocki 1989 Genus Pricella Korn 1991 Genus Protactoclymenia Wedekind 1908 Family Rectoclymeniidae Schindewolf 1923 Genus Cteroclymenia Bogoslovsky 1979 Genus Falciclymenia Schindewolf 1923 Genus Karadzharia Korn 2002 Genus Rectoclymenia Wedekind 1908 Suborder Incertae sedis Genus Borkinia Genus Gyroclymenia Genus Kazakhoclymenia Genus Miroclymenia Genus SchizoclymeniaReferences edit Dieter Korn Kenneth De Baets 2015 Klug Christian Korn Dieter De Baets Kenneth Kruta Isabelle eds Biogeography of Paleozoic Ammonoids Ammonoid Paleobiology From macroevolution to paleogeography vol 44 Dordrecht Springer Netherlands pp 145 161 doi 10 1007 978 94 017 9633 0 6 ISBN 978 94 017 9632 3 retrieved 2023 07 10 Miller Furnish and Schindewolf 1957 Paleozoic Ammonoidea Treatise on Invertebrate Paleontology Part L Ammonoidea Geological Society of America amp Univ of Kansas Evolution of Complexity in Paleozoic Ammonoid Sutures Supplementary Materia 1 GONIAT D Korn 2006 Paleozoic ammonoid classification Classification of A A Shevyrev 2006 Retrieved from https en wikipedia org w index php title Clymeniida amp oldid 1164742479, wikipedia, wiki, book, books, library,

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