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Physa

Physa is a genus of small, left-handed or sinistral, air-breathing freshwater snails, aquatic pulmonate gastropod mollusks in the subfamily Physinae of the family Physidae.[2]

Physa
Five shells of Physa fontinalis
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Heterobranchia
Superorder: Hygrophila
Superfamily: Lymnaeoidea
Family: Physidae
Genus: Physa
Draparnaud, 1801[1]
Type species
Bulla fontinalis Linnaeus, 1758
Synonyms
  • Laurentiphysa Taylor, 2003 (a junior synonym)
  • Limnea (Physa) Draparnaud, 1801
  • Physa (Diastropha) Gray, 1840
  • Physa (Gyrina) Schumacher, 1817
  • Physa (Laurentiphysa) Taylor, 2003 (a junior synonym)
  • Physa (Mediterraneophysa) Starobogatov & Budnikova, 1976 (a junior synonym)
  • Physa (Physa) Draparnaud, 1801
  • Physa (Ussuriphysa) Starobogatov & Prozorova, 1989
  • Rivicola Fitzinger, 1833 (Invalid: junior objective synonym of Physa, with the same type species)

These snails eat algae, diatoms and detritus.

Anatomy edit

Members of the freshwater pulmonate family Physidae possess a complex of muscles that is unique amongst gastropods. This complex was given the name "physid musculature". The physid musculature has two main components, the physid muscle sensu stricto and the fan muscle. The physid musculature is responsible for a unique ability of physids to rapidly flick their shells from side to side — a reaction that frequently enables them to escape predation.

Shell description edit

These small snails, like all the species in the family Physidae, have shells that are sinistral, which means that when the shell is held with the spire pointing up and the aperture facing the viewer, then the aperture is on the left-hand side.

The shells of Physa species have a long and large aperture, a pointed spire, and no operculum. The shells are thin and corneous, and rather transparent.

Species edit

Species in the genus include:

  • Physa aplexoides Yen, 1969
  • Physa arachleica Starobogatov & Prozorova, 1989
  • Physa aravanica Zharnyl'skaya, 1965
  • Physa aridi Mezzalira, 1974 - fossil from Brazil[3]
  • Physa bacca Pan & Zhu, 2007
  • Physa beijiangensis W. Yü & X.-Q. Zhang, 1982
  • Physa boreaui Coquand, 1860
  • Physa bullatula White, 1886
  • Physa canadensis Whiteaves, 1885
  • Physa carletoni Meek, 1873
  • Physa cenomanensis Repelin, 1902
  • Physa cepedaensis Perrilliat & Vega in Perrilliat et al., 2008
  • Physa changleensis Youluo, 1978
  • Physa changzhouensis W. Yü, 1977
  • Physa chippevarum (Taylor, 2003)
  • Physa clarkei Ingersoll, 1875
  • Physa cokevillensis Yen, 1954
  • Physa conispira Yen & Reeside, 1946
  • Physa copei White, 1877
  • Physa dalmatina Küster, 1844
  • Physa debilula Gu in Gu & Wang, 1989
  • Physa delecta Nicolas, 1891
  • Physa doeringi Doello-Jurado, 1927
  • Physa doliolum Matheron, 1843
  • Physa felix White, 1878
  • Physa fontinalis (Linnaeus, 1758) - Fountain bladder snail - type species
  • Physa fuxinensis X.-H. Yu, 1987
  • Physa galei Russell, 1926
  • Physa galloprovincialis Matheron, 1843
  • Physa gaoyouensis W. Yü, 1977
  • Physa gardanensis Matheron, 1843
  • Physa globosa Yen, 1954
  • Physa gracilis Nicolas, 1891
  • Physa hankensis Starobogatov & Prozorova, 1989
  • Physa jingangkouensis H.-Z. Pan, 1983
  • Physa kanabensis White, 1876
  • Physa lacryma F. Sandberger, 1871
  • Physa lacteana Russell, 1935
  • Physa liaoxiensis X.-H. Yu, 1987
  • Physa longiuscula Meek & Hayden, 1856
  • Physa megalochlamys Taylor, 1988 - Cloaked physa
  • Physa meneghinii Sacco, 1886
  • Physa mezzalirai Ghilardi, Carbonaro & Simone, 2011 - fossil from Brazil[4]
  • Physa michaudii Matheron, 1843
  • Physa micra Yen & Reeside, 1946
  • Physa minima Repelin, 1902
  • Physa miqueli Caziot, 1905
  • Physa mirollii Taylor, 2003
  • Physa montanensis Yen, 1951
  • Physa naucatica Zharnyl'skaya, 1965
  • Physa nebrascensis Meek & Hayden, 1856
  • Physa nisidai Suzuki, 1941
  • Physa nucleus Repelin, 1902
  • Physa obtusiconica X.-H. Yu, 1987
  • Physa orientalis G.-X. Zhu, 1980
  • Physa paravitimensis G.-X. Zhu, 1980
  • Physa patula Nicolas, 1891
  • Physa pijiagouensis X.-H. Yu, 1987
  • Physa pumilia Conrad, 1834
  • Physa pygmaea Nicolas, 1891
  • Physa rhomboidea Meek & Hayden, 1856
  • Physa ringentis Youluo, 1978
  • Physa saxarubrensis Russell, 1957
  • Physa secalina Evans & Shumard, 1856
  • Physa shakengensis W. Yü & X.-Q. Zhang, 1982
  • Physa shandongensis H.-Z. Pan, 1983
  • Physa shantungensis Yen, 1969
  • Physa simeyrolsensis Repelin, 1902
  • Physa sinensis Yen, 1969
  • Physa skinneri Taylor, 1954
  • Physa streletzkajae Starobogatov & Budnikova, 1967
  • Physa subacuta Benoist, 1873
  • Physa subcylindrica Repelin, 1902
  • Physa taslei Bourguignat, 1860
  • Physa tenuicostata H.-J. Wang, 1982
  • Physa usitata White, 1895
  • Physa vernalis Taylor et Jokinen, 1984
  • Physa walcotti Yen & Reeside, 1946
  • Physa wealdiana Coquand, 1856
  • Physa wichmanni Parodiz, 1961
  • Physa zhuoxianensis Yü & Pan, 1982
  • Physa sp., Lake Winnipeg Physa, lives in Manitoba in Canada. It was classified as endangered by COSEWIC. COSEWIC recommendation has been forwarded to the Minister of the Environment of Canada and a decision for listing at the List of Wildlife Species at Risk of the Canadian Species at Risk Act was pending in 2005.[5]
Synonyms
  • Physa acuta Draparnaud, 1805, Physa heterostropha (Say, 1817), Physa integra (Haldeman, 1841) and Physa natricina Taylor, 1988 are synonyms of Physella acuta (Draparnaud, 1805)
  • Physa ancillaria (Say, 1825) - Pumpkin Physa:[6] synonym of Physella ancillaria (Say, 1825) (original combination)
  • Physa carolinae Wethington, Wise & Dillon, 2009[7]: synonym of Physella carolinae (Wethington, Wise & Dillon, 2009)
  • Physa concolor Haldeman, 1841: synonym of Physella gyrina (Say, 1821)
  • Physa columbiana (Hemphill, 1890) is a synonym for Physella columbiana (Hemphill, 1890)
  • Physa gyrina (Say, 1821) is a synonym for Physella gyrina (Say, 1821)
  • Physa hordacea (I. Lea, 1864): synonym of Physella hordacea (I. Lea, 1864)
  • Physa jennessi Dall, 1919: synonym of Beringophysa jennessi (Dall, 1919)
  • Physa lordi (Baird, 1863): synonym of Physella lordi (Baird, 1863)
  • Physa marmorata Guilding, 1828[8] - synonyms: Physa mosambiquensis Clessin, 1886 and Physa waterloti:[8] synonym of Stenophysa marmorata (Guilding, 1828)
  • Physa natricina Taylor, 1988: synonym of Physella natricina (D. W. Taylor, 1988)
  • Physa nuttalli I. Lea, 1864: synonym of Physella gyrina (Say, 1821)
  • Physa propingua (Tryon, 1865): synonym of Physella propinqua (Tryon, 1865)
  • Physa sibirica Westerlund, 1876: synonym of Sibirenauta sibirica (Westerlund, 1876)

References edit

  1. ^ Draparnaud J. P. R. (1801). Tableau des mollusques terrestres et fluviatiles de la France. - pp. [1-2], 1-116. Montpellier, Paris. (Renaud; Bossange, Masson & Besson).
  2. ^ MolluscaBase eds. (2021). MolluscaBase. Physa Draparnaud, 1801. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=181551 on 2021-06-26
  3. ^ Simone L. R. L. & Mezzalira S. (1994). "Fossil Molluscs of Brazil". Boletim do Instituto Geológico 11: 1–202.
  4. ^ Ghilardi R. P., Carbonaro F. A. & Simone L. R. L. (2011). "Physa mezzalirai, a new cretaceous basommatophoran from Adamantina formation, Brazil". Strombus 18(1-2): 1-14. abstract.
  5. ^ COSEWIC. 2005. Canadian Species at Risk. Committee on the Status of Endangered Wildlife in Canada. 64 pp., page 13.
  6. ^ "Physella". NatureServe Explorer, accessed 9 April 2010.
  7. ^ Wethington A. R., Wise J. & Dillon R. T. (2009). "Genetic and morphological characterization of the Physidae of South Carolina (Pulmonata: Basommatophora), with description of a new species". The Nautilus 123: 282-292. PDF 2012-08-23 at the Wayback Machine.
  8. ^ a b Appleton C. C. & Dana P. (2005). "Re-examination of Physa mosambiquensis Clessin, 1886 and its relationship with other Aplexinae (Pulmonata: Physidae) reported from Africa". African Invertebrates 46: 71-83. abstract 2010-11-04 at the Wayback Machine
  • Janus, Horst, 1965. ‘’The young specialist looks at land and freshwater molluscs’’, Burke, London

External links edit

  • http://www.animalbase.uni-goettingen.de/zooweb/servlet/AnimalBase/list/species?taxongenus=301
  • Dillon R. T., Wethington A. R. & Lydeard C. (2011). "The evolution of reproductive isolation in a simultaneous hermaphrodite, the freshwater snail Physa". BMC Evolutionary Biology 11: 144. doi:10.1186/1471-2148-11-144.

physa, genus, small, left, handed, sinistral, breathing, freshwater, snails, aquatic, pulmonate, gastropod, mollusks, subfamily, physinae, family, physidae, five, shells, fontinalisscientific, classificationdomain, eukaryotakingdom, animaliaphylum, molluscacla. Physa is a genus of small left handed or sinistral air breathing freshwater snails aquatic pulmonate gastropod mollusks in the subfamily Physinae of the family Physidae 2 PhysaFive shells of Physa fontinalisScientific classificationDomain EukaryotaKingdom AnimaliaPhylum MolluscaClass GastropodaSubclass HeterobranchiaSuperorder HygrophilaSuperfamily LymnaeoideaFamily PhysidaeGenus PhysaDraparnaud 1801 1 Type speciesBulla fontinalis Linnaeus 1758SynonymsLaurentiphysa Taylor 2003 a junior synonym Limnea Physa Draparnaud 1801 Physa Diastropha Gray 1840 Physa Gyrina Schumacher 1817 Physa Laurentiphysa Taylor 2003 a junior synonym Physa Mediterraneophysa Starobogatov amp Budnikova 1976 a junior synonym Physa Physa Draparnaud 1801 Physa Ussuriphysa Starobogatov amp Prozorova 1989 Rivicola Fitzinger 1833 Invalid junior objective synonym of Physa with the same type species These snails eat algae diatoms and detritus Contents 1 Anatomy 2 Shell description 3 Species 4 References 5 External linksAnatomy editMembers of the freshwater pulmonate family Physidae possess a complex of muscles that is unique amongst gastropods This complex was given the name physid musculature The physid musculature has two main components the physid muscle sensu stricto and the fan muscle The physid musculature is responsible for a unique ability of physids to rapidly flick their shells from side to side a reaction that frequently enables them to escape predation Shell description editThese small snails like all the species in the family Physidae have shells that are sinistral which means that when the shell is held with the spire pointing up and the aperture facing the viewer then the aperture is on the left hand side The shells of Physa species have a long and large aperture a pointed spire and no operculum The shells are thin and corneous and rather transparent Species editSpecies in the genus include Physa aplexoides Yen 1969 Physa arachleica Starobogatov amp Prozorova 1989 Physa aravanica Zharnyl skaya 1965 Physa aridi Mezzalira 1974 fossil from Brazil 3 Physa bacca Pan amp Zhu 2007 Physa beijiangensis W Yu amp X Q Zhang 1982 Physa boreaui Coquand 1860 Physa bullatula White 1886 Physa canadensis Whiteaves 1885 Physa carletoni Meek 1873 Physa cenomanensis Repelin 1902 Physa cepedaensis Perrilliat amp Vega in Perrilliat et al 2008 Physa changleensis Youluo 1978 Physa changzhouensis W Yu 1977 Physa chippevarum Taylor 2003 Physa clarkei Ingersoll 1875 Physa cokevillensis Yen 1954 Physa conispira Yen amp Reeside 1946 Physa copei White 1877 Physa dalmatina Kuster 1844 Physa debilula Gu in Gu amp Wang 1989 Physa delecta Nicolas 1891 Physa doeringi Doello Jurado 1927 Physa doliolum Matheron 1843 Physa felix White 1878 Physa fontinalis Linnaeus 1758 Fountain bladder snail type species Physa fuxinensis X H Yu 1987 Physa galei Russell 1926 Physa galloprovincialis Matheron 1843 Physa gaoyouensis W Yu 1977 Physa gardanensis Matheron 1843 Physa globosa Yen 1954 Physa gracilis Nicolas 1891 Physa hankensis Starobogatov amp Prozorova 1989 Physa jingangkouensis H Z Pan 1983 Physa kanabensis White 1876 Physa lacryma F Sandberger 1871 Physa lacteana Russell 1935 Physa liaoxiensis X H Yu 1987 Physa longiuscula Meek amp Hayden 1856 Physa megalochlamys Taylor 1988 Cloaked physa Physa meneghinii Sacco 1886 Physa mezzalirai Ghilardi Carbonaro amp Simone 2011 fossil from Brazil 4 Physa michaudii Matheron 1843 Physa micra Yen amp Reeside 1946 Physa minima Repelin 1902 Physa miqueli Caziot 1905 Physa mirollii Taylor 2003 Physa montanensis Yen 1951 Physa naucatica Zharnyl skaya 1965 Physa nebrascensis Meek amp Hayden 1856 Physa nisidai Suzuki 1941 Physa nucleus Repelin 1902 Physa obtusiconica X H Yu 1987 Physa orientalis G X Zhu 1980 Physa paravitimensis G X Zhu 1980 Physa patula Nicolas 1891 Physa pijiagouensis X H Yu 1987 Physa pumilia Conrad 1834 Physa pygmaea Nicolas 1891 Physa rhomboidea Meek amp Hayden 1856 Physa ringentis Youluo 1978 Physa saxarubrensis Russell 1957 Physa secalina Evans amp Shumard 1856 Physa shakengensis W Yu amp X Q Zhang 1982 Physa shandongensis H Z Pan 1983 Physa shantungensis Yen 1969 Physa simeyrolsensis Repelin 1902 Physa sinensis Yen 1969 Physa skinneri Taylor 1954 Physa streletzkajae Starobogatov amp Budnikova 1967 Physa subacuta Benoist 1873 Physa subcylindrica Repelin 1902 Physa taslei Bourguignat 1860 Physa tenuicostata H J Wang 1982 Physa usitata White 1895 Physa vernalis Taylor et Jokinen 1984 Physa walcotti Yen amp Reeside 1946 Physa wealdiana Coquand 1856 Physa wichmanni Parodiz 1961 Physa zhuoxianensis Yu amp Pan 1982 Physa sp Lake Winnipeg Physa lives in Manitoba in Canada It was classified as endangered by COSEWIC COSEWIC recommendation has been forwarded to the Minister of the Environment of Canada and a decision for listing at the List of Wildlife Species at Risk of the Canadian Species at Risk Act was pending in 2005 5 SynonymsPhysa acuta Draparnaud 1805 Physa heterostropha Say 1817 Physa integra Haldeman 1841 and Physa natricina Taylor 1988 are synonyms of Physella acuta Draparnaud 1805 Physa ancillaria Say 1825 Pumpkin Physa 6 synonym of Physella ancillaria Say 1825 original combination Physa carolinae Wethington Wise amp Dillon 2009 7 synonym of Physella carolinae Wethington Wise amp Dillon 2009 Physa concolor Haldeman 1841 synonym of Physella gyrina Say 1821 Physa columbiana Hemphill 1890 is a synonym for Physella columbiana Hemphill 1890 Physa gyrina Say 1821 is a synonym for Physella gyrina Say 1821 Physa hordacea I Lea 1864 synonym of Physella hordacea I Lea 1864 Physa jennessi Dall 1919 synonym of Beringophysa jennessi Dall 1919 Physa lordi Baird 1863 synonym of Physella lordi Baird 1863 Physa marmorata Guilding 1828 8 synonyms Physa mosambiquensis Clessin 1886 and Physa waterloti 8 synonym of Stenophysa marmorata Guilding 1828 Physa natricina Taylor 1988 synonym of Physella natricina D W Taylor 1988 Physa nuttalli I Lea 1864 synonym of Physella gyrina Say 1821 Physa propingua Tryon 1865 synonym of Physella propinqua Tryon 1865 Physa sibirica Westerlund 1876 synonym of Sibirenauta sibirica Westerlund 1876 References edit Draparnaud J P R 1801 Tableau des mollusques terrestres et fluviatiles de la France pp 1 2 1 116 Montpellier Paris Renaud Bossange Masson amp Besson MolluscaBase eds 2021 MolluscaBase Physa Draparnaud 1801 Accessed through World Register of Marine Species at http www marinespecies org aphia php p taxdetails amp id 181551 on 2021 06 26 Simone L R L amp Mezzalira S 1994 Fossil Molluscs of Brazil Boletim do Instituto Geologico 11 1 202 Ghilardi R P Carbonaro F A amp Simone L R L 2011 Physa mezzalirai a new cretaceous basommatophoran from Adamantina formation Brazil Strombus 18 1 2 1 14 abstract COSEWIC 2005 Canadian Species at Risk Committee on the Status of Endangered Wildlife in Canada 64 pp page 13 Physella NatureServe Explorer accessed 9 April 2010 Wethington A R Wise J amp Dillon R T 2009 Genetic and morphological characterization of the Physidae of South Carolina Pulmonata Basommatophora with description of a new species The Nautilus 123 282 292 PDF Archived 2012 08 23 at the Wayback Machine a b Appleton C C amp Dana P 2005 Re examination of Physa mosambiquensis Clessin 1886 and its relationship with other Aplexinae Pulmonata Physidae reported from Africa African Invertebrates 46 71 83 abstract Archived 2010 11 04 at the Wayback Machine Janus Horst 1965 The young specialist looks at land and freshwater molluscs Burke LondonExternal links edit nbsp Wikimedia Commons has media related to Physa http www animalbase uni goettingen de zooweb servlet AnimalBase list species taxongenus 301 Dillon R T Wethington A R amp Lydeard C 2011 The evolution of reproductive isolation in a simultaneous hermaphrodite the freshwater snail Physa BMC Evolutionary Biology 11 144 doi 10 1186 1471 2148 11 144 Retrieved from https en wikipedia org w index php title Physa amp oldid 1180455610, wikipedia, wiki, book, books, library,

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