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Marbled cat

The marbled cat (Pardofelis marmorata) is a small wild cat native from the eastern Himalayas to Southeast Asia, where it inhabits forests up to an elevation of 2,500 m (8,200 ft). As it is present in a large range, it has been listed as Near Threatened on the IUCN Red List since 2015.[2]

Marbled cat
A marbled cat in Danum Valley, Borneo
CITES Appendix I (CITES)[2]
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Felinae
Genus: Pardofelis
Species:
P. marmorata[1]
Binomial name
Pardofelis marmorata[1]
(Martin, 1836)
Subspecies
  • P. m. charltoni
  • P. m. marmorata
Distribution of marbled cat, 2016[2]

The marbled cat is closely related to the Asian golden cat (Catopuma temminckii) and the bay cat (C. badia), all of which diverged from other felids about 9.4 million years ago.[3]

Characteristics edit

The marbled cat is similar in size to a domestic cat, but has rounded ears and a very long tail that is as long as the cat's head and body. The ground colour of its long fur varies from brownish-grey to ochreous brown above and greyish to buff below. It is patterned with black stripes on the short and round head, on the neck and back. On the tail, limbs and underbelly it has solid spots. On the flanks it has irregular dark-edged blotches that fuse to dark areas and look like a 'marbled' pattern. Its paws are webbed between the digits and are completely sheathed.[4] Its coat is thick and soft. Spots on the forehead and crown merge into narrow longitudinal stripes on the neck, and irregular stripes on the back. The legs and underparts are patterned with black dots, and the tail is marked with black spots proximally and rings distally. It has large feet and unusually large canine teeth, resembling those of the big cats, although these appear to be the result of parallel evolution. Marbled cats range from 45 to 62 cm (18 to 24 in) in head-body length with a 35 to 55 cm (14 to 22 in) long and thickly furred tail that indicates the cat's adaptation to an arboreal lifestyle, where the tail is used as a counterbalance. Recorded weights vary between 2 and 5 kg (4.4 and 11.0 lb).[5]

Distribution and habitat edit

The marbled cat occurs along the eastern Himalayan foothills and in tropical Indomalaya eastward into southwest China, and on the islands of Sumatra and Borneo. It is primarily associated with moist and mixed deciduous-evergreen tropical forests. Its distribution in India is confined to the north-eastern forests.[2]

In eastern Nepal, a marbled cat was recorded for the first time in January 2018, outside a protected area in the Kangchenjunga landscape at an altitude of 2,750 m (9,020 ft).[6]

In northeast India, marbled cats were recorded in Eaglenest Wildlife Sanctuary, Dampa and Pakke Tiger Reserves, Balpakram-Baghmara landscape and Singchung-Bugun Village Community Reserve in Arunachal Pradesh between January 2013 and March 2018.[7]

In Bhutan, it has been recorded in Royal Manas National Park, and in broadleaved and mixed conifer forests at elevations up to 3,810 m (12,500 ft) in Jigme Dorji National Park and Wangchuck Centennial National Park.[8][9][10]

In Thailand, it was recorded in a hill evergreen bamboo mixed forest in Phu Khieu Wildlife Sanctuary.[11]

In Borneo, it has also been recorded in peat swamp forest.[12] The population size of the marbled cat is not well understood. Few records were obtained during camera-trapping surveys throughout much of its range. In three areas in Sabah, the population density was estimated at 7.1 to 19.6 individuals per 100 km2 (39 sq mi), an estimate that may be higher than elsewhere in the cat's range.[13] In Kalimantan, marbled cats were recorded in mixed swamp forest and tall interior forest at altitudes below 20 m (66 ft) in the vicinity of Sabangau National Park between 2008 and 2018.[14]

Behaviour and ecology edit

Marbled cats recorded in northeastern India and Kalimantan on Borneo were active by day.[7][14]

The first-ever radio-tracked marbled cat had an overall home range of 5.8 km2 (2.2 sq mi) at an elevation of 1,000 to 1,200 m (3,300 to 3,900 ft) and was active primarily during nocturnal and crepuscular times.[11] Marbled cats recorded in northeast India were active during the day with activity peaks around noon.[7]

Forest canopies probably provide the marbled cat with much of its prey: birds, squirrels and other rodents, and reptiles.[5] In the Bukit Barisan Selatan National Park, a marbled cat was observed in a dense forest patch in an area also used by siamang.[15] In Thailand, one individual has been observed in Phu Khieo Wildlife Sanctuary preying on a Phayre's leaf monkey.[16]

A few marbled cats have been bred in captivity, with gestation estimated to be 66 to 82 days. In the few recorded instances, two kittens were born in each litter, and weighed from 61 to 85 g (2.2 to 3.0 oz). Their eyes open at around 12 days, and the kittens begin to take solid food at two months, around the time that they begin actively climbing. Marbled cats reach sexual maturity at 21 or 22 months of age, and have lived for up to 12 years in captivity.[5]

Threats edit

The primary threat to the marbled cat is loss and degradation of forest as it depends on large tracts of intact forest.[17] Forest loss is continuing across its range due to logging and expansion of human settlements and agriculture.[2] Indiscriminate snaring is prevalent throughout much of its range, and also likely poses a major threat. It is valued for its skin, meat, and bones, but infrequently observed in the illegal Asian wildlife trade.[2] During a survey in the Lower Subansiri District of Arunachal Pradesh, a marbled cat was encountered that had been killed by a local hunter for a festival celebrated by the indigenous Apatani people in March and April every year. The dead cat was used in a ceremony, and its blood was sacrificed to the deity for goodwill of their family and for ensuring a good harvest, protection from wildlife, disease and pest.[18]

Conservation edit

Pardofelis marmorata is included in CITES Appendix I and protected over parts of its range. Hunting is prohibited in Bangladesh, Cambodia, Yunnan, India, Indonesia, Malaysia, Myanmar, Nepal, and Thailand. Hunting is regulated in Laos and Singapore. In Bhutan and Brunei, the marbled cat is not legally protected outside protected areas. No information about protection status is available from Cambodia and Vietnam.[19]

Taxonomy edit

Felis marmorata was the scientific name proposed by William Charles Linnaeus Martin in 1836 for a skin of a male marbled cat from Java or Sumatra.[20]Felis longicaudata proposed by Henri Marie Ducrotay de Blainville in 1843 was a zoological specimen from India or Cochinchina.[21]Felis charltoni proposed by John Edward Gray in 1846 was a specimen from Darjeeling.[22] The generic name Pardofelis was proposed by Nikolai Severtzov in 1858.[23]

At present, two subspecies are recognized as valid:[24]

Phylogeny edit

The marbled cat was once considered to belong to the pantherine lineage of cats.[25] But results of a phylogenetic analysis indicate that it forms an evolutionary lineage with the Asian golden cat (Catopuma temminckii) and the bay cat (C. badia) that genetically diverged about 12.77 to 7.36 million years ago. The marbled cat diverged from this lineage about 8.42 to 4.27 million years ago.[3][26]

References edit

  1. ^ Wozencraft, W. C. (2005). "Species Pardofelis marmorata". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 542. ISBN 978-0-8018-8221-0. OCLC 62265494.
  2. ^ a b c d e f g Ross, J.; Brodie, J.; Cheyne, S.; Datta, A.; Hearn, A.; Loken, B.; Lynam, A.J.; McCarthy, J.; Phan, C.; Rasphone, A.; Singh, P. & Wilting, A. (2016). "Pardofelis marmorata". IUCN Red List of Threatened Species. 2016: e.T16218A97164299. doi:10.2305/IUCN.UK.2016-1.RLTS.T16218A97164299.en. Retrieved 16 January 2022.
  3. ^ a b Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E. & O'Brien, S. J. (2006). "The Late Miocene Radiation of Modern Felidae: A Genetic Assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID 16400146. S2CID 41672825.
  4. ^ Pocock, R. I. (1939). "Genus Pardofelis Severtzow". The Fauna of British India, including Ceylon and Burma. Mammalia – Volume 1. London: Taylor and Francis. pp. 253−258.
  5. ^ a b c Sunquist, M. & Sunquist, F. (2002). "Marbled cat Pardofelis marmorata (Martin 1837)". Wild Cats of the World. Chicago: University of Chicago Press. pp. 373–376. ISBN 978-0-226-77999-7.
  6. ^ Lama, S. T.; Ross, J. G.; Bista, D.; Sherpa, A. P.; Regmi, G. R.; Suwal, M. K.; Sherpa, P.; Weerman, J.; Lama, S. S.; Thapa, M.; Poudyal, L. P. (2019). "First photographic record of marbled cat Pardofelis marmorata Martin, 1837 (Mammalia, Carnivora, Felidae) in Nepal". Nature Conservation. 32: 19−34. doi:10.3897/natureconservation.32.29740. hdl:10182/10670.
  7. ^ a b c Mukherjee, S.; Singh, P.; Silva, A.; Ri, C.; Kakati, K.; Borah, B.; Tapi, T.; Kadur, S.; Choudhary, P.; Srikant, S.; Nadig, S.; Navya, R.; Björklund, R.; Ramakrishnan, U. (2019). "Activity patterns of the small and medium felid (Mammalia: Carnivora: Felidae) guild in northeastern India". Journal of Threatened Taxa. 11 (4): 13432−13447. doi:10.11609/jott.4662.11.4.13432-13447.
  8. ^ Tempa, T.; Hebblewhite, M.; Mills, L.S.; Wangchuk, T.R.; Norbu, N.; Wangchuk, T.; Nidup, T.; Dhendup, P.; Wangchuk, D.; Wangdi, Y. & Dorji, T. (2013). "Royal Manas National Park: A hotspot for wild felids, Bhutan". Oryx. 47 (2): 207–210. doi:10.1017/s0030605312001317.
  9. ^ Thinley, P.; Morreale, S.J.; Curtis, P.D.; Lassoie, J.P.; Dorji, T.; Phuntsho, S. & Dorji, N. (2015). "Diversity, occupancy, and spatio-temporal occurrences of mammalian predators in Bhutan's Jigme Dorji National Park". Bhutan Journal of Natural Resources & Development. 2 (1): 19–27.
  10. ^ Dhendup, T. (2016). "Notes on the occurrence of Marbled Cats at high altitudes in Bhutan" (PDF). NeBIO. 7 (2): 35–37.
  11. ^ a b Grassman, L. I. Jr.; Tewes, M. E. (2000). "Marbled cat in northeastern Thailand". Cat News. 33: 24.
  12. ^ Cheyne, S. M.; Macdonald, D. W. (2010). "Marbled cat in Sabangau peat-swamp forest, Indonesian Borneo" (PDF). Cat News. 52: 11.
  13. ^ Hearn, A. J.; Ross, J.; Bernard, H.; Bakar, S. A.; Hunter, Luke T. B.; Macdonald, D. W. (2016). "The First Estimates of Marbled Cat Pardofelis marmorata Population Density from Bornean Primary and Selectively Logged Forest". PLOS ONE. 11 (3): e0151046. Bibcode:2016PLoSO..1151046H. doi:10.1371/journal.pone.0151046. PMC 4805203. PMID 27007219.
  14. ^ a b Jeffers, K. A.; Adul; Cheyne, S. M. (2019). "Small cat surveys: 10 years of data from Central Kalimantan, Indonesian Borneo". Journal of Threatened Taxa. 11 (4): 13478–13491. doi:10.11609/jott.4466.11.4.13478-13491.
  15. ^ Morino, L. (2009). "Observation of a wild marbled cat in Sumatra". Cat News (50): 20.
  16. ^ Borries, C.; Primeau, Z. M.; Ossi-Lupo, K.; Dtubpraserit, S. & Koenig, A. (2014). "Possible predation attempt by a marbled cat on a juvenile Phayre's leaf monkey". Raffles Bulletin of Zoology (62): 561–565.
  17. ^ Hendry, A.; Amir, Z.; Decoeur, H.; Mendes, C. P.; Moore, J. H; Sovie, A.; Luskin, M. S. (January 2023). "Marbled cats in Southeast Asia: Are diurnal and semi-arboreal felids at greater risk from human disturbances?". Ecosphere. 14 (1): e4338. doi:10.1002/ecs2.4338.
  18. ^ Selvan, K. M.; Gopi, G. V.; Habib, B.; Lyngdoh, S. (2013). "Hunting record of endangered Marbled Cat Pardofelis marmorata in the Ziro Valley of Lower Subansiri, Arunachal Pradesh, India". Journal of Threatened Taxa. 5 (1): 3583–3584. doi:10.11609/JoTT.o3208.100.
  19. ^ Nowell, K. and Jackson, P. (1996). Marbled Cat Felis marmorata. in: Wild Cats. Status Survey and Conservation Action Plan. IUCN/SSC Cat Specialist Group, Gland, Switzerland and Cambridge, UK.
  20. ^ Martin, W. C. (1836). "Description of a new species of Felis". Proceedings of the Zoological Society of London. IV (XLVII): 107–108.
  21. ^ Blainville, H. M. D. (1843). "Os du squelette des Felis". Ostéographie ou description iconographique comparée du squelette et du système dentaire des cinques classes d'animaux vertébrés récents et fossils pour servir de base a la zoologie et la géologie. Volume 2: Mammifères. Carnassiers. Paris: Arthus Bertrand. pp. 1–196.
  22. ^ Gray, J. E. (1846). "New species of Mammalia". The Annals and Magazine of Natural History; Zoology, Botany, and Geology. 18 (118): 211−212. doi:10.1080/037454809494412.
  23. ^ Severtzow, M. N. (1858). "Notice sur la classification multisériale des Carnivores, spécialement des Félidés, et les études de zoologie générale qui s'y rattachent". Revue et Magasin de Zoologie Pure et Appliquée. X: 385–396.
  24. ^ Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O’Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.; Nowell, K.; Timmons, Z.; Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group" (PDF). Cat News (Special Issue 11): 34−35.
  25. ^ Hemmer, H. (1978). "The evolutionary systematics of living Felidae: Present status and current problems". Carnivore. 1: 71–79.
  26. ^ Werdelin, L.; Yamaguchi, N.; Johnson, W. E.; O'Brien, S. J. (2010). "Phylogeny and evolution of cats (Felidae)". In Macdonald, D. W.; Loveridge, A. J. (eds.). Biology and Conservation of Wild Felids. Oxford, UK: Oxford University Press. pp. 59–82. ISBN 978-0-19-923445-5.

External links edit

  • "Marbled cat Pardofelis marmorata". IUCN/SSC Cat Specialist Group.

marbled, marbled, pardofelis, marmorata, small, wild, native, from, eastern, himalayas, southeast, asia, where, inhabits, forests, elevation, present, large, range, been, listed, near, threatened, iucn, list, since, 2015, marbled, danum, valley, borneoconserva. The marbled cat Pardofelis marmorata is a small wild cat native from the eastern Himalayas to Southeast Asia where it inhabits forests up to an elevation of 2 500 m 8 200 ft As it is present in a large range it has been listed as Near Threatened on the IUCN Red List since 2015 2 Marbled catA marbled cat in Danum Valley BorneoConservation statusNear Threatened IUCN 3 1 2 CITES Appendix I CITES 2 Scientific classificationDomain EukaryotaKingdom AnimaliaPhylum ChordataClass MammaliaOrder CarnivoraSuborder FeliformiaFamily FelidaeSubfamily FelinaeGenus PardofelisSpecies P marmorata 1 Binomial namePardofelis marmorata 1 Martin 1836 SubspeciesP m charltoni P m marmorataDistribution of marbled cat 2016 2 The marbled cat is closely related to the Asian golden cat Catopuma temminckii and the bay cat C badia all of which diverged from other felids about 9 4 million years ago 3 Contents 1 Characteristics 2 Distribution and habitat 3 Behaviour and ecology 4 Threats 5 Conservation 6 Taxonomy 6 1 Phylogeny 7 References 8 External linksCharacteristics editThe marbled cat is similar in size to a domestic cat but has rounded ears and a very long tail that is as long as the cat s head and body The ground colour of its long fur varies from brownish grey to ochreous brown above and greyish to buff below It is patterned with black stripes on the short and round head on the neck and back On the tail limbs and underbelly it has solid spots On the flanks it has irregular dark edged blotches that fuse to dark areas and look like a marbled pattern Its paws are webbed between the digits and are completely sheathed 4 Its coat is thick and soft Spots on the forehead and crown merge into narrow longitudinal stripes on the neck and irregular stripes on the back The legs and underparts are patterned with black dots and the tail is marked with black spots proximally and rings distally It has large feet and unusually large canine teeth resembling those of the big cats although these appear to be the result of parallel evolution Marbled cats range from 45 to 62 cm 18 to 24 in in head body length with a 35 to 55 cm 14 to 22 in long and thickly furred tail that indicates the cat s adaptation to an arboreal lifestyle where the tail is used as a counterbalance Recorded weights vary between 2 and 5 kg 4 4 and 11 0 lb 5 Distribution and habitat editThe marbled cat occurs along the eastern Himalayan foothills and in tropical Indomalaya eastward into southwest China and on the islands of Sumatra and Borneo It is primarily associated with moist and mixed deciduous evergreen tropical forests Its distribution in India is confined to the north eastern forests 2 In eastern Nepal a marbled cat was recorded for the first time in January 2018 outside a protected area in the Kangchenjunga landscape at an altitude of 2 750 m 9 020 ft 6 In northeast India marbled cats were recorded in Eaglenest Wildlife Sanctuary Dampa and Pakke Tiger Reserves Balpakram Baghmara landscape and Singchung Bugun Village Community Reserve in Arunachal Pradesh between January 2013 and March 2018 7 In Bhutan it has been recorded in Royal Manas National Park and in broadleaved and mixed conifer forests at elevations up to 3 810 m 12 500 ft in Jigme Dorji National Park and Wangchuck Centennial National Park 8 9 10 In Thailand it was recorded in a hill evergreen bamboo mixed forest in Phu Khieu Wildlife Sanctuary 11 In Borneo it has also been recorded in peat swamp forest 12 The population size of the marbled cat is not well understood Few records were obtained during camera trapping surveys throughout much of its range In three areas in Sabah the population density was estimated at 7 1 to 19 6 individuals per 100 km2 39 sq mi an estimate that may be higher than elsewhere in the cat s range 13 In Kalimantan marbled cats were recorded in mixed swamp forest and tall interior forest at altitudes below 20 m 66 ft in the vicinity of Sabangau National Park between 2008 and 2018 14 Behaviour and ecology editMarbled cats recorded in northeastern India and Kalimantan on Borneo were active by day 7 14 The first ever radio tracked marbled cat had an overall home range of 5 8 km2 2 2 sq mi at an elevation of 1 000 to 1 200 m 3 300 to 3 900 ft and was active primarily during nocturnal and crepuscular times 11 Marbled cats recorded in northeast India were active during the day with activity peaks around noon 7 Forest canopies probably provide the marbled cat with much of its prey birds squirrels and other rodents and reptiles 5 In the Bukit Barisan Selatan National Park a marbled cat was observed in a dense forest patch in an area also used by siamang 15 In Thailand one individual has been observed in Phu Khieo Wildlife Sanctuary preying on a Phayre s leaf monkey 16 A few marbled cats have been bred in captivity with gestation estimated to be 66 to 82 days In the few recorded instances two kittens were born in each litter and weighed from 61 to 85 g 2 2 to 3 0 oz Their eyes open at around 12 days and the kittens begin to take solid food at two months around the time that they begin actively climbing Marbled cats reach sexual maturity at 21 or 22 months of age and have lived for up to 12 years in captivity 5 Threats editThe primary threat to the marbled cat is loss and degradation of forest as it depends on large tracts of intact forest 17 Forest loss is continuing across its range due to logging and expansion of human settlements and agriculture 2 Indiscriminate snaring is prevalent throughout much of its range and also likely poses a major threat It is valued for its skin meat and bones but infrequently observed in the illegal Asian wildlife trade 2 During a survey in the Lower Subansiri District of Arunachal Pradesh a marbled cat was encountered that had been killed by a local hunter for a festival celebrated by the indigenous Apatani people in March and April every year The dead cat was used in a ceremony and its blood was sacrificed to the deity for goodwill of their family and for ensuring a good harvest protection from wildlife disease and pest 18 Conservation editPardofelis marmorata is included in CITES Appendix I and protected over parts of its range Hunting is prohibited in Bangladesh Cambodia Yunnan India Indonesia Malaysia Myanmar Nepal and Thailand Hunting is regulated in Laos and Singapore In Bhutan and Brunei the marbled cat is not legally protected outside protected areas No information about protection status is available from Cambodia and Vietnam 19 Taxonomy editFelis marmorata was the scientific name proposed by William Charles Linnaeus Martin in 1836 for a skin of a male marbled cat from Java or Sumatra 20 Felis longicaudata proposed by Henri Marie Ducrotay de Blainville in 1843 was a zoological specimen from India or Cochinchina 21 Felis charltoni proposed by John Edward Gray in 1846 was a specimen from Darjeeling 22 The generic name Pardofelis was proposed by Nikolai Severtzov in 1858 23 At present two subspecies are recognized as valid 24 P marmorata marmorata Martin 1836 from the Malay Peninsula to Sumatra and Borneo P marmorata longicaudata Blainville 1843 from Nepal to north of the Isthmus of KraPhylogeny edit The marbled cat was once considered to belong to the pantherine lineage of cats 25 But results of a phylogenetic analysis indicate that it forms an evolutionary lineage with the Asian golden cat Catopuma temminckii and the bay cat C badia that genetically diverged about 12 77 to 7 36 million years ago The marbled cat diverged from this lineage about 8 42 to 4 27 million years ago 3 26 References edit Wozencraft W C 2005 Species Pardofelis marmorata In Wilson D E Reeder D M eds Mammal Species of the World A Taxonomic and Geographic Reference 3rd ed Johns Hopkins University Press p 542 ISBN 978 0 8018 8221 0 OCLC 62265494 a b c d e f g Ross J Brodie J Cheyne S Datta A Hearn A Loken B Lynam A J McCarthy J Phan C Rasphone A Singh P amp Wilting A 2016 Pardofelis marmorata IUCN Red List of Threatened Species 2016 e T16218A97164299 doi 10 2305 IUCN UK 2016 1 RLTS T16218A97164299 en Retrieved 16 January 2022 a b Johnson W E Eizirik E Pecon Slattery J Murphy W J Antunes A Teeling E amp O Brien S J 2006 The Late Miocene Radiation of Modern Felidae A Genetic Assessment Science 311 5757 73 77 Bibcode 2006Sci 311 73J doi 10 1126 science 1122277 PMID 16400146 S2CID 41672825 Pocock R I 1939 Genus Pardofelis Severtzow The Fauna of British India including Ceylon and Burma Mammalia Volume 1 London Taylor and Francis pp 253 258 a b c Sunquist M amp Sunquist F 2002 Marbled cat Pardofelis marmorata Martin 1837 Wild Cats of the World Chicago University of Chicago Press pp 373 376 ISBN 978 0 226 77999 7 Lama S T Ross J G Bista D Sherpa A P Regmi G R Suwal M K Sherpa P Weerman J Lama S S Thapa M Poudyal L P 2019 First photographic record of marbled cat Pardofelis marmorata Martin 1837 Mammalia Carnivora Felidae in Nepal Nature Conservation 32 19 34 doi 10 3897 natureconservation 32 29740 hdl 10182 10670 a b c Mukherjee S Singh P Silva A Ri C Kakati K Borah B Tapi T Kadur S Choudhary P Srikant S Nadig S Navya R Bjorklund R Ramakrishnan U 2019 Activity patterns of the small and medium felid Mammalia Carnivora Felidae guild in northeastern India Journal of Threatened Taxa 11 4 13432 13447 doi 10 11609 jott 4662 11 4 13432 13447 Tempa T Hebblewhite M Mills L S Wangchuk T R Norbu N Wangchuk T Nidup T Dhendup P Wangchuk D Wangdi Y amp Dorji T 2013 Royal Manas National Park A hotspot for wild felids Bhutan Oryx 47 2 207 210 doi 10 1017 s0030605312001317 Thinley P Morreale S J Curtis P D Lassoie J P Dorji T Phuntsho S amp Dorji N 2015 Diversity occupancy and spatio temporal occurrences of mammalian predators in Bhutan s Jigme Dorji National Park Bhutan Journal of Natural Resources amp Development 2 1 19 27 Dhendup T 2016 Notes on the occurrence of Marbled Cats at high altitudes in Bhutan PDF NeBIO 7 2 35 37 a b Grassman L I Jr Tewes M E 2000 Marbled cat in northeastern Thailand Cat News 33 24 Cheyne S M Macdonald D W 2010 Marbled cat in Sabangau peat swamp forest Indonesian Borneo PDF Cat News 52 11 Hearn A J Ross J Bernard H Bakar S A Hunter Luke T B Macdonald D W 2016 The First Estimates of Marbled Cat Pardofelis marmorata Population Density from Bornean Primary and Selectively Logged Forest PLOS ONE 11 3 e0151046 Bibcode 2016PLoSO 1151046H doi 10 1371 journal pone 0151046 PMC 4805203 PMID 27007219 a b Jeffers K A Adul Cheyne S M 2019 Small cat surveys 10 years of data from Central Kalimantan Indonesian Borneo Journal of Threatened Taxa 11 4 13478 13491 doi 10 11609 jott 4466 11 4 13478 13491 Morino L 2009 Observation of a wild marbled cat in Sumatra Cat News 50 20 Borries C Primeau Z M Ossi Lupo K Dtubpraserit S amp Koenig A 2014 Possible predation attempt by a marbled cat on a juvenile Phayre s leaf monkey Raffles Bulletin of Zoology 62 561 565 Hendry A Amir Z Decoeur H Mendes C P Moore J H Sovie A Luskin M S January 2023 Marbled cats in Southeast Asia Are diurnal and semi arboreal felids at greater risk from human disturbances Ecosphere 14 1 e4338 doi 10 1002 ecs2 4338 Selvan K M Gopi G V Habib B Lyngdoh S 2013 Hunting record of endangered Marbled Cat Pardofelis marmorata in the Ziro Valley of Lower Subansiri Arunachal Pradesh India Journal of Threatened Taxa 5 1 3583 3584 doi 10 11609 JoTT o3208 100 Nowell K and Jackson P 1996 Marbled Cat Felis marmorata in Wild Cats Status Survey and Conservation Action Plan IUCN SSC Cat Specialist Group Gland Switzerland and Cambridge UK Martin W C 1836 Description of a new species of Felis Proceedings of the Zoological Society of London IV XLVII 107 108 Blainville H M D 1843 Os du squelette des Felis Osteographie ou description iconographique comparee du squelette et du systeme dentaire des cinques classes d animaux vertebres recents et fossils pour servir de base a la zoologie et la geologie Volume 2 Mammiferes Carnassiers Paris Arthus Bertrand pp 1 196 Gray J E 1846 New species of Mammalia The Annals and Magazine of Natural History Zoology Botany and Geology 18 118 211 212 doi 10 1080 037454809494412 Severtzow M N 1858 Notice sur la classification multiseriale des Carnivores specialement des Felides et les etudes de zoologie generale qui s y rattachent Revue et Magasin de Zoologie Pure et Appliquee X 385 396 Kitchener A C Breitenmoser Wursten C Eizirik E Gentry A Werdelin L Wilting A Yamaguchi N Abramov A V Christiansen P Driscoll C Duckworth J W Johnson W Luo S J Meijaard E O Donoghue P Sanderson J Seymour K Bruford M Groves C Hoffmann M Nowell K Timmons Z Tobe S 2017 A revised taxonomy of the Felidae The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group PDF Cat News Special Issue 11 34 35 Hemmer H 1978 The evolutionary systematics of living Felidae Present status and current problems Carnivore 1 71 79 Werdelin L Yamaguchi N Johnson W E O Brien S J 2010 Phylogeny and evolution of cats Felidae In Macdonald D W Loveridge A J eds Biology and Conservation of Wild Felids Oxford UK Oxford University Press pp 59 82 ISBN 978 0 19 923445 5 External links edit nbsp Wikispecies has information related to Pardofelis marmorata nbsp Wikimedia Commons has media related to Pardofelis marmorata Marbled cat Pardofelis marmorata IUCN SSC Cat Specialist Group Retrieved from https en wikipedia org w index php title Marbled cat amp oldid 1195324461, wikipedia, wiki, book, books, library,

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