List of pterosaur genera

This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered pterosaurian. The list currently includes 263 genera.

Scope and terminology

There is no official, canonical list of pterosaur genera, but the most thorough attempts can be found at the Pterosauria section of Mikko Haaramo's Phylogeny Archive,^[1] the Genus Index at Mike Hanson's The Pterosauria,^[2] supplemented by the Pterosaur Species List,^[3] and in the fourth supplement of Donald F. Glut's Dinosaurs: The Encyclopedia series.^[4]

Authors and year

The authors column lists the authors of the formal description responsible for the erection of the genus listed. They are not necessarily the same as the authors of the type species as sometimes a species from one genus is determined sufficiently distinct to warrant the erection of a new genus to house it. If this is the case, only the latter authors will be listed. The year column notes the year the description was published.

Status

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

• Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
• Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
• Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
• Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

Age

The age column denotes the epoch of geologic time to which the fossils date. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as age N/A because there was never a time in which a pterosaur by that generic name actually lived.

Location and notes

The location column designates the geographic region where remains of the relevant genus have been found. The regions used are continents except in the case of smaller landmasses (e.g. Cuba.) Political bodies, being non-existent in the Mesozoic are not used to indicate genera locations. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as location N/A because there was never a place in which a pterosaur by that generic name actually lived. The notes column is a collection of annotations on the scientific significance and taxonomic history of listed genera, as well as elaborations on the information presented in other columns.

The list

Genera

Genus Authors Year Status Age Location Notes Aerodactylus^[5] Vidovic Martill 2014 Valid Late Jurassic Europe Aerodraco^[6] Holgado Pêgas 2020 Valid Early Cretaceous Europe Aerotitan^[7] Novas et al. 2012 Valid Late Cretaceous S. America Aetodactylus^[8] Myers 2010 Valid Late Cretaceous N. America Afrotapejara^[9] Martill et al 2020 Valid Late Cretaceous Africa Aidachar^[10] Nesov 1981 Misidentification N/A N/A Actually a teleost fish whose remains were originally mistaken for jaw fragments of a ctenochasmatid. The mistake was corrected in 1986. Alamodactylus^[11] Andres Myers 2013 Valid Late Cretaceous^[11] N. America^[11] Alanqa^[12] Ibrahim et al. 2010 Valid Late Cretaceous Africa Albadraco^[13] Solomon et al. 2019 Valid Late Cretaceous Europe Alcione^[14] Longrich et al. 2018 Valid Late Cretaceous Africa Allkaruen Codorniú et al. 2016 Valid Early Jurassic or Middle Jurassic S. America Altmuehlopterus^[15] Vidovic Martill 2017 Valid Late Jurassic Europe Amblydectes Hooley 1914 Nomen dubium Early Cretaceous Europe Angustinaripterus^[16] He Xinlu et al. 1983 Valid Middle Jurassic Asia Known from a single skull recovered from the Shaximiao Formation. Its Dorygnathus-like teeth suggest it had a piscivorous diet. Anhanguera^[17] Campos Kellner 1985 Valid Early Cretaceous S. America A piscivorous anhanguerid from the Santana Formation. A study of its anatomy helped resolve controversy regarding pterodacyloids' posture while on the ground. Anurognathus^[18] Döderlein 1923 Valid Late Jurassic.^[19] Europe^[19] A tiny (50 cm wingspan) insectivore known only from two skeletons recovered from the Solnhofen Formation. Apatomerus^[20] Williston 1903 Misidentification^[2] Early Cretaceous N. America Probably a misidentified plesiosaur.^[2] Apatorhamphus^[21] McPhee et al. 2020 Valid Early Cretaceous Africa Arambourgiania^[22] Nesov Kanznyshkina Cherepanov 1987 Valid Late Cretaceous Asia Aralazhdarcho^[23] Averianov 2007 Valid Late Cretaceous Asia Araripedactylus Wellnhofer 1977 Valid Early Cretaceous S. America Araripesaurus Price 1971 Valid Early Cretaceous S. America Archaeoistiodactylus Lü Fucha 2011 Valid Middle Jurassic Asia Arcticodactylus Kellner 2015 Valid Late Triassic Europe Ardeadactylus^[24] Bennett 2013 Valid Late Jurassic Europe Argentinadraco Kellner & Calvo 2017 Valid Late Cretaceous S. America Arthurdactylus Frey Martill 1994 Valid Early Cretaceous S. America Aurorazhdarcho^[25] Frey Meyer Tischlinger 2011 Valid Late Jurassic.^[25] Europe^[25] Aussiedraco^[26] Kellner Rodrigues Costa 2011 Valid Early Cretaceous Australia Austriadactylus^[16] Dalla Vecchia Wild Reitner 2002 Valid Late Triassic Europe Austriadraco Kellner 2015 Valid Late Triassic Europe Avgodectes^[27] Peters 2004 Jr. snyonym N/A N/A Probable junior synonym of Haopterus. Aymberedactylus Pêgas et al. 2016 Valid Early Cretaceous S. America Azhdarcho^[28] Nesov 1984 Valid Late Cretaceous Asia Bakonydraco^[29] Ösi Weishampel Jianu 2005 Valid Late Cretaceous Europe Banguela^[30] Headden Campos 2015 Jr. snyonym Early Cretaceous S. America Subjective junior synonym of Thalassodromeus. Barbaridactylus^[14] Longrich et al. 2018 Valid Late Cretaceous Africa Barbosania^[31] Elgin Frey 2011 Valid Late Cretaceous S. America Batrachognathus^[32] Rjabinin 1948 Valid Late Jurassic.^[33] Asia^[33] Beipiaopterus^[34] Lü 2003 Valid Early Cretaceous Asia Belonochasma^[2] Broili 1939 Misidentification^[2] N/A N/A A non-pterosaurian gnathostome.^[2] Bellubrunnus^[35] Hone et al. 2012 Valid Late Jurassic Europe Bennettazhia Nesov 1991 Valid Early Cretaceous N. America Bergamodactylus^[36] Kellner 2015 Valid Late Triassic Europe Bogolubovia^[37] Nesov A. A. Yarkov 1989 Valid Late Cretaceous Asia Boreopterus^[38] Lü Q. Ji 2005 Valid Early Cretaceous Asia Brachytrachelus Giebel 1852 Preoccupied N/A N/A preoccupied name; now known as Scaphognathus Brasileodactylus^[39] Kellner 1984 Valid Early Cretaceous S. America Cacibupteryx^[40] Gasparini Fernández de la Fuente 2004 Valid Late Jurassic Cuba Caelestiventus^[41] Britt Dalla Vecchia Chure et al. 2018 Valid Late Triassic North America Caiuajara^[42] Manzig et al. 2014 Valid Late Cretaceous S. America Camposipterus^[43] Rodrigues Kellner 2013 Valid Early Cretaceous Europe "Campylognathus"^[44] Plieninger 1894 Preoccupied N/A N/A preoccupied name; now known as Campylognathoides Campylognathoides^[16] Strand 1928 Valid Early Jurassic^[45] Europe^[45] Carniadactylus^[46] Dalla Vecchia 2009 Valid Late Triassic Europe Formerly Eudimorphodon rosenfeldi.^[46] Cascocauda^[47] Yang et al. 2022 Valid Middle Jurassic-Late Jurassic Asia Cathayopterus^[48]^[49] Wang Zhou 2006 Valid Early Cretaceous Asia Caulkicephalus^[50] Steel Martill et al. 2005 Valid Early Cretaceous Europe Caupedactylus^[51] Kellner 2013 Valid Early Cretaceous S. America Caviramus^[52] Fröbisch Fröbisch 2006 Valid Late Triassic Europe Cearadactylus^[53] Leonardi Borgomanero 1985 Valid Early Cretaceous S. America Changchengopterus^[54] Lü 2009 Valid Late Jurassic Asia Chaoyangopterus^[55] Wang Zhou 2003 Valid Early Cretaceous Asia Cimoliopterus^[43] Rodrigues Kellner 2013 Valid Late Cretaceous Europe Cimoliornis Owen 1846 Nomen nudum N/A N/A Coloborhynchus^[16] Owen 1874 Valid Early Cretaceous Europe N. America S. America Africa Comodactylus^[56] Galton 1981 Nomen dubium Late Jurassic N. America Cretornis Fritsch 1880 Valid Late Cretaceous Europe Criorhynchus Owen 1874 Jr. snyonym N/A N/A Jr. synonym of Ornithocheirus. Cryodrakon^[57] Hone et al. 2019 Valid Late Cretaceous N. America Ctenochasma^[58] von Meyer 1852 Valid Late Jurassic.^[59] Europe^[59] Cuspicephalus^[60] Martill Etches 2013 Valid Late Jurassic Europe Cycnorhamphus^[16] Seeley 1870 Valid Late Jurassic.^[61] Europe^[61] "Daitingopterus"^[62] Maisch Matzke Ge Sun 2004 Nomen nudum Late Jurassic Europe Daohugoupterus Cheng et al. 2015 Valid Late Jurassic Asia Darwinopterus^[63] Lü Unwin et al. 2009 Valid Middle Jurassic Asia Intermediate form between rhamphorhynchoids and pterodactyloids. Dawndraco^[64] Kellner 2010 Valid Late Cretaceous N. America Possible junior synonym of Geosternbergia. Dearc^[65] Jagielska et al. 2022 Valid Middle Jurassic Isle of Skye Dendrorhynchoides^[66] S.-A. Ji Q. Ji Padian 1999 Valid Late Jurassic or Early Cretaceous Asia "Dendrorhynchus"^[67] S.-A. Ji Q. Ji 1998 Preoccupied N/A N/A Preoccupied name; now known as Dendrorhynchoides. Dermodactylus Marsh 1881 Nomen dubium Late Jurassic N. America Dimorphodon^[16] Owen 1859 Valid Early Jurassic^[68] Europe^[68] Diopecephalus Seeley 1871 Valid Late Jurassic Europe Dolicorhamphus^[69] Seeley 1875 Nomen dubium Middle Jurassic Europe Domeykodactylus^[70] Martill Frey et al. 2000 Valid Early Cretaceous S. America Doratorhynchus^[71] Seeley 1875 Nomen vanum Late Jurassic or Early Cretaceous Europe Dorygnathus^[16] Wagner 1860 Valid Early Jurassic^[72] Europe^[72] Douzhanopterus^[73] Wang et al. 2017 Valid Late Jurassic Asia Draigwenia^[74] Holgado 2021 Valid Early Cretaceous Europe Dsungaripterus^[16] Young 1964 Valid Late Jurassic-Early Cretaceous^[75] Asia^[75] Elanodactylus^[76] Andres Ji, Q. 2008 Valid Early Cretaceous Asia Eoazhdarcho^[77] Lü Ji 2005 Valid Early Cretaceous Asia Eopteranodon^[78] Lü Zhang 2005 Valid Early Cretaceous Asia Eosipterus^[16] Ji Ji 1997 Valid Early Cretaceous Asia Epapatelo^[79] Fernandes et al. 2022 Valid Late Cretaceous Africa Eudimorphodon^[16] Zambelli 1973 Valid Late Triassic^[80] Europe^[80] Europejara^[81] Vullo et al. 2012 Valid Early Cretaceous Europe Eurazhdarcho^[82] Vremir et al. 2013 Valid Late Cretaceous Europe Eurolimnornis^[83] Kessler Jurcsák 1986 Valid Early Cretaceous Europe Originally described as a bird, subsequently reinterpreted as a pterosaur.^[84] Faxinalipterus^[85] Bonaparte, et al. 2010 Misidentification N/A N/A Originally described as a pterosaur, but has since been reinterpreted as a non-pterosaur pterosauromorph.^[86] Feilongus^[87] Wang, Kellner, et al. 2005 Valid Early Cretaceous Asia Fenghuangopterus^[88] Lü Fucha Chen 2010 Valid Middle Jurassic Asia Ferrodraco^[89] Pentland et al. 2019 Valid Late Cretaceous Australia Forfexopterus Jiang et al. 2016 Valid Early Cretaceous Asia Gallodactylus Fabre 1974 Jr. snyonym N/A N/A Jr. synonym of Cycnorhamphus. Geosternbergia Miller 1978 Valid N/A N/A Probable Jr. synonym of Pteranodon. Gegepterus^[90] X. Wang Kellner et al. 2007 Valid Early Cretaceous Asia Germanodactylus^[16] Yang 1964 Valid Late Jurassic.^[91] Europe^[91] Gladocephaloideus^[92] Lü Ji Wei Liu 2011 Valid Early Cretaceous Asia Gnathosaurus^[16] von Meyer 1833 Valid Late Jurassic^[93] Europe^[93] Guidraco^[94] Wang et al. 2012 Valid Early Cretaceous Asia Gwawinapterus^[95] Arbour Currie 2011 Misidentification Late Cretaceous^[95] N. America^[95] Initially thought to be an istiodactylid pterosaur,^[95] but subsequently reinterpreted as an indeterminate saurodontid fish.^[96] Hamipterus^[97] Wang et al. 2014 Valid Early Cretaceous Asia Haopterus^[98] Wang X. Lü 2001 Valid Early Cretaceous Asia Harpactognathus^[99] Carpenter Unwin et al. 2003 Valid Late Jurassic N. America Hatzegopteryx^[100] Buffetaut Grigorescu Csiki 2002 Valid Late Cretaceous Europe Herbstosaurus^[16] Casamiquela 1974 Valid Late Jurassic S. America Hongshanopterus^[101] X. Wang Campos et al. 2008 Valid Early Cretaceous Asia Huanhepterus^[102] Dong 1982 Valid Late Jurassic Asia Huaxiapterus^[103] Lü C. Yuan 2005 Valid Early Cretaceous Asia Iberodactylus^[104] Holgado et al. 2019 Valid Early Cretaceous Europe Ikrandraco^[105] Wang et al. 2014 Valid Early Cretaceous Asia Ingridia^[106] Unwin Martill 2007 Jr. snyonym N/A N/A Objective Jr. synonym of Tupandactylus. Istiodactylus^[16] Howse Milner Martill 2001 Valid Early Cretaceous Europe Javelinadactylus Campos 2021 Jr. synonym N/A N/A Junior synonym of Wellnhopterus. Although named a few months earlier, its description was retracted over allegations that the describer did not have access to its holotype, and seemingly wrote it to beat out a major series of papers which the Wellnhopterus description was part of. Jeholopterus^[107] X. Wang Zhou 2002 Valid Middle Jurassic or Early Cretaceous Asia Jianchangnathus^[108] Cheng Wang Jiang Kellner 2012 Valid Middle Jurassic Asia Jianchangopterus^[109] Lü Bo 2011 Valid Middle Jurassic Asia Jidapterus^[110] Dong Sun Wu 2003 Valid Early Cretaceous Asia Kariridraco^[111] Cerqueira et al. 2021 Valid Early Cretaceous S. America Kepodactylus^[112] Harris Carpenter 1996 Valid Late Jurassic N. America Keresdrakon^[113] Kellner et al. 2019 Valid Early Cretaceous to Late Cretaceous S. America Klobiodon^[114] O'Sullivan Martill 2018 Valid Middle Jurassic Europe Kunpengopterus^[115] X. Wang Kellner et al. 2010 Valid Middle Jurassic Asia Kryptodrakon^[116] Andres Clark Xu 2014 Valid Middle Jurassic and/or Late Jurassic Asia Lacusovagus^[117] Witton 2008 Valid Early Cretaceous S. America Laopteryx Marsh 1881 Nomen dubium Late Jurassic N. America Leptostomia^[118] Smith et al. 2020 Valid Early or Late Cretaceous Africa Liaodactylus^[119] C.-F. Zhou et al. 2017 Valid Late Jurassic Asia Liaoningopterus^[120] X.-L. Wang Z.-H. Zhou 2003 Valid Early Cretaceous Asia Liaoxipterus^[121] Dong Lü 2005 Valid Early Cretaceous Asia Limnornis^[122] Kessler Jurcsák 1984 Preoccupied Early Cretaceous Europe Originally described as a bird; fossils later renamed Palaeolimnornis. Linlongopterus^[123] Rodrigues et al. 2015 Valid Early Cretaceous Asia Lingyuanopterus^[124] Xu et al. 2022 Valid Early Cretaceous Asia "Lithosteornis"^[125] Gervais 1844 Nomen nudum N/A N/A Nomen nudum. Lonchodectes^[16] Hooley 1914 Valid Early Cretaceous to Late Cretaceous Europe Lonchodraco^[43] Rodrigues Kellner 2013 Valid Early Cretaceous to Late Cretaceous Europe Lonchognathosaurus^[126] Maisch Matzke Ge Sun 2004 Valid Early Cretaceous Asia Longchengpterus^[127] L. Wang L. Li et al. 2006 Valid Early Cretaceous Asia Luchibang^[128] Hone L. Li et al. 2020 Valid Early Cretaceous Asia Ludodactylus^[129] Frey Martill Buchy 2003 Valid Early Cretaceous S. America Luopterus^[130] Hone 2020 Valid Late Jurassic Asia Maaradactylus^[131] Bantim et al. 2014 Valid Early Cretaceous S. America Macrotrachelus Giebel 1852 Jr. snyonym N/A N/A Jr. synonym of Pterodactylus. Mesadactylus^[132] Jensen Padian 1989 Valid Late Jurassic N. America Microtuban^[133] Elgin Frey 2011 Valid Late Cretaceous Africa Mimodactylus^[134] Kellner et al. 2019 Valid Late Cretaceous Lebanon Mistralazhdarcho^[135] Vullo Garcia Godefroit et al. 2018 Valid Late Cretaceous Europe Moganopterus^[136] Lü et al. 2012 Valid Early Cretaceous Asia Montanazhdarcho^[137] Padian Horner de Ricqlès 1993 Valid Late Cretaceous N. America Muzquizopteryx^[138] Frey Buchy et al. 2006 Valid Late Cretaceous N. America Mythunga^[139] Molnar Thulborn 2008 Valid Early Cretaceous Australia Navajodactylus^[140] Sullivan Fowler 2011 Valid Late Cretaceous N. America Nemicolopterus^[141] X. Wang Kellner et al. 2008 Valid Early Cretaceous Asia Nesodactylus^[142] Colbert 1969 Valid Late Jurassic Cuba Nesodon^[143] Jensen Ostrom 1977 Lapsus calami N/A N/A Misspelling of Nesodactylus, also preoccupied by a toxodont. Nicorhynchus^[6] Holgado Pêgas 2020 Valid Early Cretaceous to Late Cretaceous Africa Europe Ningchengopterus^[144] Lü 2009 Valid Early Cretaceous Asia Noripterus^[16] Yang 1973 Valid Early Cretaceous Asia Normannognathus^[126] Buffetaut J.-J. Lepage G. Lepage 1998 Valid Late Jurassic Europe Nurhachius Wang Kellner et al. 2003 Valid Early Cretaceous Asia Nyctodactylus^[145] Marsh 1881 Jr. snyonym N/A N/A Jr. synonym of Nyctosaurus. Nyctosaurus^[146] Marsh 1876 Valid Late Cretaceous^[147] N. America,^[147] S. America^[147] "Odontorhynchus" Stolley 1936 Jr. snyonym N/A N/A Jr. synonym of Rhamphorhynchus. "Oolithorhynchus"^[2] Whalley 2000 Nomen nudum^[2] Manuscript name^[2] about which almost nothing is known. Orientognathus Lü et al. 2015 Valid Late Jurassic Asia Ordosipterus^[148] Ji 2020 Valid Early Cretaceous Asia Ornithocephalus^[149] von Sömmering 1812 Jr. snyonym N/A N/A Jr. synonym of Pterodactylus. Ornithocheirus^[16] Seeley 1869 Valid Early Cretaceous.^[43] Europe^[43] Ornithodesmus^[2] Seeley 1887 Misidentification^[2] N/A N/A Misidentified dromaeosaurid.^[2] Ornithopterus^[150] Fitzinger 1843 Jr. snyonym N/A N/A Jr. synonym of Rhamphorhynchus. Ornithostoma^[16] Seeley 1871 Valid Late Cretaceous Europe "Osteornis"^[125] Gervais 1844 Nomen nudum N/A Europe Nomen ex dissertationae for pterosaur remains later named Palaeornis and Cimoliornis. Otogopterus^[151] Ji Zhang 2020 Valid Early Cretaceous Asia Pachagnathus^[152] Martínez et al. 2022 Valid Late Triassic S. America Pachyrhamphus^[150] Fitzinger 1843 Preoccupied N/A N/A Preoccupied name; now known as Scaphognathus Palaeocursornis^[83] Kessler Jurcsák 1986 Valid Early Cretaceous Europe Originally described under the name Limnornis as a bird; Limnornis was preoccupied, and the fossils were subsequently reinterpreted as pterosaurian.^[84] Palaeornis Mantell 1844 Preoccupied N/A N/A preoccupied name. Pangupterus Lü et al. 2016 Valid Early Cretaceous Asia Paranurognathus^[153] Peters 2005 Jr. snyonym N/A N/A Jr. synonym of Anurognathus. Parapsicephalus^[154] von Arthaber 1919 Valid Early Jurassic Europe Peteinosaurus^[16] Wild 1978 Valid Late Triassic^[155] Europe^[155] "Phobetor"^[2]^[16] Bakhurina 1986 Jr. snyonym N/A N/A Jr. synonym of Noripterus.^[156] Phosphatodraco^[157] Pereda-Suberbiola Bardet et al. 2003 Valid Late Cretaceous Africa Piksi^[158] Varricchio 2002 Valid Late Cretaceous N. America Originally described as a bird, subsequently reinterpreted as a pterosaur.^[84] Plataleorhynchus^[159] Howse Milner 1995 Valid Late Jurassic to Early Cretaceous Europe Prejanopterus^[160] Vidarte Calvo 2010 Valid Early Cretaceous Europe Preondactylus^[16] Wild 1983 Valid Late Triassic^[161] Europe^[161] "Pricesaurus"^[162] Bonaparte Sanchez 1986 Nomen nudum Early Cretaceous S. America Procoelosaurus^[163] Atanassov 2002 Nomen ex dissertatione^[2] Ptenodactylus^[164] Seeley 1869 Preoccupied N/A N/A Preoccupied name. Ptenodracon^[165] Lydekker 1888 Jr. synonym N/A N/A Junior synonym of Ctenochasma. Pteranodon^[16] Marsh 1876 Valid Late Cretaceous^[166] N. America.^[166] Ptéro-dactyle Cuvier 1809 N/A N/A Renamed Pterodactylus. Pterodactylus Rafinesque 1815 Valid Late Jurassic.^[167] Africa,^[167] Europe^[167] Pterodaustro^[16] Bonaparte 1970 Valid Early Cretaceous^[168] S. America^[168] Pterofiltrus^[169] Jiang Wang 2011 Valid Early Cretaceous Asia Pteromimus^[163] Atanassov 2002 Nomen ex dissertatione^[2] Pteromonodactylus Teryaev 1967 Jr. snyonym N/A N/A Jr. synonym of Rhamphorhynchus. Pterorhynchus^[170] Czerkas Q. Ji 2002 Valid Late Jurassic Asia Pterotherium^[171] Fischer von Waldheim 1813 Jr. snyonym N/A N/A Jr. synonym of Pterodactylus. Puntanipterus^[172] Bonaparte Sanchez 1975 Valid Late Jurassic to Early Cretaceous S. America Qinglongopterus^[173] Lü Unwin et al. 2012 Valid Middle Jurassic Asia Quetzalcoatlus^[16] Lawson 1975 Valid Late Cretaceous^[174] N. America^[174] Radiodactylus^[11] Andres Myers 2013 Valid Early Cretaceous N. America Raeticodactylus^[175] Stecher 2008 Valid N/A N/A Possible junior synonym of Caviramus Rhabdopelix^[2] Cope 1870 Misidentification^[2] N/A N/A At first it was thought to be a Triassic pterosaur, but is now known to be (at least in part) a kuehneosaurid.^[2] Rhamphinion^[176] Padian 1984 Valid Early Jurassic N. America Rhamphocephalus Seeley 1880 Misidentification Middle Jurassic Europe Originally assigned to Pterosauria, later reassigned to Thalattosuchia.^[177] Rhamphorhynchus^[16] von Meyer 1846 Valid Late Jurassic.^[178] Africa,^[178] Europe^[178] Samrukia^[179] Naish et al. 2012 Valid Late Cretaceous Asia Originally described as a bird but reinterpreted as a pterosaur. Santanadactylus^[180] de Buisonjé 1980 Valid Early Cretaceous S. America Scaphognathus^[16] Wagner 1861 Valid Late Jurassic.^[181] Europe^[181] Seazzadactylus^[182] Dalla Vecchia 2019 Valid Late Triassic Europe Sericipterus Andres Clark X. Xing 2010 Valid Late Jurassic Asia Serradraco^[183] Rigal Martill Sweetman 2017 Valid Late Jurassic. Europe Shenzhoupterus^[184] Lü Unwin et al. 2008 Valid Early Cretaceous Asia Simurghia^[14] Longrich et al. 2018 Valid Late Cretaceous Africa Sinomacrops^[185] Wei et al. 2021 Valid Middle Jurassic to Late Jurassic Asia Sinopterus^[186] X. Wang Z. Zhou 2003 Valid Early Cretaceous Asia Siroccopteryx^[187] Mader Kellner 1999 Valid N/A N/A Possible junior synonym of Coloborhynchus Sordes^[16] Sharov 1971 Valid Late Jurassic.^[188] Asia^[188] Sultanuvaisia^[10] Nesov 1981 Misidentification Late Cretaceous Asia. Actually a fish. Tacuadactylus^[189] Soto et al. 2021 Valid Late Jurassic S. America Tapejara^[16] Kellner 1989 Valid Early Cretaceous S. America Targaryendraco^[190] Pêgas et al. 2019 Valid Early Cretaceous Europe Tendaguripterus^[191] Unwin Heinrich 1999 Valid Late Jurassic Africa Tethydraco^[14] Longrich et al. 2018 Valid Late Cretaceous Africa Thalassodromeus^[192] Kellner Campos 2002 Valid Early Cretaceous S. America Thanatosdrakon Ortiz David et al. 2022 Valid Late Cretaceous South America Thapunngaka^[193] Richards et al. 2021 Valid Early Cretaceous Australia "Titanopteryx"^[194] Arambourg 1959 Preoccupied Late Cretaceous Asia Preoccupied by a simuliid blackfly. It was later renamed Arambourgiania. Tribelesodon Bassani 1886 Misidentification N/A N/A At first it was thought to be a Triassic pterosaur but is now known to be a misinterpreted specimen of the prolacertiform Tanystropheus. Tropeognathus Wellnhofer 1987 Valid Early Cretaceous S. America One of the largest anhanguerids. Tupandactylus^[195] Kellner Campos 2007 Valid Early Cretaceous S. America Tupuxuara^[196] Kellner Campos 1988 Valid Late Cretaceous S. America Uktenadactylus^[197] Rodrigues Kellner 2008 Valid Early Cretaceous N. America Possible junior synonym of Coloborhynchus Unwindia^[198] Martill 2011 Valid Late Cretaceous S. America Utahdactylus^[199] Czerkas Mickelson 2002 Valid Late Jurassic N. America Vectidraco^[200] Naish Simpson Dyke 2013 Valid Early Cretaceous Europe Vesperopterylus^[201] Lü et al. 2017 Valid Early Cretaceous Asia Volgadraco^[202] Averianov Arkhangelsky Pervushov 2008 Valid Late Cretaceous Asia Wellnhopterus^[203] Andres, Langston Jr. 2021 Valid Late Cretaceous N. America Wenupteryx^[204] Codorniú Gasparini 2013 Valid Late Jurassic S. America Wightia^[205] Martill et al. 2020 Valid Early Cretaceous Europe Wukongopterus X. Wang Kellner et al. 2009 Valid Middle Jurassic Asia "Wyomingopteryx"^[206] Bakker 1994 Nomen nudum Xericeps^[207] D. Martill et al. 2017 Valid Early Cretaceous to Late Cretaceous Africa Yelaphomte^[152] Martínez et al. 2022 Valid Late Triassic S. America Yixianopterus^[208] Lü S. Ji et al. 2006 Valid Early Cretaceous Asia Zhejiangopterus^[209] Z. Cai F. Wei 1994 Valid Late Cretaceous Asia Zhenyuanopterus^[210] Lü 2010 Valid Early Cretaceous Asia		Angustinaripterus   Anhanguera   Anurognathus   Aurorazhdarcho   Bakonydraco   Campylognathoides   Cearadactylus   Coloborhynchus   Ctenochasma   Dimorphodon   Dorygnathus   Dsungaripterus   Eudimorphodon   Feilongus   Germanodactylus   Haopterus   Hatzegopteryx   Istiodactylus   Jeholopterus   Liaoningopterus   Lonchodectes   Nemicolopterus   Ningchengopterus   Noripterus   Nurhachius   Nyctosaurus   Ornithocheirus   Preondactylus   Pteranodon   Pterodactylus   Pterodaustro   Quetzalcoatlus   Rhamphorhynchus   Scaphognathus   Sordes   Tapejara   Thalassodromeus   Tupandactylus   Tupuxuara   Zhejiangopterus

Ichnogenera

Genus Authors Year Status Age Location Notes Agadirichnus Ambroggi Lapparent 1954 Valid uK Africa Haenamichnus^[211] Hwang Huh et al. 2002 Valid uK^[211] Asia.^[211] More than 5 times as large as Pteraichnus, these tracks were probably made by azhdarchids.^[211] Kouphichnium Nopcsa 1923 Misidentification N/A N/A The trackmaker was probably a limulid. Pteraichnus Stokes 1957 Valid uJ Asia^[212] Europe N. America Some tracks attributed to Pteraichnus have been considered crocodilian in origin by some scientists (Padian, 1984) however, recent studies in the past decade have overturned Padian's thoughts. Purbeckopus Delair 1963 Nomen dubium lK Europe.		Haenamichnus

Oogenera

Although pterosaur eggs are known, some with complete embryos, no oogenera have been erected to house them. The holotype of the oospecies Oolithes sphaericus was briefly considered by Harry Govier Seeley to be pterosaurian in origin, although this attribution was dismissed before the formal erection of that oogenus.

See also

•  Paleontology portal

• List of dinosaurs
• List of plesiosaurs
• List of pterosaur classifications
• Pterosaur
• Timeline of pterosaur research

References

1. Haaramo, Mikko (15 November 2005). . Mikko's Phylogeny Archive. Archived from the original on 29 March 2007. Retrieved 29 March 2007.
2. Hanson, Mike (12 October 2005). . The Pterosauria. Archosauria.org. Archived from the original on 1 July 2007. Retrieved 29 March 2007.
3. Hanson, Mike (30 September 2006). (PDF). The Pterosauria. Archosauria.org. Archived from the original (PDF) on 29 July 2007. Retrieved 27 July 2007.
4. Glut, Donald F. (2006). "Appendix One: Pterosaurs". Dinosaurs: The Encyclopedia. 4th Supplement. Jefferson, North Carolina: McFarland & Company, Inc. pp. 583–633. ISBN 978-0-7864-2295-1.
5. Vidovic, S. U.; Martill, D. M. (2014). "Pterodactylus scolopaciceps Meyer, 1860 (Pterosauria, Pterodactyloidea) from the Upper Jurassic of Bavaria, Germany: The Problem of Cryptic Pterosaur Taxa in Early Ontogeny". PLOS ONE. 9 (10): e110646. Bibcode:2014PLoSO...9k0646V. doi:10.1371/journal.pone.0110646. PMC 4206445. PMID 25337830.
6. Borja Holgado; Rodrigo V. Pêgas (2020). "A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae". Acta Palaeontologica Polonica. 65. doi:10.4202/app.00751.2020.
7. Fernando E. Novas; Martin Kundrat; Federico L. Agnolín; Martín D. Ezcurra; Per Erik Ahlberg; Marcelo P. Isasi; Alberto Arriagada; Pablo Chafrat (2012). "A new large pterosaur from the Late Cretaceous of Patagonia". Journal of Vertebrate Paleontology. 32 (6): 1447–1452. doi:10.1080/02724634.2012.703979. S2CID 84340520.
8. Myers, Timothy S. (2010). "A new ornithocheirid pterosaur from the Upper Cretaceous (Cenomanian–Turonian) Eagle Ford Group of Texas". Journal of Vertebrate Paleontology. 30 (1): 280–287. doi:10.1080/02724630903413099. S2CID 130367018.
9. David M. Martill; Roy Smith; David M. Unwin; Alexander Kao; James McPhee; Nizar Ibrahim (2020). "A new tapejarid (Pterosauria, Azhdarchoidea) from the mid-Cretaceous Kem Kem beds of Takmout, southern Morocco". Cretaceous Research. 112: Article 104424. doi:10.1016/j.cretres.2020.104424. S2CID 216303122.
10. Nesov, Lev A. (1981). "[Flying reptiles from the Late Cretaceous of Kyzyl-Kum]". Paleontologicheskii Zhurnal (in Russian). 15: 98–104.
11. Andres, Brian; Myers, Timothy S. (2013). "Lone Star Pterosaurs". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 383–398. doi:10.1017/S1755691013000303. S2CID 84617119.
12. Ibrahim, N.; Unwin, D.M.; Martill, D.M.; Baidder, L.; Zouhri, S. (2010). "A New Pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco". PLOS ONE. 5 (5): e10875. Bibcode:2010PLoSO...510875I. doi:10.1371/journal.pone.0010875. PMC 2877115. PMID 20520782.
13. Alexandru A. Solomon; Vlad A. Codrea; Márton Venczel; Gerald Grellet-Tinner (2019). "A new species of large-sized pterosaur from the Maastrichtian of Transylvania (Romania)". Cretaceous Research. 110: Article 104316. doi:10.1016/j.cretres.2019.104316. S2CID 213808137.
14. Nicholas R. Longrich; David M. Martill; Brian Andres (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3): e2001663. doi:10.1371/journal.pbio.2001663. PMC 5849296. PMID 29534059.
15. Steven U. Vidovic; David M. Martill (2017). "The taxonomy and phylogeny of Diopecephalus kochi (Wagner, 1837) and Germanodactylus rhamphastinus (Wagner, 1851)" (PDF). In D. W. E. Hone; M. P. Witton; D. M. Martill (eds.). New Perspectives on Pterosaur Palaeobiology (PDF). Geological Society of London, Special Publications. Vol. 455. The Geological Society of London. pp. 125–147. Bibcode:2018GSLSP.455..125V. doi:10.1144/SP455.12. S2CID 219204038.
16. Unwin, David M. Appendix: List of Valid Pterosaur Species. Pterosaurs From Deep Time. New York: Pi Press, 2006. ISBN 0-13-146308-X
17. Campos; Kellner (1985). "Panorama of the Flying Reptiles Study in Brazil and South America (Pterosauria/ Pterodactyloidea/ Anhangueridae)". Anais da Academia Brasileira de Ciências. 57 (4): 141–142 & 453–466.
18. Döderlein, L. (1923). "Anurognathus ammoni ein neuer Flugsaurier." Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Abteilung der Bayerischen Akademie der Wissenschaften zu München, 1923, 117–164.
19. "Anurognathus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 306–307.
20. Williston, Samuel W. (1903). "On the osteology of Nyctosaurus (Nyctodactylus), with notes on American pterosaurs". Fieldiana Geology. 2 (3): 125–163.
21. James McPhee; Nizar Ibrahim; Alex Kao; David M. Unwin; Roy Smith; David M. Martill (2020). "A new ?chaoyangopterid (Pterosauria: Pterodactyloidea) from the Cretaceous Kem Kem beds of Southern Morocco". Cretaceous Research. 110: Article 104410. doi:10.1016/j.cretres.2020.104410. S2CID 213739173.
22. Nesov, L.A., Kanznyshkina, L.F., and Cherepanov, G.O. (1987). "Dinosaurs, crocodiles and other archosaurs from the Late mesozoic of central Asia and their place in ecosystems." Abstracts of the 33rd session of the All-Union Palaeontological Society, Leningrad, pp. 46–47. [In Russian].
23. Averianov, A.O. (2007). "New records of azhdarchids (Pterosauria, Azhdarchidae) from the late Cretaceous of Russia, Kazakhstan, and Central Asia". Paleontological Journal. 41 (2): 189–197. doi:10.1134/S0031030107020098. S2CID 128637719.
24. Bennett, S. Christopher (2013). "New information on body size and cranial display structures of Pterodactylus antiquus, with a revision of the genus". Paläontologische Zeitschrift. 87 (2): 269–289. doi:10.1007/s12542-012-0159-8. S2CID 83722829.
25. Eberhard Frey, Christian A. Meyer and Helmut Tischlinger (2011). "The oldest azhdarchoid pterosaur from the Late Jurassic Solnhofen Limestone (Early Tithonian) of Southern Germany". Swiss Journal of Geosciences. 104: 35–55. doi:10.1007/s00015-011-0073-1. S2CID 129784137.
26. Kellner, Alexander W.A.; Taissa Rodrigues; Fabiana R. Costa (2011). "Short note on a pteranodontoid pterosaur (Pterodactyloidea) from western Queensland, Australia" (PDF). Anais da Academia Brasileira de Ciências. 83 (1): 301–308. doi:10.1590/S0001-37652011000100018. PMID 21437387.
27. Peters, David (2004). . Prehistoric Times. 67: 21. Archived from the original on 2005-02-11.
28. Nesov (1984). . Paleontologicheskii Zhurnal. 1984 (1): 47–57. Archived from the original on 2009-01-05.
29. Ösi, Attila; Weishampel, David B.; Jianu, Coralia M. (2005). "First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary" (PDF). Acta Palaeontologica Polonica. 50 (4): 777–787. Retrieved 2016-11-30.
30. Jaime A. Headden & Hebert B.N. Campos (2015). "An unusual edentulous pterosaur from the Early Cretaceous Romualdo Formation of Brazil". Historical Biology: An International Journal of Paleobiology. 27 (7): 815–826 1–12. doi:10.1080/08912963.2014.904302. S2CID 129306469.
31. Elgin, Ross A.; Frey, Eberhard (2011). "A new ornithocheirid, Barbosania gracilirostris gen. et sp. nov. (Pterosauria, Pterodactyloidea) from the Santana Formation (Cretaceous) of NE Brazil". Swiss Journal of Palaeontology. 130 (2): 259–275. doi:10.1007/s13358-011-0017-4. S2CID 89178816.
32. Rjabinin, A. N. (1948). "Remarks on a flying reptile from the Jurassic of the Kara-Tau". Akademia Nauk. 15 (1): 86–93.
33. "Batrachognathus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 320–321.
34. Lü, J.-C. (2003). "A new pterosaur: Beipiaopterus chenianus, gen. et sp. nov. (Reptilia: Pterosauria) from Western Liaoning Province, China". Memoir of the Fukui Prefectural Dinosaur Museum. 2: 153–160.
35. David W. E. Hone; Helmut Tischlinger; Eberhard Frey; Martin Röper (2012). "A New Non-Pterodactyloid Pterosaur from the Late Jurassic of Southern Germany". PLOS ONE. 7 (7): e39312. Bibcode:2012PLoSO...739312H. doi:10.1371/journal.pone.0039312. PMC 3390345. PMID 22792168.
36. Vecchia, Fabio M. Dalla (July 2018). "COMMENTS ON TRIASSIC PTEROSAURS WITH A COMMENTARY ON THE "ONTOGENETIC STAGES" OF KELLNER (2015) AND THE VALIDITY OF BERGAMODACTYLUS WILDI". Rivista Italiana di Paleontologia e Stratigrafia (Research in Paleontology and Stratigraphy). 124 (2): 317–341. doi:10.13130/2039-4942/10099. ISSN 2039-4942.
37. Nesov; Yarkov, A.A. (1989). "New Birds from the Cretaceous–Paleogene of the USSR and Some Remarks on the History of Origin and Evolution of the Class". Tr. Zool. Inst. Akad. Nauk SSSR. 197: 78–97.
38. Lü, Junchang; Ji, Qiang (2005). "A new ornithocheirid from the Early Cretaceous of Liaoning Province, China". Acta Geologica Sinica. 79 (2): 157–163. doi:10.1111/j.1755-6724.2005.tb00877.x. S2CID 129084078.
39. Kellner, A. W. A. (1984). "Ocorrencia de uma mandibula de pterosauria (Brasileodactylus araripensis, nov. gen.; nov. sp.) na Formacao Santana, Cretaceo da Chapada do Araripe, Ceara-Brasil." Anais XXXIII Cong. Brasil. de Geol, 578–590. Rio de Janeiro.
40. Gasparini, Zulma; Fernández, Marta; de la Fuente, Marcelo (2004). "A new pterosaur from the Jurassic of Cuba". Palaeontology. 47 (4): 919–927. doi:10.1111/j.0031-0239.2004.00399.x.
41. Brooks B. Britt; Fabio M. Dalla Vecchia; Daniel J. Chure; George F. Engelmann; Michael F. Whiting; Rodney D. Scheetz (2018). "Caelestiventus hanseni gen. et sp. nov. extends the desert-dwelling pterosaur record back 65 million years". Nature Ecology & Evolution. 2 (9): 1386–1392. doi:10.1038/s41559-018-0627-y. PMID 30104753. S2CID 51984440.
42. Paulo C. Manzig; Alexander W. A. Kellner; Luiz C. Weinschütz; Carlos E. Fragoso; Cristina S. Vega; Gilson B. Guimarães; Luiz C. Godoy; Antonio Liccardo; João H. Z. Ricetti; Camila C. de Moura (2014). "Discovery of a Rare Pterosaur Bone Bed in a Cretaceous Desert with Insights on Ontogeny and Behavior of Flying Reptiles". PLOS ONE. 9 (8): e100005. Bibcode:2014PLoSO...9j0005M. doi:10.1371/journal.pone.0100005. PMC 4131874. PMID 25118592.
43. Rodrigues, Taissa; Kellner, Alexander Wilhelm Armin (2013). "Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England". ZooKeys (308): 1–112. doi:10.3897/zookeys.308.5559. PMC 3689139. PMID 23794925.
44. Plieninger, F. 1895 "Campylognathus Zitteli, ein neuer Flugsaurier aus dem obersten Lias Schwabens", Paläontographica 41, 193–222 & pl. 19.
45. "Campylognathoides." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 296–297.
46. Dalla Vecchia, Fabio M. (2009). "Anatomy and systematics of the pterosaur Carniadactylus (gen. n.) rosenfeldi (Dalla Vecchia, 1995)". Rivista Italiana de Paleontologia e Stratigrafia. 115 (2): 159–188.
47. Yang, Zixiao; Benton, Michael J.; Hone, David W. E.; Xu, Xing; McNamara, Maria E.; Jiang, Baoyu (2022-03-03). "Allometric analysis sheds light on the systematics and ontogeny of anurognathid pterosaurs". Journal of Vertebrate Paleontology. 41 (5): e2028796. doi:10.1080/02724634.2021.2028796. ISSN 0272-4634. S2CID 247262846.
48. Xiaolin, Wang; Zhonghe, Zhou (2006). "Pterosaur assemblages of the Jehol Biota and their implication for the Early Cretaceous pterosaur radiation". Geological Journal. 41 (3–4): 405–418. doi:10.1002/gj.1046. S2CID 131602205.
49. Xiaolin, Wang; Zhou, Zhonghe (2006). Rong, Jiayu; Fang, Zongjie; Zhou, Zhonghe; Zhan, Renbin; Yuan, Xunalai (eds.). Originations and Radiations—Evidences from the Chinese Fossil Record. Beijing: Science Press. pp. 665–689, 937–938.
50. Steel, L.; Martill; Unwin; Winch, J. D. (2005). "A new pterodactyloid pterosaur from the Wessex Formation (Lower Cretaceous) of the Isle of Wight, England". Cretaceous Research. 26 (4): 686–698. doi:10.1016/j.cretres.2005.03.005.
51. Alexander W. A. Kellner (2013). "A new unusual tapejarid (Pterosauria, Pterodactyloidea) from the Early Cretaceous Romualdo Formation, Araripe Basin, Brazil". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 409–421. doi:10.1017/S1755691013000327. S2CID 131574480.{{cite journal}}: CS1 maint: uses authors parameter (link)
52. Fröbisch, N.B.; Fröbisch, J. (2006). "A new basal pterosaur genus from the Upper Triassic of the Northern Calcareous Alps of Switzerland". Palaeontology. 49 (5): 1081–1090. doi:10.1111/j.1475-4983.2006.00581.x.
53. Leonardi, G.; Borgomanero, G. (1985). "Cearadactylus atrox nov. gen., nov. sp.: novo Pterosauria (Pterodactyloidea) da Chapada do Araripe, Ceara, Brasil". Resumos dos communicaçoes VIII Congresso bras. De Paleontologia e Stratigrafia. 27: 75–80.
54. Lü, J. (2009). "A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China". Acta Geologica Sinica. 83 (2): 189–199. doi:10.1111/j.1755-6724.2009.00062.x. S2CID 247739978.
55. Xiao-Lin, Wang; Zhong-He, Zhou (2003). "Two new pterodactyloid pterosaurs from the Early Cretaceous Jiufotang Formation of Western Liaoning, China". Vertebrata PalAsiatica. 41 (1): 34–41.
56. Galton, P.M. (1981). "A rhamphorhynchoid pterosaur from the Upper Jurassic of North America". Journal of Paleontology. 55: 1117–1122.
57. David W. E. Hone; Michael B. Habib; François Therrien (2019). "Cryodrakon boreas gen. et sp. nov. a Late Cretaceous Canadian azhdarchid pterosaur". Journal of Vertebrate Paleontology. 39 (3): e1649681. doi:10.1080/02724634.2019.1649681. S2CID 203406859.
58. von Meyer, C. E. H. 1852, Ctenochasma Roemeri. Paläontographica 2, 82–84 & pl. 13.
59. "Ctenochasma." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 316–317.
60. Martill, David M.; Etches, Steve (2013). "A new monofenestratan pterosaur from the Kimmeridge Clay Formation (Upper Jurassic, Kimmeridgian) of Dorset, England" (PDF). Acta Palaeontologica Polonica. 58 (2): 285–294. doi:10.4202/app.2011.0071. S2CID 55336060.
61. "Gallodactylus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 312–313.
62. Maisch, M.W.; Matzke, A.T.; Ge Sun (2004). "A new dsungaripteroid pterosaur from the Lower Cretaceous of the southern Junggar Basin, north-west China". Cretaceous Research. 25 (5): 625–634. doi:10.1016/j.cretres.2004.06.002.
63. Lü, J.; Unwin, D. M.; Jin, X.; Liu, Y.; Ji, Q. (2009). "Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull". Proceedings of the Royal Society B. 277 (1680): 383–9. doi:10.1098/rspb.2009.1603. PMC 2842655. PMID 19828548.
64. Kellner, A.W.A. (2010). "Comments on the Pteranodontidae (Pterosauria, Pterodactyloidea) with the description of two new species". Anais da Academia Brasileira de Ciências. 82 (4): 1063–1084. doi:10.1590/s0001-37652010000400025. PMID 21152777.
65. Jagielska, Natalia; O’Sullivan, Michael; Funston, Gregory F.; Butler, Ian B.; Challands, Thomas J.; Clark, Neil D. L.; Fraser, Nicholas C.; Penny, Amelia; Ross, Dugald A.; Wilkinson, Mark; Brusatte, Stephen L. (2022-02-22). "A skeleton from the Middle Jurassic of Scotland illuminates an earlier origin of large pterosaurs". Current Biology. 32 (6): 1446–1453.e4. doi:10.1016/j.cub.2022.01.073. ISSN 0960-9822. PMID 35196508. S2CID 247013664.
66. Ji, S.-A.; Ji, Q.; Padian (1999). "Biostratigraphy of new pterosaurs from China". Nature. 398 (6728): 573–574. Bibcode:1999Natur.398..573J. doi:10.1038/19221. S2CID 4316586.
67. Ji, S.-A.; Ji, Q. (1998). "A new fossil pterosaur (Rhamphorhynchoidea) from Liaoning". Jiangsu Geology. 22 (4): 199–206.
68. "Dimorphodon." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 288–291.
69. Andres, Brian (2021-12-07). "Phylogenetic systematics of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea)". Journal of Vertebrate Paleontology. 41 (sup1): 203–217. doi:10.1080/02724634.2020.1801703. ISSN 0272-4634. S2CID 245078533.
70. Martill; Frey, E.; Diaz, G.C.; Bell, C.M. (2000). "Reinterpretation of a Chilean pterosaur and the occurrence of Dsungeripteridae in South America". Geological Magazine. 137 (1): 19–25. Bibcode:2000GeoM..137...19M. doi:10.1017/s0016756800003502. S2CID 128593191.
71. Seeley, H. G. (1875). "On an Ornithosaurian (Doratorhynchus validus) from the Purbeck Limestone of Langton near Swanage". Quarterly Journal of the Geological Society. 31 (1–4): 465–468. doi:10.1144/GSL.JGS.1875.031.01-04.35. S2CID 131057548.
72. "Dorygnathus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 292–295.
73. Xiaoli Wang, Shunxing Jiang, Junqiang Zhang, Xin Cheng, Xuefeng Yu, Yameng Li, Guangjin Wei & Xiaolin Wang, 2017, "New evidence from China for the nature of the pterosaur evolutionary transition", Scientific Reports 7, Article number: 42763
74. Holgado, Borja (2021-12-03). "On the validity of the genus Amblydectes Hooley 1914 (Pterodactyloidea, Anhangueridae) and the presence of Tropeognathinae in the Cambridge Greensand". Anais da Academia Brasileira de Ciências. 93 (suppl 2): e20201658. doi:10.1590/0001-3765202120201658. ISSN 0001-3765. PMID 34877964. S2CID 244884444.
75. "Dsungaripterus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 334–335.
76. Andres, B.; Ji, Q. (2008). "A new pterosaur from the Liaoning Province of China, the phylogeny of the Pterodactyloidea, and convergence in their cervical vertebrae". Palaeontology. 51 (2): 453–469. doi:10.1111/j.1475-4983.2008.00761.x.
77. Lü, Junchang; Qiang Ji (2005). "New azhdarchid pterosaur from the Early Cretaceous of western Liaoning". Acta Geologica Sinica. 79 (3): 301–307. doi:10.1111/j.1755-6724.2005.tb00893.x. S2CID 128958556.
78. Lü, J.C.; B.K. Zhang (2005). "New pterodactyloid pterosaur from the Yixian Formation of western Liaoning". Geological Review. 51 (4): 458–462.
79. Fernandes AE, Mateus O, Andres B, Polcyn MJ, Schulp AS, Gonçalves AO, Jacobs LL (2022). "Pterosaurs from the Late Cretaceous of Angola". Diversity. 14 (9). 741. doi:10.3390/d14090741.
80. "Eudimorphodon." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 280–281.
81. Romain Vullo; Jesús Marugán-Lobón; Alexander W. A. Kellner; Angela D. Buscalioni; Bernard Gomez; Montserrat de la Fuente; José J. Moratalla (2012). "A New Crested Pterosaur from the Early Cretaceous of Spain: The First European Tapejarid (Pterodactyloidea: Azhdarchoidea)". PLOS ONE. 7 (7): e38900. Bibcode:2012PLoSO...738900V. doi:10.1371/journal.pone.0038900. PMC 3389002. PMID 22802931.
82. Mátyás Vremir, Alexander W. A. Kellner, Darren Naish, Gareth J. Dyke (2013). "A New Azhdarchid Pterosaur from the Late Cretaceous of the Transylvanian Basin, Romania: Implications for Azhdarchid Diversity and Distribution". PLOS ONE. 8 (1): e54268. Bibcode:2013PLoSO...854268V. doi:10.1371/journal.pone.0054268. PMC 3559652. PMID 23382886.{{cite journal}}: CS1 maint: uses authors parameter (link)
83. Kessler, E.; Jurcsák, T. (1986). "New contributions to the knowledge of the Lower Cretaceous bird remains from Cornet (Romania)". Travaux du Muséum National d'Histoire Naturelle "Grigore Antipa". 28: 289–295.
84. Federico L. Agnolin & David Varricchio (2012). "Systematic reinterpretation of Piksi barbarulna Varricchio, 2002 from the Two Medicine Formation (Upper Cretaceous) of Western USA (Montana) as a pterosaur rather than a bird". Geodiversitas. 34 (4): 883–894. doi:10.5252/g2012n4a10. S2CID 56002643. Archived from the original on 2013-01-07.
85. Bonaparte, J., Schultz, C., and Soares, M. (2010). "Pterosauria from the Late Triassic of Southern Brazil." In : Bandyopadhyay, S. (ed). New Aspects of Mesozoic Biodiversity. Berlin/Heidelberg: Springer Pp. 63–71.
86. Kellner, A.W.A.; Holgado, B.; Grillo, O.; Pretto, F.A.; Kerber, L.; Pinheiro, F.L.; Soares, M.B.; Schultz, C.L.; Lopes, R.T.; Araújo, A.; Müller, R.T. (2022). "Reassessment of Faxinalipterus minimus, a purported Triassic pterosaur from southern Brazil with the description of a new taxon". PeerJ. 10: e13276. doi:10.7717/peerj.13276. PMC 9074864. PMID 35529502.
87. Wang, Xiaolin; Campos, Diogenes (October 2005). "Pterosaur diversity and faunal turnover in Cretaceous terrestrial ecosystems in China". Nature. 437 (7060): 875–879. Bibcode:2005Natur.437..875W. doi:10.1038/nature03982. PMID 16208369. S2CID 23146503.
88. Lü, J.; Fucha, X.; Chen, J. (2010). "A new scaphognathine pterosaur from the Middle Jurassic of western Liaoning, China". Acta Geoscientica Sinica. 31 (2): 263–266.
89. Adele H. Pentland; Stephen F. Poropat; Travis R. Tischler; Trish Sloan; Robert A. Elliott; Harry A. Elliott; Judy A. Elliott; David A. Elliott (2019). "Ferrodraco lentoni gen. et sp. nov., a new ornithocheirid pterosaur from the Winton Formation (Cenomanian–lower Turonian) of Queensland, Australia". Scientific Reports. 9 (1): Article number 13454. Bibcode:2019NatSR...913454P. doi:10.1038/s41598-019-49789-4. PMC 6776501. PMID 31582757.
90. Wang, X.; Kellner; Zhou, Z.; Campos (2007). "A new pterosaur (Ctenochasmatidae, Archaeopterodactyloidea) from the Lower Cretaceous Yixian Formation of China". Cretaceous Research. 28 (2): 2245–260. doi:10.1016/j.cretres.2006.08.004.
91. "Germanodactylus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 310–311.
92. Lü Junchang; Ji Qiang; Wei Xuefang; Liu Yongqing (2011). "A new ctenochasmatoid pterosaur from the Early Cretaceous Yixian Formation of western Liaoning, China". Cretaceous Research. 34: 26–30. doi:10.1016/j.cretres.2011.09.010.
93. "Gnathosaurus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 314–315.
94. Xiaolin Wang; Alexander W. A. Kellner; Shunxing Jiang; Xin Cheng (2012). "New toothed flying reptile from Asia: close similarities between early Cretaceous pterosaur faunas from China and Brazil". Naturwissenschaften. 99 (4): 249–257. Bibcode:2012NW.....99..249W. doi:10.1007/s00114-012-0889-1. PMID 22354475. S2CID 7323552.
95. Victoria M. Arbour; Philip J. Currie (2011). "An istiodactylid pterosaur from the Upper Cretaceous Nanaimo Group, Hornby Island, British Columbia, Canada". Canadian Journal of Earth Sciences. 48 (1): 63–69. Bibcode:2011CaJES..48...63S. doi:10.1139/E10-083.^{[permanent dead link]}
96. Romain Vullo, Eric Buffetaut & Michael J. Everhart (2012). "Reappraisal of Gwawinapterus beardi from the Late Cretaceous of Canada: a saurodontid fish, not a pterosaur". Journal of Vertebrate Paleontology. 32 (5): 1198–1201. doi:10.1080/02724634.2012.681078. S2CID 129180570.
97. Xiaolin Wang; Alexander W.A. Kellner; Shunxing Jiang; Qiang Wang; Yingxia Ma; Yahefujiang Paidoula; Xin Cheng; Taissa Rodrigues; Xi Meng; Jialiang Zhang; Ning Li; Zhonghe Zhou (2014). "Sexually Dimorphic Tridimensionally Preserved Pterosaurs and Their Eggs from China". Current Biology. 24 (12): 1323–1330. doi:10.1016/j.cub.2014.04.054. PMID 24909325.
98. Wang, X.; Lü, J. (2001). "Discovery of a pterodactloid pterosaur from the Yixian Formation of western Liaoning, China". Chinese Science Bulletin. 45 (12): 447–454.
99. Carpenter, K., Unwin, D.M., Cloward, K., Miles, C.A., and Miles, C. (2003). A new scaphognathine pterosaur from the Upper Jurassic Formation of Wyoming, USA. In: Buffetaut, E., and Mazin, J.- M. (eds.). Evolution and Palaeobiology of Pterosaurs. Geological Society of London, Special Publications 217:45–54.
100. Buffetaut, E.; Grigorescu, D.; Csiki, Z. (2002). "A new giant pterosaur with a robust skull from the latest Cretaceous of Romania". Naturwissenschaften. 89 (4): 180–184. Bibcode:2002NW.....89..180B. doi:10.1007/s00114-002-0307-1. PMID 12061403. S2CID 15423666.
101. Xiaolin Wang; Diogenes de Almeida Kellner (2008). "A primitive istiodactylid pterosaur (Pterodactyloidea) from the Jiufotang Formation (Early Cretaceous), northeast China". Zootaxa. 1813: 1–18. doi:10.11646/zootaxa.1813.1.1.
102. Dong, Z.-M. (1982). "On a new Pterosauria (Huanhepterus quingyangensis gen.et. sp.nov.) from Ordos, China". Vertebrata PalAsiatica. 20 (2): 115–121.
103. Lü, J.; Yuan, C. (2005). "New tapejarid pterosaur from Western Liaoning, China". Acta Geologica Sinica. 79 (4): 453–458. doi:10.1111/j.1755-6724.2005.tb00911.x. S2CID 129245203.
104. Borja Holgado; Rodrigo V. Pêgas; José Ignacio Canudo; Josep Fortuny; Taissa Rodrigues; Julio Company; Alexander W. A. Kellner (2019). "On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria". Scientific Reports. 9 (1): Article number 4940. Bibcode:2019NatSR...9.4940H. doi:10.1038/s41598-019-41280-4. PMC 6426928. PMID 30894614.
105. Xiaolin Wang; Taissa Rodrigues; Shunxing Jiang; Xin Cheng; Alexander W. A. Kellner (2014). "An Early Cretaceous pterosaur with an unusual mandibular crest from China and a potential novel feeding strategy". Scientific Reports. 4: Article number 6329. Bibcode:2014NatSR...4E6329W. doi:10.1038/srep06329. PMC 5385874. PMID 25210867.
106. Unwin, D. M. and Martill, D. M. (2007). "Pterosaurs of the Crato Formation." In Martill, D. M., Bechly, G. and Loveridge, R. F. (eds), The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press (Cambridge), pp. 475–524.
107. Wang, X.; Zhou, Z.; Zhang, F.; Xu, X. (2002). "A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and 'hairs' from Inner Mongolia, northeast China". Chinese Science Bulletin. 47 (3): 226–232. Bibcode:2002ChSBu..47..226W. doi:10.1360/02tb9054. S2CID 86641794.
108. Xin Cheng; Xiaolin Wang; Shunxing Jiang; Alexander W.A. Kellner (2012). "A new scaphognathid pterosaur from western Liaoning, China". Historical Biology. 24: 101–111. doi:10.1080/08912963.2011.635423. S2CID 128539358.
109. Lü Junchang & Bo Xue (2011). "A New Rhamphorhynchid Pterosaur (Pterosauria) from the Middle Jurassic Tiaojishan Formation of Western Liaoning, China". Acta Geologica Sinica. 85 (5): 977–983. doi:10.1111/j.1755-6724.2011.00531.x. S2CID 140196879.
110. Dong, Z.; Sun, Y.; Wu, S. (2003). "On a new pterosaur from the Lower Cretaceous of Chaoyang Basin, Western Liaoning, China". Global Geology. 22 (1): 1–7.
111. Cerqueira GM, Santos MA, Marks MF, Sayão JM, Pinheiro FL (2021). "A new azhdarchoid pterosaur from the Lower Cretaceous of Brazil and the paleobiogeography of the Tapejaridae". Acta Palaeontologica Polonica. 66. doi:10.4202/app.00848.2020..
112. Harris, J.D., and Carpenter, K. (1996). A large pterodactyloid from the Morrison Formation (Late Jurassic) of Garden Park, Colorado. Neues Jahrbuch für Geologie und Paläontologie Monatshefte 1996(8):473–484.
113. Alexander W.A. Kellner; Luiz C. Weinschütz; Borja Holgado; Renan A. M. Bantim; Juliana M. Sayão (2019). "A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert". Anais da Academia Brasileira de Ciências. 91 (Suppl. 2): e20190768. doi:10.1590/0001-3765201920190768. PMID 31432888.
114. Michael O’Sullivan; David M. Martill (2018). "Pterosauria of the Great Oolite Group (Bathonian, Middle Jurassic) of Oxfordshire and Gloucestershire, England". Acta Palaeontologica Polonica. Online edition. doi:10.4202/app.00490.2018.
115. Wang, X.; Kellner, A.W.A.; Jiang, S.; Cheng, X.; Meng, X.; Rodrigues, T. (2010). "New long-tailed pterosaurs (Wukongopteridae) from western Liaoning, China". Anais da Academia Brasileira de Ciências. 82 (4): 1045–1062. doi:10.1590/S0001-37652010000400024. PMID 21152776.
116. Brian Andres, James Clark & Xing Xu (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24 (9): 1011–1016. doi:10.1016/j.cub.2014.03.030. PMID 24768054.
117. Witton, Mark P. (2008). "A new azhdarchoid pterosaur from the Crato Formation (Lower Cretaceous, Aptian?) of Brazil". Palaeontology. 51 (6): 1289–1300. doi:10.1111/j.1475-4983.2008.00811.x.
118. Roy E. Smith; David M. Martill; Alexander Kao; Samir Zouhri; Nicholas Longrich (2020). "A long-billed, possible probe-feeding pterosaur (Pterodactyloidea: ?Azhdarchoidea) from the mid-Cretaceous of Morocco, North Africa". Cretaceous Research. 118: Article 104643. doi:10.1016/j.cretres.2020.104643. S2CID 225201538.
119. Chang-Fu Zhou; Ke-Qin Gao; Hongyu Yi; Jinzhuang Xue; Quanguo Li; Richard C. Fox (2017). "Earliest filter-feeding pterosaur from the Jurassic of China and ecological evolution of Pterodactyloidea". Royal Society Open Science. 4 (2): 160672. Bibcode:2017RSOS....460672Z. doi:10.1098/rsos.160672. PMC 5367317. PMID 28386425.
120. Wang, X.-L.; Zhou, Z.-H. (2003). "Two new pterodactyloid pterosaurs from the Early Cretaceous Jiufotang Formation of Western Liaoning, China". Vertebrata PalAsiatica. 41 (1): 34–41.
121. Dong, Z.; Lü, J. (2005). "A New Ctenochasmatid Pterosaur from the Early Cretaceous of Liaoning Province". Acta Geologica Sinica. 79 (2): 164–167. doi:10.1111/j.1755-6724.2005.tb00878.x. S2CID 129501612.
122. Kessler, E.; Jurcsák, T. (1984). "Fossil bird remains in the bauxite from Cornet (Romania, Bihor County)". Travaux du Muséum National d'Histoire Naturelle "Grigore Antipa". 25: 393–401.
123. Rodrigues, Taissa; Jiang, Shunxing; Cheng, Xin; Wang, Xiaolin; Kellner, Alexander W.A. (2015). "A new toothed pteranodontoid (Pterosauria, Pterodactyloidea) from the Jiufotang Formation (Lower Cretaceous, Aptian) of China and comments on Liaoningopterus gui Wang and Zhou, 2003". Historical Biology. 27 (6): 782–795. doi:10.1080/08912963.2015.1033417. S2CID 129062416.
124. Yizhi Xu; Shunxing Jiang; Xiaolin Wang (26 July 2022). "A new istiodactylid pterosaur, Lingyuanopterus camposi gen. et sp. nov., from the Jiufotang Formation of western Liaoning, China". PeerJ. 10: e13819. doi:10.7717/peerj.13819. PMC 9336611. PMID 35910775. Retrieved 26 July 2022.
125. Gervais, F. L. P. 1844. Remarques sur les oiseaux fossiles. Thèse, Fac. Sci., Univ. Paris. [Abstract in Bulletin of the Société Philomathique de Paris, pp. 67–70 & L'Institut 12:293–294.]
126. Buffetaut, E.; Lepage, J.-J.; Lepage, G. (1998). "A new pterodactyloid pterosaur from the Kimmeridgian of the Cap de la Hève (Normandy, France)". Geological Magazine. 135 (5): 719–722. Bibcode:1998GeoM..135..719B. doi:10.1017/s0016756898001575. S2CID 129836562.
127. Wang, L.; Li, L.; Duan, Y.; Cheng, S.L. (2006). "A new istiodactylid pterosaur from western Liaoning, China". Geological Bulletin of China. 25 (6): 737–740.
128. David W. E. Hone; Adam J. Fitch; Feimin Ma; Xing Xu (2020). "An unusual new genus of istiodactylid pterosaur from China based on a near complete specimen". Palaeontologia Electronica. 23 (1): Article number 23(1):a09. doi:10.26879/1015.
129. Frey, E., Martill, D., and Buchy, M. (2003). A new crested ornithocheirid from the Lower Cretaceous of northeastern Brazil and the unusual death of an unusual pterosaur. In: Buffetaut, E., and Mazin, J.-M. (eds.). Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publication 217:56–63. ISBN 1-86239-143-2.
130. David W. E. Hone (2020). "A review of the taxonomy and palaeoecology of the Anurognathidae (Reptilia, Pterosauria)". Acta Geologica Sinica (English Edition). 94 (5): 1676–1692. doi:10.1111/1755-6724.14585. S2CID 225169094.
131. Bantim, Renan A. M.; Saraiva, Antônio A. F.; Oliveira, Gustavo R.; Sayão, Juliana M. (2014). "A new toothed pterosaur (Pterodactyloidea: Anhangueridae) from the Early Cretaceous Romualdo Formation, NE Brazil". Zootaxa. 3869 (3): 201–223. doi:10.11646/zootaxa.3869.3.1. PMID 25283914.
132. Jensen, J. A.; Padian (1989). "Small pterosaurs and dinosaurs from the Uncompahgre fauna (Brushy Basin Member, Morrison Formation: ?Tithonian), late Jurassic, western Colorado". Journal of Paleontology. 63 (3): 363–374. doi:10.1017/S0022336000019533. S2CID 130246056.
133. Ross A. Elgin & Eberhard Frey (2011). "A new azhdarchoid pterosaur from the Cenomanian (Late Cretaceous) of Lebanon". Swiss Journal of Geosciences. 104: 21–33. doi:10.1007/s00015-011-0081-1. S2CID 128405107.
134. Alexander W. A. Kellner; Michael W. Caldwell; Borja Holgado; Fabio M. Dalla Vecchia; Roy Nohra; Juliana M. Sayão & Philip J. Currie (2019). "First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity". Scientific Reports. 9 (1): 17875. Bibcode:2019NatSR...917875K. doi:10.1038/s41598-019-54042-z. PMC 6884559. PMID 31784545. S2CID 208497557.
135. Romain Vullo; Géraldine Garcia; Pascal Godefroit; Aude Cincotta; Xavier Valentin (2018). "Mistralazhdarcho maggii, gen. et sp. nov., a new azhdarchid pterosaur from the Upper Cretaceous of southeastern France". Journal of Vertebrate Paleontology. 38 (4): (1)-(16). doi:10.1080/02724634.2018.1502670. S2CID 91265861.
136. Lü Junchang, Pu Hanyong, Xu Li, Wu Yanhua and Wei Xuefang (2012). "Largest toothed pterosaur skull from the Early Cretaceous Yixian Formation of Western Liaoning, China, with comments on the family Boreopteridae". Acta Geologica Sinica. 86 (2): 287–293. doi:10.1111/j.1755-6724.2012.00658.x. S2CID 130163272.{{cite journal}}: CS1 maint: uses authors parameter (link)
137. Padian; Horner, J.R.; de Ricqlès, A.J. (1993). "A new azhdarchid pterosaur from the Two Medicine Formation (Late Cretaceous, Campanian) of Montana, identified on the basis of bone histology". Journal of Vertebrate Paleontology. 13: 52A. doi:10.1080/02724634.1993.10011533.
138. Frey, E.; Buchy, M.-C.; Stinnesbeck, W.; González, A.G.; Stefano, A. (2006). "Muzquizopteryx coahuilensis n.g., n. sp., a nyctosaurid pterosaur with soft tissue preservation from the Coniacian (Late Cretaceous) of northeast Mexico (Coahuila)". Oryctos. 6: 19–39.
139. Molnar, Ralph E.; Thulborn, R.A. (2008). "An incomplete pterosaur skull from the Cretaceous of north-central Queensland, Australia". Arquivos do Museu Nacional, Rio de Janeiro. 65 (4): 461–470.
140. Sullivan, Robert M.; Fowler, Denver W. (2011). "Navajodactylus boerei, n. gen., n. sp., (Pterosauria, ?Azhdarchidae) from the Upper Cretaceous Kirtland Formation (upper Campanian) of New Mexico" (PDF). Fossil Record 3. New Mexico Museum of Natural History and Science, Bulletin. 53: 393–404.
141. Wang, X.; Kellner; Zhou, Z.; Campos, D.A. (2008). "Discovery of a rare arboreal forest-dwelling flying reptile (Pterosauria, Pterodactyloidea) from China". Proceedings of the National Academy of Sciences. 105 (6): 1983–1987. Bibcode:2008PNAS..105.1983W. doi:10.1073/pnas.0707728105. PMC 2538868. PMID 18268340.
142. Colbert, Edwin H. (1969). "A Jurassic pterosaur from Cuba" (PDF). American Museum Novitates (2370): 1–26. Retrieved 2007-03-03.
143. Jensen, J. A.; Ostrom, J. H. (1977). "A second Jurassic pterosaur from North America". Journal of Paleontology. 51 (4): 867–870.
144. Lu, J. (2009). "A baby pterodactyloid pterosaur from the Yixian Formation of Ningcheng, Inner Mongolia, China". Acta Geologica Sinica. 83 (1): 1–8. doi:10.1111/j.1755-6724.2009.00001.x. S2CID 85074573.
145. Marsh, O. C. 1881. "Note on American pterodactyls". American Journal of Science (Series 3) 21:342–343.
146. Marsh, O. C. 1876. Principal characters of American pterodactyls 2011-07-21 at the Wayback Machine. American Journal of Science, Series 3, 12:479–480.
147. "Nyctosaurus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 348–349.
148. Shu-an Ji (2020). "First record of Early Cretaceous pterosaur from the Ordos Region, Inner Mongolia, China". China Geology. 3 (1): 1–7. doi:10.31035/cg2020007.
149. von Sömmering, S. T. 1812. "Über einen Ornithocephalus oder über das unbekannten Thier der Vorwelt, dessen Fossiles Gerippe Collini im 5. Bande der Actorum Academiae Theodoro-Palatinae nebst einer Abbildung in natürlicher Grösse im Jahre 1784 beschrieb, und welches Gerippe sich gegenwärtig in der Naturalien-Sammlung der königlichen Akademie der Wissenschaften zu München befindet". Denkschriften der königlichen bayerischen Akademie der Wissenschaften München, mathematisch-physikalische Classe 3:89–158.
150. Fitzinger, L. J. 1843. Systema Reptilium. Braumüller et Seidel, Wien. 106 pages. [Pachyrhamphus reference on p. 35]
151. Shu’an Ji; Lifu Zhang (2020). "A new Early Cretaceous pterosaur from the Ordos region, Inner Mongolia". Earth Science Frontiers. 27 (6): 365–370. doi:10.13745/j.esf.sf.2020.6.14.
152. Martínez, R. N.; Andres, B.; Apaldetti, C.; Cerda, I. A. (2022). "The dawn of the flying reptiles: first Triassic record in the southern hemisphere". Papers in Palaeontology. 8 (2): e1424. doi:10.1002/spp2.1424. ISSN 2056-2799. S2CID 247494547.
153. Peters, D. 2005. Pterosaur ptrouble?. Prehistoric Times 72: 44–47.
154. von Arthaber, G. (1919). "Studien über Flugsaurier auf Grund der Bearbeitung des Wiener exemplars von Dorygnathus banthensis Theod Sp". Denkschriften der königlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftlichen Klasse 97:391–464. [German]
155. "Peteinosaurus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 282–283.
156. "New material of dsungaripterid pterosaurs (Pterosauria: Pterodactyloidea) from western Mongolia and its palaeoecological implications." Geological Magazine, 146(5): 690–700.
157. Pereda-Suberbiola, Xabier; Bardet, N., Jouve, S., Iarochène, M., Bouya, B., and Amaghzaz, M. (2003). "A new azhdarchid pterosaur from the Late Cretaceous phosphates of Morocco", in Buffetaut, E., and Mazin, J.-M. (eds.): "Evolution and Palaeobiology of Pterosaurs", Geological Society of London, Special Publications, 217. London: Geological Society of London, 80–90.
158. Varricchio, David J (2002). "A new bird from the Upper Cretaceous Two Medicine Formation of Montana". Canadian Journal of Earth Sciences. 39 (1): 19–26. Bibcode:2002CaJES..39...19V. doi:10.1139/e01-057.
159. Howse, S.C.B.; Milner, A.R. (1995). "The pterodactyloids from the Purbeck Limestone Formation of Dorset". Bulletin of the Natural History Museum, Geology Series. 51 (1): 73–88.
160. Fuentes Vidarte, C., Meijide Calvo, M. (2010). (PDF). Boletín Geológico y Minero. 121 (3): 311–328. Archived from the original (PDF) on 2011-07-20.{{cite journal}}: CS1 maint: multiple names: authors list (link)
161. "Preondactylus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 284–285.
162. Martins Neto, R.G. (1986). Pricesaurus megalodon nov. gen. nov. sp. (Pterosauria, Pterodactyloidea), Cretaceo Inferior, chapada do Araripe (NE-Brasil). Ciência e Cultura 38(7):756–757. [Portuguese]
163. Atanassov, M. N. 2002. Two new archosaur reptiles from the Late Triassic of Texas. PhD dissertation, Texas Tech University, Lubbock, Texas. 352 pages.
164. Seeley, H. G. 1869. Index to the Fossil Remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata, Arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge. 143 pages./Gray, J. E. 1845. (This book can be freely downloaded from Google Books.)
165. Lydekker, R. 1888. Catalogue of the fossil Reptilia and Amphibia in the British Museum (Natural History). I. London. pp. 2–42.
166. "Pteranodon." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 346–347 and 350–351.
167. "Pterodactylus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 298–301.
168. "Pterodaustro." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 342–345.
169. Jiang Shunxing & Wang Xiaolin (2011). "A new ctenochasmatid pterosaur from the Lower Cretaceous, western Liaoning, China". Anais da Academia Brasileira de Ciências. 83 (4): 1243–1249. doi:10.1590/s0001-37652011000400011. ISSN 0001-3765. PMID 22146956.
170. Czerkas, S.A., and Ji, Q. (2002). A new rhamphorhynchoid with a headcrest and complex integumentary structures. In: Czerkas, S.J. (Ed.). Feathered Dinosaurs and the Origin of Flight. The Dinosaur Museum:Blanding, Utah, 15–41. ISBN 1-932075-01-1.
171. Fischer von Waldheim, J. G. 1813. Zoognosia tabulis synopticus illustrata, in usum praelectionum Academiae Imperialis Medico-Chirurgicae Mosquenis edita. 3rd edition, volume 1. 466 pages.
172. Bonaparte, J.F., and Sanchez, T.M. (1975). "Restos de un pterosaurio Puntanipterus globosus de la formación La Cruz provincia San Luis, Argentina". Actas Primo Congresso Argentino de Paleontologia e Biostratigraphica 2:105–113. [Spanish]
173. Lü, J., Unwin, D.M., Zhao, B., Gao, C. and Shen, C. (2012). "A new rhamphorhynchid (Pterosauria: Rhamphorhynchidae) from the Middle/Upper Jurassic of Qinglong, Hebei Province, China" (PDF). Zootaxa. 3158: 1–19. doi:10.11646/zootaxa.3158.1.1.{{cite journal}}: CS1 maint: multiple names: authors list (link)
174. "Quetzalcoatlus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 352–355.
175. Stecher, Rico (2008). "A new Triassic pterosaur from Switzerland (Central Austroalpine, Grisons), Raeticodactylus filisurensis gen. et sp. nov". Swiss Journal of Geosciences. 101 (1): 185–201. doi:10.1007/s00015-008-1252-6. S2CID 128980861. Online First
176. Padian, K. (1984). Pterosaur remains from the Kayenta Formation (?early Jurassic) of Arizona 2007-09-27 at the Wayback Machine. Palaeontology 27(2):407–413. [if you get an I/O error message, push "OK" and it should work]
177. http://www.app.pan.pl/archive/published/app63/app004902018.pdf^{[bare URL PDF]}
178. "Rhamphorhynchus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 302–305.
179. Naish, D., Dyke, G., Cau, A., Escuillié, F., and Godefroit, P. (2012). "A gigantic bird from the Upper Cretaceous of Central Asia". Biology Letters. 8 (1): 97–100. doi:10.1098/rsbl.2011.0683. PMC 3259976. PMID 21835881.{{cite journal}}: CS1 maint: uses authors parameter (link)
180. De Buisonjé, P.H. (1980). "Santanadactylus brasiliensis nov.gen. nov.sp. a longnecked, large pterosaur from the Aptian of Brazil". Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen. Series B. 83 (2): 145–172.
181. "Scaphognathus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 308–309.
182. Fabio Marco Dalla Vecchia (2019). "Seazzadactylus venieri gen. et sp. nov., a new pterosaur (Diapsida: Pterosauria) from the Upper Triassic (Norian) of northeastern Italy". PeerJ. 7: e7363. doi:10.7717/peerj.7363. PMC 6661147. PMID 31380153.
183. Stanislas Rigal; David M. Martill; Steven C. Sweetman (2017). "A new pterosaur specimen from the Upper Tunbridge Wells Sand Formation (Cretaceous, Valanginian) of southern England and a review of Lonchodectes sagittirostris (Owen 1874)". In D. W. E. Hone; M. P. Witton; D. M. Martill (eds.). New Perspectives on Pterosaur Palaeobiology. Geological Society of London, Special Publications. Vol. 455. The Geological Society of London. pp. 221–232. Bibcode:2018GSLSP.455..221R. doi:10.1144/SP455.5. S2CID 133080548.
184. Lü J.; D.M. Unwin; Xu L.; Zhang X. (2008). "A new azhdarchoid pterosaur from the Lower Cretaceous of China and its implications for pterosaur phylogeny and evolution". Naturwissenschaften. 95 (9): 891–7. Bibcode:2008NW.....95..891L. doi:10.1007/s00114-008-0397-5. PMID 18509616. S2CID 13458087.
185. Wei, X.; Pêgas, R. V.; Shen, C.; Guo, Y.; Ma, W.; Sun, D.; Zhou, X. (2021). "Sinomacrops bondei, a new anurognathid pterosaur from the Jurassic of China and comments on the group". PeerJ. 9: e11161. doi:10.7717/peerj.11161. PMC 8019321. PMID 33850665.
186. Wang, X.; Zhou, Z. (2003). (PDF). Chinese Science Bulletin. 48 (1): 16–23. Bibcode:2003ChSBu..48...16W. doi:10.1007/bf03183326. S2CID 43993745. Archived from the original (PDF) on 2007-10-27.
187. Mader, B. J. and A. W. A. Kellner. 1999. "A new Anhanguerid pterosaur from the Cretaceous of Morocco". Boletim do Museu Nacional, Nova Série, Geologia 45:1–11.
188. "Sordes." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 318–319.
189. Soto M, Montenegro F, Toriño P, Mesa V, Perea D (2021). "A new ctenochasmatid (Pterosauria, Pterodactyloidea) from the late Jurassic of Uruguay". Journal of South American Earth Sciences. 111: Article 103472. Bibcode:2021JSAES.11103472S. doi:10.1016/j.jsames.2021.103472.
190. Rodrigo Pêgas; Borja Holgado; Maria Eduarda C. Leal (2019). "On Targaryendraco wiedenrothi gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids". Historical Biology: An International Journal of Paleobiology. 33 (8): 1–15. doi:10.1080/08912963.2019.1690482. S2CID 209595986.
191. Unwin, David M.; Heinrich, Wolf-Dieter (1999). "On a pterosaur jaw from the Upper Jurassic of Tendaguru (Tanzania)". Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaftliche Reihe. 2: 121–134.
192. Kellner; Campos (2002). "The function of the cranial crest and jaws of a unique pterosaur from the early Cretaceous of Brazil". Science. 297 (5580): 389–392. Bibcode:2002Sci...297..389K. doi:10.1126/science.1073186. PMID 12130783. S2CID 45430990.
193. Richards, T.M.; Stumkat, P.E.; Salisbury, S.W. (2021). "A new species of crested pterosaur (Pterodactyloidea, Anhangueridae) from the Lower Cretaceous (upper Albian) of Richmond, North West Queensland, Australia". Journal of Vertebrate Paleontology. 41 (3): e1946068. doi:10.1080/02724634.2021.1946068.
194. Arambourg, C. (1959). "Titanopteryx philadelphiae nov. gen., nov. sp. Ptérosaurien géant". Notes Mém. Moyen-Orient. 7: 229–234.
195. Campos, D.A. (2007). "Short note on the ingroup relationships of the Tapejaridae (Pterosauria, Pterodactyloidea)". Boletim do Museu Nacional. 75: 1–14.
196. Kellner, A.W.A., and Campos, D.A. (1988). "Sobre un novo pterossauro com crista sagital da Bacia do Araripe, Cretaceo Inferior do Nordeste do Brasil. (Pterosauria, Tupuxuara, Cretaceo, Brasil)." Anais da Academia Brasileira de Ciências, 60: 459–469. [in Portuguese]
197. Rodrigues, T.; Kellner, A. W. A. (2008). "Review of the pterodactyloid pterosaur Coloborhynchus". Zitteliana B. 28: 219–228.
198. David M. Martill (2011). "A new pterodactyloid pterosaur from the Santana Formation (Cretaceous) of Brazil". Cretaceous Research. In. 32 (2): 236–243. doi:10.1016/j.cretres.2010.12.008.^{[dead link]}
199. Czerkas, Stephen A.; Mickelson, Debra L. (2002). "The first occurrence of skeletal pterosaur remains in Utah". In Czerkas, Sylvia J. (ed.). Feathered Dinosaurs and the Origin of Flight. Blanding, Utah: The Dinosaur Museum. pp. 3–13. ISBN 978-1-932075-01-4.
200. Darren Naish, Martin Simpson, Gareth Dyke (2013). "A New Small-Bodied Azhdarchoid Pterosaur from the Lower Cretaceous of England and Its Implications for Pterosaur Anatomy, Diversity and Phylogeny". PLOS ONE. 8 (3): e58451. Bibcode:2013PLoSO...858451N. doi:10.1371/journal.pone.0058451. PMC 3601094. PMID 23526986.{{cite journal}}: CS1 maint: uses authors parameter (link)
201. Junchang Lü; Qingjin Meng; Baopeng Wang; Di Liu; Caizhi Shen; Yuguang Zhang (2017). "Short note on a new anurognathid pterosaur with evidence of perching behaviour from Jianchang of Liaoning Province, China". In D. W. E. Hone; M. P. Witton; D. M. Martill (eds.). New Perspectives on Pterosaur Palaeobiology. Geological Society of London, Special Publications. Vol. 455. The Geological Society of London. pp. 95–104. Bibcode:2018GSLSP.455...95L. doi:10.1144/SP455.16. S2CID 219196969.
202. Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". Paleontological Journal. 42 (6): 634–642. doi:10.1134/S0031030108060099. S2CID 129558986.
203. Andres, Brian; Langston, Wann (2021-12-14). "Morphology and taxonomy of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea)". Journal of Vertebrate Paleontology. 41 (sup1): 46–202. doi:10.1080/02724634.2021.1907587. ISSN 0272-4634. S2CID 245125409.
204. Codorniú, Laura; Gasparini, Zulma (2013). "The Late Jurassic pterosaurs from northern Patagonia, Argentina". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 399–408. doi:10.1017/S1755691013000388. S2CID 130885184.
205. David M. Martill; Mick Green; Roy E. Smith; Megan L. Jacobs; John Winch (2020). "First tapejarid pterosaur from the Wessex Formation (Wealden Group: Lower Cretaceous, Barremian) of the United Kingdom". Cretaceous Research. 113: Article 104487. doi:10.1016/j.cretres.2020.104487. S2CID 219099220.
206. Bakker, R.T. (1994) Unearthing the Jurassic. In: Science Year 1995. World Book Inc.:Chicago, London, Sydney, Toronto, 76–89. ISBN 0-7166-0595-3.
207. David M. Martill; David M. Unwin; Nizar Ibrahim; Nick Longrich (2017). "A new edentulous pterosaur from the Cretaceous Kem Kem beds of south eastern Morocco". Cretaceous Research. 84: 1–12. doi:10.1016/j.cretres.2017.09.006. hdl:2381/41058.
208. Lü, J., Ji, S., Yuan, C., Gao, Y., Sun, Z. and Ji, Q. 2006. New pterodactyloid pterosaur from the Lowe Cretaceous Yixian Formation of Western Liaoning, p. 195–203. In: Lü, J., Kobayashi, Y., Huang, D. and Lee, Y. (eds), Papers form the 2005 Heyuan International Dinosaur Symposium. Beijing: Geological Publishing House.
209. Cai, Z.; Wei, F. (1994). "On a new pterosaur (Zhejiangopterus linhaiensis gen. et sp. nov.) from Upper Cretaceous in Linhai, Zhejiang, China". Vertebrata PalAsiatica. 32: 181–194.
210. Lü, J. (2010). "A new boreopterid pterodactyloid pterosaur from the Early Cretaceous Yixian Formation of Liaoning Province, northeastern Chian". Acta Geologica Sinica. 84 (2): 241–246. doi:10.1111/j.1755-6724.2010.00204.x. S2CID 140600398.
211. Hwang, K.-G.; et al. (2002). "New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, southwestern Korea". Geological Magazine. 139 (4): 421–435. Bibcode:2002GeoM..139..421H. doi:10.1017/s0016756802006647. S2CID 54996027.
212. Kim, J. Y.; et al. (2006). "The oldest record of webbed bird and pterosaur tracks from South Korea (Cretaceous Haman Formation, Changseon and Sinsu Islands): More evidence of high avian diversity in East Asia". Cretaceous Research. 27 (1): 56–69. doi:10.1016/j.cretres.2005.10.005.

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