Isaria

Isaria is a genus of fungi mostly in the order Hypocreales and family Clavicipitaceae, or by some authorities the Cordycipitaceae.^[1] It includes a large number of entomopathogenic species, some of them exploited as biopesticides (e.g. I. fumosorosea): often previously assigned to the genus Paecilomyces.

Isaria
Isaria sinclairii showing the fruiting bodies and parasitised cicada nymphs
Scientific classification
Domain:	Eukaryota
Kingdom:	Fungi
Division:	Ascomycota
Class:	Sordariomycetes
Order:	Hypocreales
Family:	Cordycipitaceae
Genus:	Isaria Persoon in Roemer, 1794

The teleomorph of this genus appears to be Ophiocordyceps.^[2]

Moth infected with Isaria farinosa.

Species

The Encyclopaedia of Life lists the following species:^[3]

• Isaria abietina Höhn. 1909
• Isaria abutii Lloyd 1923
• Isaria acaricida Pat.
• Isaria acervata Massee 1901
• Isaria acicularis Pat. 1897
• Isaria acuta Preuss
• Isaria agaricina Pers. 1797
  • Isaria agaricina var. agaricina Pers. 1797
  • Isaria agaricina var. petiolaris Alb. & Schwein. 1805
• Isaria aggregata Cooke & Massee 1890
• Isaria alba J. F. H. Beyma 1928
• Isaria albida (Fr.) Sacc.
• Isaria albiziae Pat.
• Isaria alborosea Penz. & Sacc. 1904
• Isaria ambigua P. Karst.
• Isaria americana P. Karst.
• Isaria amoene-rosea Henn. 1902
• Isaria amorpha Höhn. 1909
• Isaria anisopliae (Metschn.) R. H. Pettit 1895
  • Isaria anisopliae var. americana R.H. Pettit 1895
  • Isaria anisopliae var. anisopliae (Metschn.) R.H. Pettit 1895
• Isaria arachnicida Speg. 1898
• Isaria arachnophila Ditmar 1817
• Isaria aranearum Schwein. 1822
• Isaria araneosa Speg. 1910
• Isaria arborea Pat.
• Isaria arbuscula Bres. & Roum. or Har. 1893
• Isaria arcyrioides Berk. & Broome 1875
• Isaria argentina Speg. 1898
• Isaria asamaensis Aoki 1957
• Isaria aspergilliformis Rostr.
• Isaria aurantiaca Berk. & Broome 1875
• Isaria aviaria Sacc.
• Isaria barberi Giard 1894
• Isaria basili (Taylor) Kobayasi 1941
• Isaria bialowieziensis Koval 1977
• Isaria bicolor Schwein. 1832
• Isaria brachiata (Batsch) Schumach. 1803
• Isaria brevis Berk. & M.A. Curtis 1874
• Isaria briardii (Vuill.) Sacc. & D. Sacc.
• Isaria bulbosa Nees 1816
• Isaria buntingii Lloyd 1916
• Isaria buquetii (Mont. & C. P. Robin) Lloyd 1922
• Isaria byssoidea Pers.
• Isaria calva (Alb. & Schwein.) Fr. 1832
• Isaria candida Schwein. 1832
• Isaria capitata Ellis & Everh. 1898
• Isaria carnea Pers. 1796 or Schwein.
• Isaria cateniannulata (Z.Q. Liang) Samson & Hywel-Jones 2005
• Isaria cateniobliqua (Z.Q. Liang) Samson & Hywel-Jones 2005
• Isaria ceratoides Speg. 1880
• Isaria chrysopoda Bres. 1900
• Isaria cicadae Miq. 1838
• Isaria cinnabarina Preuss
• Isaria citrina Pers. 1801
• Isaria citrinula Speg. 1910
• Isaria cladogena Sacc.
• Isaria clavata Ditmar 1817
• Isaria clonostachoides F. J. Pritch. & Porte 1922
• Isaria cocoa Lloyd 1920
• Isaria coleopterorum (Samson & H.C. Evans) Samson & Hywel-Jones 2005
• Isaria congesta Berk. & M.A. Curtis 1874 or Berk. & Broome 1875
• Isaria congregata Berk. & Broome 1875
• Isaria corallina Fr.
• Isaria coralloidea Kalchbr. & Cooke 1880
• Isaria cornea Lloyd 1919
• Isaria cosmopsaltriae Yasuda 1918
• Isaria crassa Pers. 1797
• Isaria cretacea J. F. H. Beyma 1935
• Isaria crinita Lloyd 1920
• Isaria cryptotympanae Sawada 1959
• Isaria cuneispora Boud. 1887
• Isaria dedawensis Henn.
• Isaria dendroidea Jungh. 1838
• Isaria densa (Link) Giard 1891
• Isaria destructor Metschn.
• Isaria dubia Delacr. 1893
• Isaria dussii Pat. 1900
• Isaria edessicola Speg. 1910
• Isaria edwalliana Henn. 1904
• Isaria elata (Kalchbr.) Sacc. & Traverso 1910
• Isaria elegans Beeli 1924
• Isaria elegantula Syd. 1909
• Isaria eleutheratorum Nees 1816
• Isaria empetri Lind 1928
• Isaria epiphylla Pers. 1797
  • Isaria epiphylla var. acuta Tassi
  • Isaria epiphylla var. epiphylla Pers. 1797
  • Isaria epiphylla var. fasciculata Sommerf.
• Isaria episphaeria Desm. 1843
• Isaria erastalidis Yasuda
• Isaria eriopoda Syd. 1912
• Isaria eristalidis Yasuda 1921
• Isaria exigua (Brond.) Sacc. 1892
• Isaria exoleta Fr. 1832
• Isaria farinosa (Holmsk.) Fr. 1832
  • Isaria farinosa f. crassa (Pers.) Fr. 1832
  • Isaria farinosa f. farinosa (Holmsk.) Fr. 1832
  • Isaria farinosa f. truncata (Pers.) Fr. 1832
  • Isaria farinosa f. velutipes (Link) Fr. 1832
  • Isaria farinosa var. citriformis Kobayasi 1941
  • Isaria farinosa var. farinosa (Holmsk.) Fr. 1832
• Isaria fasciculata Sommerf. 1826
• Isaria felina (DC.) Fr. 1832
  • Isaria felina var. aviaria Sacc. 1882
  • Isaria felina var. cuniculina Ferraris 1910
  • Isaria felina var. domestica Speg. 1912
  • Isaria felina var. felina (DC.) Fr. 1832
  • Isaria felina var. pirina Ellis & Marchal
  • Isaria felina var. pirina Marchal & É.J. Marchal 1921
  • Isaria felina var. suina Sacc. 1882
• Isaria filamentosa Sacc. 1886
• Isaria filiformis Wallr. 1833
  • Isaria filiformis var. cladogena Sacc. 1886
  • Isaria filiformis var. filiformis Wallr. 1833
• Isaria fimicola Sternon 1923
• Isaria flabelliformis (Schwein.) Lloyd 1916
• Isaria flavoviridis Rick 1922
• Isaria floccosa Berk. & Broome or Fr. 1829
• Isaria friesii Mont. 1836
• Isaria froggattii Lloyd 1922
• Isaria fruticosa Demelius 1923
• Isaria fuciformis Berk. 1872
• Isaria fulvipes Peck 1879
• Isaria fumosorosea Wize 1904
• Isaria funicularis Wallr.
• Isaria furcata Schwein. 1832
• Isaria furcellata G. Martin
• Isaria fusca J. N. Kapoor & Munjal 1967
• Isaria geophila Speg. 1898
• Isaria ghanensis (Samson & H.C. Evans) Samson & Hywel-Jones 2005
• Isaria gigantea Mont. 1856
• Isaria glauca Ditmar 1817
  • Isaria glauca var. americana P. Karst.
  • Isaria glauca var. glauca Ditmar 1817
• Isaria glaucocephala Link 1825
• Isaria globosa Schwein. 1832
• Isaria gossypina Pat.
• Isaria gracilioides Kobayasi 1941
• Isaria gracilis Speg. 1886
• Isaria gracilis Vosseler 1902
• Isaria graminiperda Berk. & F. Muell. 1873
  • Isaria graminiperda var. fuciformis (Berk.) Cooke 1892
  • Isaria graminiperda var. graminiperda Berk. & F. Muell. 1873
• Isaria grassa Pers.
• Isaria guignardii Maheu 1906
  • Isaria guignardii f. guignardii Maheu 1906
  • Isaria guignardii f. major Martinez{?} & Guinea 1931
• Isaria hariotii Arnaud 1915
• Isaria holmbergii Speg. 1880
• Isaria hydnoides Link 1825
• Isaria hypoxyli Kalchbr. 1865
• Isaria hystricina P. Karst.
• Isaria intricata Fr. 1832
• Isaria japonica Yasuda 1915
• Isaria javanica (Frieder. & Bally) Samson & Hywel-Jones 2005
• Isaria kogane Hasegawa & Koyama 1941
• Isaria kunitatiensis Kobayasi 1941
• Isaria lanuginosa Petch 1917
• Isaria lecaniicola Jaap 1909
• Isaria leprosa Fr. 1832
  • Isaria leprosa var. corralina (Fr.) Sacc. 1886
  • Isaria leprosa var. leprosa Fr. 1832
  • Isaria leprosa ß corralina Fr. 1832
• Isaria locusticola Z.Q. Liang, X.Y. He & Y.F. Han 2008
• Isaria lutea (Brond.) Sacc. 1892
• Isaria macroscyticola Kobayasi 1941
• Isaria melanopus Speg. 1889
• Isaria meliolae Hansf. 1943
• Isaria micromegala Berl. 1887
  • Isaria micromegala var. micromegala Berl. 1887
  • Isaria micromegala var. ulmicola Cif. 1922
• Isaria microscopica Grev. 1823
• Isaria mitruliformis Henn. 1906
• Isaria mokanshawii Lloyd 1919
• Isaria monilioides Alb. & Schwein. 1805
• Isaria moriokaensis Aoki 1957
• Isaria mucida Pers. 1794 or Schumach. 1803
• Isaria murina Oudem. 1892
• Isaria musasiensis Aoki 1942
• Isaria muscigena Cooke & Müll. {?}
• Isaria mycelioides Lloyd 1923
• Isaria myrmicidae Lloyd 1920
• Isaria nigra Yakush. & Kumaz. 1930
• Isaria nigripes Schwein. 1832
• Isaria nipponica Kobayasi 1939
• Isaria ochracea Boud. 1903
• Isaria odora Raf. 1813
• Isaria oncopterae McAlpine
• Isaria oncotympanae Kobayasi 1939
• Isaria orthopterorum Petch 1933
• Isaria ovi Teng 1932
• Isaria pachylomerica Kawam. 1927
• Isaria palmae F. Stevens & C. J. King 1927
• Isaria palmatifida Henn. 1902
• Isaria patrobus Lloyd 1924
• Isaria pattersonii Massee 1912
• Isaria peckoltii Lloyd 1919
• Isaria penicilliformis Peck 1898
• Isaria perexigua Kobayasi 1941
• Isaria perpusilla Speg. 1922
• Isaria phalangiophila Link 1825
• Isaria pistillariiformis Pat.
• Isaria plumosa Cooke
• Isaria psychidae A. Evans
• Isaria puberula Berk. 1841
• Isaria pulcherrima Berk. & Broome 1875
• Isaria radians Berk.
• Isaria radiata Berk. & M. A. Curtis 1874
• Isaria ramosissima Zoll.
• Isaria repens Cooke
• Isaria rhodosperma Bres.
• Isaria rosea (Berk. & M. A. Curtis) Sacc.
• Isaria saccharina Pers. 1801
• Isaria saussurei Cooke 1892
• Isaria schweinitzii Sacc.
• Isaria shiotae Kuru 1932
• Isaria sinclairii (Berk.) Lloyd 1923
• Isaria smilanensis Wize 1904
• Isaria sphaerocephala Petch 1942
• Isaria sphecophila Ditmar 1817
• Isaria sphingophila Link 1824
• Isaria sphingum Schwein. 1822
• Isaria spiculosa Wallr.
• Isaria splendens Rabenh. 1844
• Isaria spumarioides Cooke
• Isaria stellata Cooke
• Isaria stemonitis Pers. 1797
• Isaria stenobothri Hollande & Moreau 1922
• Isaria stenocori (Quél.) Kobayasi 1982
• Isaria stilbiformis Speg. 1879
• Isaria straminipes Ellis & Everh. 1888
• Isaria strigosa Fr. 1832
• Isaria subsimplex Schumach. 1803
• Isaria suffruticosa Cooke & Massee 1890
• Isaria suina Sacc.
• Isaria sulfurea Fiedl. 1859 : Isaria sulphurea is an Orthographic variant
  • Isaria sulfurea var. ossicola Speg. 1912
  • Isaria sulfurea var. sulfurea Fiedl. 1859
  • Isaria sulphurea var. sulphurea Fiedl. 1859 (legitimate)^[4]
• Isaria surinamensis Vosseler 1902
• Isaria tartarea Wallr.
• Isaria tax Kumar D, Roy CS, Khan ZR, Yazdani, Hameed, Mahmood M. 1983
• Isaria tenella (Sacc.) Giard 1891
• Isaria tenuipes Peck 1879
• Isaria tenuis F. Heim 1893
• Isaria terrestris Fr. 1825
• Isaria thelephoroides Bres. 1911
• Isaria thyrosoidea Penz. & Sacc. 1904
• Isaria tinearum Speg. 1898
• Isaria tomentella Fr. 1832
• Isaria tremellosa Schumach. 1803
• Isaria truncata Pers. 1801 or Briard
  • Isaria truncata var. damaecornu Pers. 1822
  • Isaria truncata var. truncata Pers. 1801
• Isaria umbrina Pers. 1801
  • Isaria umbrina var. parasitica Alb. & Schwein. 1805
  • Isaria umbrina var. umbrina Pers. 1801
• Isaria valsicola Kirschst. 1939
• Isaria vassiljevae Koval 1967
• Isaria velutipes Link 1809
• Isaria verticillata G. F. Atk.
• Isaria vexans R.H. Pettit 1895
• Isaria virescens Elenkin & A.L. Danilov 1922
• Isaria virginiensis Ellis & Everh. 1893
• Isaria xylariiformis Lloyd 1923
• Isaria yokohamensis Kobayasi 1941

Species formerly described in Isaria

• Family Ophiocordycipitaceae
  • Genus Purpureocillium: Isaria takamizusanensis and I. atypicola.

References

1. eol.org http://eol.org/pages/36823/overview:. Retrieved 2017-06-14. {{cite web}}: Missing or empty |title= (help)^{[title missing]}
2. Sung G-H, Hywel-Jones NL, Sung J-M, Luangsa-ard JJ, Shrestha B, Spatafora JW (2007). "Phylogenetic classification of Cordyceps and the clavicipitaceous fungi". Studies in Mycology. 57: 5–59. doi:10.3114/sim.2007.57.01. PMC 2104736. PMID 18490993.
3. EoL: Isaria
4. MycoBank

•   Media related to Isaria at Wikimedia Commons
•   Data related to Clavicipitaceae at Wikispecies