Amphibamidae has traditionally included small-bodied, terrestrial dissorophoids. The name is attributed to Moodie (1909),[2] but it was rarely used because it originally referred only to Amphibamus. Similar monogeneric families were also erected for other small, terrestrial dissorophoids (e.g., Doleserpetontidae), and most of the taxa now recognized as amphibamiforms were placed within the Dissorophidae.
Clack & Milner (1993) revived the Amphibamidae to include Amphibamus, Platyrhinops, Doleserpeton, and Tersomius[3]. Daly (1994) further expanded the composition of the Amphibamidae to include the newly described Eoscopus as well as the Early Triassic form Micropholis.[4] She suggested that the micromelerpetids were also amphibamids, which has not been validated by more recent workers. Subsequent phylogenetic work verified the monophyly of the Amphibamidae,[5] with recent analyses also recovering the branchiosaurids nested within the amphibamids.[6]
Schoch (2018) erected the new clade Amphibamiformes to include the traditional amphibamids and the nested branchiosaurids and subsequently restricted the Amphibamidae to two taxa: Doleserpeton annectens from the Dolese Brothers Limestone Quarry near Richards Spur, Oklahoma and Amphibamus grandiceps from Mazon Creek, Illinois. These taxa are united by several features, such as a medially expanded choana.
Phylogeny of amphibamiforms from Schoch (2018)edit
The Amphibamidae have typically been recovered close to some or all of the lissamphibian crown. Until the description of Gerobatrachus, Doleserpeton was considered to be the closest extinct relative to the crown group.[7] Below is a modified cladogram from Anderson et al. (2008) showing Batrachia nested in the Amphibamidae, with Gerobatrachus as the sister taxon of Batrachia (anurans and caudates) and Doleserpeton and Amphibamus as successive outgroups:[7]
There is continued debate over the origin of lissamphibians, including whether they are monophyletic or whether batrachians and caecilians are descended from different clades of tetrapods or temnospondyls.[8] If they are monophyletic, there is also a debate about which clade they are nested in, with some hypotheses suggesting a lepospondyl origin, in which case amphibamids would not be closely related to any extant amphibians.[9]
Referencesedit
^ abSchoch, Rainer R. (2018). "The putative lissamphibian stem-group: phylogeny and evolution of the dissorophoid temnospondyls". Journal of Paleontology. 93 (1): 137–156. doi:10.1017/jpa.2018.67. ISSN 0022-3360.
^Moodie, Roy L. (1909). "A contribution to a monograph of the extinct Amphibia of North America. New forms from the Carboniferous". Journal of Geology. 17 (1): 38–82. Bibcode:1909JG.....17...38M. doi:10.1086/621585.
^Clack, Jennifer A.; Milner, Andrew R. (1993). "Platyrhinops from the Upper Carboniferous of Linton and Nyrany and the family Amphibamidae (Amphibia: Temnospondyli)". New Research on Permo-Carboniferous Faunas. 29: 185–191.
^Daly, Eleanor (1994). "The Amphibamidae (Amphibia: Temnospondyli), with a description of a new genus from the Upper Pennsylvanian of Kansas". The University of Kansas Miscellaneous Publications. 85: 1–59.
^Schoch, Rainer R.; Rubidge, Bruce S. (2005). "The amphibamid Micropholis from the Lystrosaurus Assemblage Zone of South Africa". Journal of Vertebrate Paleontology. 25 (3): 502–522. doi:10.1671/0272-4634(2005)025[0502:tamftl]2.0.co;2.
^Schoch, Rainer R.; Milner, Andrew R. (2008). "The intrarelationships and evolutionary history of the temnospondyl family branchiosauridae". Journal of Systematic Palaeontology. 6 (4): 409–431. doi:10.1017/s1477201908002460. ISSN 1477-2019. S2CID 86207414.
^ abAnderson, J.S.; Reisz, R.R.; Scott, D.; Fröbisch, N.B.; Sumida, S.S. (2008). (PDF). Nature. 453 (7194): 515–518. Bibcode:2008Natur.453..515A. doi:10.1038/nature06865. PMID 18497824. S2CID 205212809. Archived from the original (PDF) on 2015-06-26. Retrieved 2015-06-25.
^Pardo, Jason D.; Small, Bryan J.; Huttenlocker, Adam K. (2017). "Stem caecilian from the Triassic of Colorado sheds light on the origins of Lissamphibia". Proceedings of the National Academy of Sciences. 114 (27): E5389–E5395. doi:10.1073/pnas.1706752114. ISSN 0027-8424. PMC5502650. PMID 28630337.
^Marjanović, David; Laurin, Michel (2019). "Phylogeny of Paleozoic limbed vertebrates reassessed through revision and expansion of the largest published relevant data matrix". PeerJ. 6 (e5565): e5565. doi:10.7717/peerj.5565. PMC6322490. PMID 30631641.
amphibamidae, ancient, family, dissorophoid, temnospondyls, known, from, late, carboniferous, early, permian, strata, united, states, temporal, range, late, carboniferous, early, permian, possible, descendant, taxon, lissamphibia, survives, present, preꞒ, amph. The Amphibamidae are an ancient family of dissorophoid temnospondyls known from Late Carboniferous Early Permian strata in the United States 1 AmphibamidaeTemporal range Late Carboniferous Early Permian 311 45 289 Ma Possible descendant taxon Lissamphibia survives to present PreꞒ Ꞓ O S D C P T J K Pg N Amphibamus grandiceps Scientific classification Domain Eukaryota Kingdom Animalia Phylum Chordata Order Temnospondyli Clade Amphibamiformes Family AmphibamidaeMoodie 1909 Subgroups See text Synonyms Doleserpetontidae Bolt 1969 Peliontidae Cope MS Contents 1 Classification 1 1 Phylogeny of amphibamiforms from Schoch 2018 1 2 Relationship to the Batrachia 2 References 3 External linksClassification editAmphibamidae has traditionally included small bodied terrestrial dissorophoids The name is attributed to Moodie 1909 2 but it was rarely used because it originally referred only to Amphibamus Similar monogeneric families were also erected for other small terrestrial dissorophoids e g Doleserpetontidae and most of the taxa now recognized as amphibamiforms were placed within the Dissorophidae Clack amp Milner 1993 revived the Amphibamidae to include Amphibamus Platyrhinops Doleserpeton and Tersomius 3 Daly 1994 further expanded the composition of the Amphibamidae to include the newly described Eoscopus as well as the Early Triassic form Micropholis 4 She suggested that the micromelerpetids were also amphibamids which has not been validated by more recent workers Subsequent phylogenetic work verified the monophyly of the Amphibamidae 5 with recent analyses also recovering the branchiosaurids nested within the amphibamids 6 Schoch 2018 erected the new clade Amphibamiformes to include the traditional amphibamids and the nested branchiosaurids and subsequently restricted the Amphibamidae to two taxa Doleserpeton annectens from the Dolese Brothers Limestone Quarry near Richards Spur Oklahoma and Amphibamus grandiceps from Mazon Creek Illinois These taxa are united by several features such as a medially expanded choana Phylogeny of amphibamiforms from Schoch 2018 edit Xerodromes Olsoniformes Amphibamiformes Eoscopus Platyrhinops Micropholidae Tersomius Pasawioops Micropholis Amphibamidae Doleserpeton Amphibamus Gerobatrachus Georgenthalia Lissamphibia Eocaecilia Karaurus Triadobatrachus Branchiosauridae Branchiosaurus Apateon Leptorophus Schoenfelderpeton 1 Relationship to the Batrachia edit The Amphibamidae have typically been recovered close to some or all of the lissamphibian crown Until the description of Gerobatrachus Doleserpeton was considered to be the closest extinct relative to the crown group 7 Below is a modified cladogram from Anderson et al 2008 showing Batrachia nested in the Amphibamidae with Gerobatrachus as the sister taxon of Batrachia anurans and caudates and Doleserpeton and Amphibamus as successive outgroups 7 Amphibamidae Tersomius Micropholis Eoscopus Platyrhinops Amphibamus Doleserpeton Gerobatrachus Batrachia Anura Triadobatrachus Caudata Albanerpetontidae There is continued debate over the origin of lissamphibians including whether they are monophyletic or whether batrachians and caecilians are descended from different clades of tetrapods or temnospondyls 8 If they are monophyletic there is also a debate about which clade they are nested in with some hypotheses suggesting a lepospondyl origin in which case amphibamids would not be closely related to any extant amphibians 9 References edit a b Schoch Rainer R 2018 The putative lissamphibian stem group phylogeny and evolution of the dissorophoid temnospondyls Journal of Paleontology 93 1 137 156 doi 10 1017 jpa 2018 67 ISSN 0022 3360 Moodie Roy L 1909 A contribution to a monograph of the extinct Amphibia of North America New forms from the Carboniferous Journal of Geology 17 1 38 82 Bibcode 1909JG 17 38M doi 10 1086 621585 Clack Jennifer A Milner Andrew R 1993 Platyrhinops from the Upper Carboniferous of Linton and Nyrany and the family Amphibamidae Amphibia Temnospondyli New Research on Permo Carboniferous Faunas 29 185 191 Daly Eleanor 1994 The Amphibamidae Amphibia Temnospondyli with a description of a new genus from the Upper Pennsylvanian of Kansas The University of Kansas Miscellaneous Publications 85 1 59 Schoch Rainer R Rubidge Bruce S 2005 The amphibamid Micropholis from the Lystrosaurus Assemblage Zone of South Africa Journal of Vertebrate Paleontology 25 3 502 522 doi 10 1671 0272 4634 2005 025 0502 tamftl 2 0 co 2 Schoch Rainer R Milner Andrew R 2008 The intrarelationships and evolutionary history of the temnospondyl family branchiosauridae Journal of Systematic Palaeontology 6 4 409 431 doi 10 1017 s1477201908002460 ISSN 1477 2019 S2CID 86207414 a b Anderson J S Reisz R R Scott D Frobisch N B Sumida S S 2008 A stem batrachian from the Early Permian of Texas and the origin of frogs and salamanders PDF Nature 453 7194 515 518 Bibcode 2008Natur 453 515A doi 10 1038 nature06865 PMID 18497824 S2CID 205212809 Archived from the original PDF on 2015 06 26 Retrieved 2015 06 25 Pardo Jason D Small Bryan J Huttenlocker Adam K 2017 Stem caecilian from the Triassic of Colorado sheds light on the origins of Lissamphibia Proceedings of the National Academy of Sciences 114 27 E5389 E5395 doi 10 1073 pnas 1706752114 ISSN 0027 8424 PMC 5502650 PMID 28630337 Marjanovic David Laurin Michel 2019 Phylogeny of Paleozoic limbed vertebrates reassessed through revision and expansion of the largest published relevant data matrix PeerJ 6 e5565 e5565 doi 10 7717 peerj 5565 PMC 6322490 PMID 30631641 External links editAmphibamidae in the Paleobiology Database Retrieved from https en wikipedia org w index php title Amphibamidae amp oldid 1215041619, wikipedia, wiki, book, books, library,
