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Helochares

April 11, 2024

Helochares is a genus of water scavenger beetles in the family Hydrophilidae, represented by 161 described species. It is distributed across the Afrotropical, Australasian, Indo-Malayan, Nearctic, and Palearctic realms. [1]

Helochares
Helochares lividus
Scientific classification
Kingdom:
Animalia
Phylum:
Arthropoda
Class:
Insecta
Order:
Coleoptera
Family:
Hydrophilidae
Subfamily:
Acidocerinae
Genus:
Helochares

Mulsant, 1844
Diversity
158 species + 3 subspecies
Synonyms
  • Helophilus Mulsant, 1844
  • Pylophilus Motschulsky, 1845
  • Hydrobaticus MacLeay, 1871
  • Helocharimorphus Kuwert, 1890
  • Graphelochares Kuwert, 1890
  • Grapidelochares Ganglbauer, 1904

Taxonomy edit

Helochares is one of the largest and most taxonomically problematic genera within the Acidocerinae. It was originally described by Étienne Mulsant in 1844. [2]

For a long time Helochares contained five subgenera: Batochares (currently recognized as a separate genus [3]), Helochares, Helocharimorphus (now synonymized under Helochares [1]), Hydrobaticus (now synonymized under Helochares [1]), and Sindolus (currently regognized as a separate genus [3]).

Several taxa that used to be recognized as typical Helochares are now assigned to newly created genera (e.g., Colossochares, Novochares) or re-assigned to exinsting genera (i.e., Peltochares), based on a phylogenetic analysis based on molecular data. [3]

Description edit

As currently defined, Helochares contains small to medium-sized beetles (2–7 mm), which are yellowish to brown in coloration. There is a lot of variation in the impression of the elytral punctation, and the aedeagal forms are also highly variable (see Figure 37 in Girón and Short 2021).[1] A detailed diagnosis can be found in Girón and Short 2021. [1]

Species edit

  1. Helochares aeacus Balfour-Browne, 1952 [4]
  2. Helochares aethiopicus d'Orchymont, 1939 [5]
  3. Helochares alberti d'Orchymont, 1943 [6]
  4. Helochares alcimus d'Orchymont, 1943 [7]
  5. Helochares alcinous Balfour-Browne, 1948 [8]
  6. Helochares altus d'Orchymont, 1943 [9]
  7. Helochares anchoralis Sharp, 1890 [10]
  8. Helochares ancoroides Hebauer, 2001 [11]
  9. Helochares anchoralis expansus Knisch, 1921
  10. Helochares andreinii d'Orchymont, 1939 [12]
  11. Helochares androgynus Hebauer, 1996 [13]
  12. Helochares anthonyae Watts, 1995 [14]
  13. Helochares balfourbrownei Hansen, 1999 [15]
  14. Helochares basilewskyi Balfour-Browne, 1957 [16]
  15. Helochares bilardoi Hebauer, 2009 [17]
  16. Helochares blaesus d'Orchymont, 1936
  17. Helochares bohemani d'Orchymont, 1936
  18. Helochares camerunensis d'Orchymont, 1939
  19. Helochares cancellatus Hebauer, 1998 [18]
  20. Helochares championi Sharp, 1882 [19]
  21. Helochares chappuisi Balfour-Browne, 1952 [20]
  22. Helochares clypeatus (Blackburn, 1891)
  23. Helochares collarti d'Orchymont, 1939
  24. Helochares compactus Hebauer, 2001 [11]
  25. Helochares conformis Hebauer, 1995 [21]
  26. Helochares congoensis d'Orchymont, 1939
  27. Helochares congruens d'Orchymont, 1939
  28. Helochares conjectus d'Orchymont, 1939
  29. Helochares crenatostriatus Régimbart, 1903
  30. Helochares crenatuloides d'Orchymont, 1943
  31. Helochares crenatus Régimbart, 1903
  32. Helochares crepitus Balfour-Browne, 1950 [22]
  33. Helochares cresphontes d'Orchymont, 1939
  34. Helochares crespulus d'Orchymont, 1939
  35. Helochares crispus d'Orchymont, 1939
  36. Helochares dalhuntyi Watts, 1995 [14]
  37. Helochares densepunctus Régimbart, 1907
  38. Helochares densus Sharp, 1890 [10]
  39. Helochares dentalus d'Orchymont, 1943
  40. Helochares denudatus d'Orchymont, 1943
  41. Helochares depactus d'Orchymont, 1939
  42. Helochares diductus d'Orchymont, 1939
  43. Helochares didymoides Balfour-Browne, 1947
  44. Helochares didymus d'Orchymont, 1939
  45. Helochares difficilis d'Orchymont, 1939
  46. Helochares dilutus (Erichson, 1843)
  47. Helochares dilutus consputus Boheman, 1851 [23]
  48. Helochares dimorphus d'Orchymont, 1939
  49. Helochares distinctus Jia & Tang, 2021 [24]
  50. Helochares dollmani Balfour-Browne, 1950
  51. Helochares dolus d'Orchymont, 1939
  52. Helochares egregius Balfour-Browne, 1952
  53. Helochares endroedyi Hebauer, 1996 [13]
  54. Helochares fratris Hebauer, 2003 [25]
  55. Helochares fulgurans Hebauer, 1995 [26]
  56. Helochares fuliginosus d'Orchymont, 1932 [27]
  57. Helochares goticus Hebauer, 1996
  58. Helochares guoi Yang & Jia, 2021 [24]
  59. Helochares hainanensis Dong and Bian, 2021 [28]
  60. Helochares hiekei Hebauer, 1995 [26]
  61. Helochares insolitus d'Orchymont, 1925
  62. Helochares interjectus Hebauer, 1998 [18]
  63. Helochares iteratus Hebauer, 1996 [13]
  64. Helochares itylus Balfour-Browne, 1952
  65. Helochares ivani Hebauer, 1996
  66. Helochares kerstinneumanni Hebauer, 2009 [17]
  67. Helochares knischi d'Orchymont, 1939
  68. Helochares laevis Short and Girón, 2018 [29]
  69. Helochares lamprus d'Orchymont, 1940
  70. Helochares lentus Sharp, 1890 [10]
  71. Helochares lepidus d'Orchymont, 1943
  72. Helochares leptinus d'Orchymont, 1943
  73. Helochares letus d'Orchymont, 1943
  74. Helochares livianus d'Orchymont, 1939
  75. Helochares lividoides Hansen and Hebauer, 1988 [30]
  76. Helochares lividus (Forster, 1771)
  77. Helochares lobatus d'Orchymont, 1948
  78. Helochares lollius d'Orchymont, 1939
  79. Helochares loticus Hebauer, 1998 [18]
  80. Helochares loweryae Watts, 1995 [14]
  81. Helochares luridus (MacLeay, 1871)
  82. Helochares lutulentus Balfour-Browne, 1952
  83. Helochares maculatus Hebauer, 1988 [31]
  84. Helochares maculicollis Mulsant, 1844 [2]
  85. Helochares madli Hebauer, 2002 [32]
  86. Helochares marreensis Watts, 1995 [14]
  87. Helochares mecarus d'Orchymont, 1939
  88. Helochares mediastinus d'Orchymont, 1939
  89. Helochares melanophthalmus (Mulsant, 1844) [2]
  90. Helochares mendosus Hebauer, 1996 [13]
  91. Helochares mentinotus Kuwert, 1888
  92. Helochares menulus d'Orchymont, 1943
  93. Helochares meracus Balfour-Browne, 1950
  94. Helochares mersus d'Orchymont, 1939
  95. Helochares minax d'Orchymont, 1939
  96. Helochares minor d'Orchymont, 1925
  97. Helochares minusculus d'Orchymont, 1943
  98. Helochares namcatensis Hebauer, 2002 [32]
  99. Helochares nebridius d'Orchymont, 1940
  100. Helochares negatus Hebauer, 1995
  101. Helochares neglectus (Hope, 1845)
  102. Helochares nexus Short and Girón, 2018 [29]
  103. Helochares nigrifrons Brancsik, 1893
  104. Helochares nigripalpis Hebauer and Hendrich, 1999 [33]
  105. Helochares nigritulus Kuwert, 1889
  106. Helochares nigroseriatus Hebauer, 1998 [18]
  107. Helochares niobelus d'Orchymont, 1939
  108. Helochares nipponicus Hebauer, 1995 [26]
  109. Helochares normatus (LeConte, 1861)
  110. Helochares notaticollis Régimbart, 1906
  111. Helochares notaticollis curtus Régimbart, 1906
  112. Helochares obliquus Mart, İncekara and Karaca, 2010 [34]
  113. Helochares obscurus (Müller, 1776)
  114. Helochares opacus Hebauer, 2009 [17]
  115. Helochares pallens (MacLeay, 1825)
  116. Helochares parallelus Hebauer, 1999 [35]
  117. Helochares percyi Watts, 1995 [14]
  118. Helochares perminutus Hebauer, 1996 [13]
  119. Helochares phallicus d'Orchymont, 1936
  120. Helochares politus Short and Girón, 2018 [29]
  121. Helochares punctatus Sharp, 1869 [36]
  122. Helochares rugipennis Balfour-Browne, 1958
  123. Helochares salvazai d'Orchymont, 1919
  124. Helochares sauteri d'Orchymont, 1943
  125. Helochares schoedli Hebauer, 1996 [13]
  126. Helochares schwendingeri Hebauer, 1995 [26]
  127. Helochares scitulus Balfour-Browne, 1952
  128. Helochares sechellensis Régimbart, 1903
  129. Helochares serpentinus Hebauer, 1998 [18]
  130. Helochares sharpi (Kuwert, 1890) [37]
  131. Helochares silvester Hebauer, 2009 [17]
  132. Helochares simulator Knisch, 1922
  133. Helochares skalei Hebauer, 2002 [32]
  134. Helochares songi Jia and Tang, 2018
  135. Helochares steffani Hebauer, 2002 [32]
  136. Helochares stenius d'Orchymont, 1943
  137. Helochares striatus (Boheman, 1851) [23]
  138. Helochares strictus d'Orchymont, 1939
  139. Helochares strigellus Hebauer, 2002 [32]
  140. Helochares structus d'Orchymont, 1936
  141. Helochares sublineatus Hebauer, 2002 [32]
  142. Helochares subseriatus Hebauer, 2009 [17]
  143. Helochares subtilis d'Orchymont, 1936
  144. Helochares sufflavus Balfour-Browne, 1952
  145. Helochares sylvaticus Balfour-Browne, 1957
  146. Helochares tamsi Balfour-Browne, 1947
  147. Helochares tatei (Blackburn, 1896)
  148. Helochares tengchongensis Dong and Bian, 2021 [28]
  149. Helochares tenuistriatus Régimbart, 1908
  150. Helochares tertius Hebauer, 1996 [13]
  151. Helochares thurmerae Watts, 1995 [14]
  152. Helochares tristis (MacLeay, 1871)
  153. Helochares trujillo Short and Girón, 2018 [29]
  154. Helochares uenoi Matsui, 1995 [38]
  155. Helochares uhligi Hebauer, 1999 [35]
  156. Helochares vitalisi d'Orchymont, 1919
  157. Helochares wagneri Hebauer, 2002 [32]
  158. Helochares wattsi Hebauer and Hendrich, 1999 [33]
  159. Helochares wuzhifengensis Dong and Bian, 2021 [28]
  160. Helochares yangae Hebauer, Hendrich, and Balke, 1999 [39]
  161. Helochares zamora Short and Girón, 2018 [29]


References edit

 
Wikimedia Commons has media related to Helochares.
  1. ^ a b c d e Girón, Jennifer C.; Short, Andrew Edward Z. (2021-06-18). "The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species". ZooKeys (1045): 1–236. doi:10.3897/zookeys.1045.63810. ISSN 1313-2970. PMC 8233300. PMID 34228772.
  2. ^ a b c Mulsant, E. (1844). Histoire naturelle des coléoptères de France: Palpicornes. Vol. 5. Maison.
  3. ^ a b c Short, Andrew Edward Z.; Girón, Jennifer C.; Toussaint, Emmanuel F. A. (2021). "Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America". Systematic Entomology. 46 (2): 380–395. doi:10.1111/syen.12467. ISSN 1365-3113. S2CID 232264478.
  4. ^ Balfour-Browne, J. (1952). "Contribution à l'étude du peuplement de la Mauritanie. Coléoptères Hydrophilides". Bulletin de l'Institut français d'Afrique noire. 14 (2): 513–517.
  5. ^ d’Orchymont, A. (1939). "L'Helochares minutissimus Kuwert, vrai". Bulletin et Annales de la Société Entomologique de Belgique. 79: 196–198.
  6. ^ d’Orchymont, A. (1943). "Nouvelles notes sur les Helochares (Hydrobaticus) (Coleoptera Palpi-cornia Hydrophilidae)". Bulletin du Musée royal d'Histoire naturelle de Belgique. 19 (20): 1–12.
  7. ^ d’Orchymont, A. (1943). "Nouvelles notes sur les Helochares (Hydrobaticus). (Coleoptera Palpicornia Hydrophilidae)". Bulletin du Musée royal d'Histoire naturelle de Belgique. 19 (20): 1–12.
  8. ^ Balfour-Browne, J. (1947-12-01). "LXXV.—New East African Palpicornia". Annals and Magazine of Natural History. 14 (120): 817–833. doi:10.1080/00222934708654695. ISSN 0374-5481.
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  10. ^ a b c Sharp, David (1890). "IX. On some aquatic Coleoptera from Ceylon". Transactions of the Royal Entomological Society of London. 38 (2): 339–359. doi:10.1111/j.1365-2311.1890.tb03026.x. ISSN 1365-2311.
  11. ^ a b Hebauer, F. (2001). "Beitrag zur Kenntnis der Hydrophilidae von Neuguinea. Ergebnisse der zoologischen Forschungsreisen von M. Balke und L. Hendrich nach West Neuguinea (Irian Jaya) in den Jahren 1990–1998 (Coleoptera: Hydrophilidae)". Acta Coleopterologica. 17 (1): 3–72.
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  13. ^ a b c d e f g Hebauer, F. (1996). "Synopsis der afrikanischen Arten der Gattung Helochares Mulsant (Coleoptera, Hydrophilidae)". Acta Coleopterologica. 12: 3–38.
  14. ^ a b c d e f Watts, C.H.S. (1995). "Revision of the Australasian genera Agraphydrus Rogimbart, Chasmogenus Sharp and Helochares Mulsant (Coleoptera: Hydrophilidae)". Records of the South Australian Museum. 28 (1): 113–130.
  15. ^ Hansen, M. (1999). World catalogue of insects (Volume 2). Hydrophiloidea (s. str.) (Coleoptera). Stenstrup: Apollo Books. pp. 414 pp.
  16. ^ Balfour-Browne, J (1957). "Contributions à l'étude de la faune entomologique du Ruanda-Urun-di (Mission P. Basilewsky 1953). CXVIII. Coleoptera Hydrophilidae". Annales du Musée Royal du Congo Belge, Zool. 58: 14–25.
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  18. ^ a b c d e Hebauer, F. (1998). "Six new species of the genus Helochares Mulsant, 1844, subgenus Hydrobaticus Mac Leay, 1871 from Africa and Asia (Coleoptera: Hydrophilidae)". Acta Coleopterologica. 14: 41–46.
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  21. ^ Hebauer, Franz (1995). "Die Hydrophilidae und Spercheidae Namibias Unter Berücksichtigung des Materials der Namibia-Expedition des Museums für Naturkunde Berlin 1992, ergänzt durch Nachweise aus früheren Namibia- und Kalahari-Expeditionen (Coleoptera, Hydrophiloidea : Hydrophilidae et Spercheidae)". Mitteilungen aus dem Museum für Naturkunde in Berlin. Zoologisches Museum und Institut für Spezielle Zoologie (Berlin) (in German). 71 (2): 251–275. doi:10.1002/mmnz.19950710206. ISSN 1860-0743.
  22. ^ Balfour-Browne, J. (1950). "On the aquatic Coleoptera of Northern Rhodesia (Dytiscidae, Gyr-inidae and Palpicornia)". Occasional Papers of the National Museums of Southern Rhodesia. 2 (16): 359–399.
  23. ^ a b Boheman, C. H. (1851). Insecta Caffrariæ annis 1838–1845 a J. A. Wahlberg col-lecta (Vol. 1). Coleoptera. Holmiæ: Fritze & Norstedt. pp. 625 pp.
  24. ^ a b Yang, Zhenming; Jia, Fenglong; Tang, Yudan; Jiang, Lu (2021-12-16). "Two new species of Helochares, with additional faunistic records from China (Coleoptera, Hydrophilidae, Acidocerinae)". ZooKeys (1078): 57–83. doi:10.3897/zookeys.1078.73458. ISSN 1313-2970. PMC 8702532. PMID 35068953.
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  26. ^ a b c d Hebauer, F. (1995). "Neues zu den Acidocerina Hansen (Helocharae d'Orchymont) der indomalaiischen Region (Coleoptera, Hydrophilidae)". Acta Coleopterologica. 11 (3): 3–14.
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  29. ^ a b c d e Short, A. E. Z.; Girón, J. C. (2018-04-09). "Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)". Zootaxa. 4407 (1): 29–50. doi:10.11646/zootaxa.4407.1.2. ISSN 1175-5334. PMID 29690204.
  30. ^ Hansen, Michael; Hebauer, Franz (1988-01-01). "A new species of Helochares from Israel, with a key to the European and some Near East species (Coleoptera: Hydrophilidae)". Insect Systematics & Evolution. 19 (1): 27–30. doi:10.1163/187631289X00023. ISSN 1876-312X.
  31. ^ Hebauer, F. (1988). "Hydrophiloidea aus Namibia (Coleoptera; Hydrophilidae, Spercheidae)". Bonner zoologische Beiträge. 39: 153–161.
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  33. ^ a b Hebauer, Franz; Hendrich, Lars (1999). "Two new species of Helochares from Northern Australia (Coleoptera: Hydrophilidae)". Entomological Problems. 30 (1): 47–51.
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  38. ^ Matsui, E. (1995). "A new species of the genus Helochares (Coleoptera, Hydrophilidae) from Japan, with a key to the Japanese species of the subgenus Hydrobaticus". Special Bulletin of the Japanese Society of Coleopterology. 4: 317–322.
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April 11, 2024
helochares, genus, water, scavenger, beetles, family, hydrophilidae, represented, described, species, distributed, across, afrotropical, australasian, indo, malayan, nearctic, palearctic, realms, lividusscientific, classificationkingdom, animaliaphylum, arthro. Helochares is a genus of water scavenger beetles in the family Hydrophilidae represented by 161 described species It is distributed across the Afrotropical Australasian Indo Malayan Nearctic and Palearctic realms 1 HelocharesHelochares lividusScientific classificationKingdom AnimaliaPhylum ArthropodaClass InsectaOrder ColeopteraFamily HydrophilidaeSubfamily AcidocerinaeGenus HelocharesMulsant 1844Diversity158 species 3 subspeciesSynonymsHelophilus Mulsant 1844 Pylophilus Motschulsky 1845 Hydrobaticus MacLeay 1871 Helocharimorphus Kuwert 1890 Graphelochares Kuwert 1890 Grapidelochares Ganglbauer 1904 Contents 1 Taxonomy 2 Description 3 Species 4 ReferencesTaxonomy editHelochares is one of the largest and most taxonomically problematic genera within the Acidocerinae It was originally described by Etienne Mulsant in 1844 2 For a long time Helochares contained five subgenera Batochares currently recognized as a separate genus 3 Helochares Helocharimorphus now synonymized under Helochares 1 Hydrobaticus now synonymized under Helochares 1 and Sindolus currently regognized as a separate genus 3 Several taxa that used to be recognized as typical Helochares are now assigned to newly created genera e g Colossochares Novochares or re assigned to exinsting genera i e Peltochares based on a phylogenetic analysis based on molecular data 3 Description editAs currently defined Helochares contains small to medium sized beetles 2 7 mm which are yellowish to brown in coloration There is a lot of variation in the impression of the elytral punctation and the aedeagal forms are also highly variable see Figure 37 in Giron and Short 2021 1 A detailed diagnosis can be found in Giron and Short 2021 1 Species editHelochares aeacus Balfour Browne 1952 4 Helochares aethiopicus d Orchymont 1939 5 Helochares alberti d Orchymont 1943 6 Helochares alcimus d Orchymont 1943 7 Helochares alcinous Balfour Browne 1948 8 Helochares altus d Orchymont 1943 9 Helochares anchoralis Sharp 1890 10 Helochares ancoroides Hebauer 2001 11 Helochares anchoralis expansus Knisch 1921 Helochares andreinii d Orchymont 1939 12 Helochares androgynus Hebauer 1996 13 Helochares anthonyae Watts 1995 14 Helochares balfourbrownei Hansen 1999 15 Helochares basilewskyi Balfour Browne 1957 16 Helochares bilardoi Hebauer 2009 17 Helochares blaesus d Orchymont 1936 Helochares bohemani d Orchymont 1936 Helochares camerunensis d Orchymont 1939 Helochares cancellatus Hebauer 1998 18 Helochares championi Sharp 1882 19 Helochares chappuisi Balfour Browne 1952 20 Helochares clypeatus Blackburn 1891 Helochares collarti d Orchymont 1939 Helochares compactus Hebauer 2001 11 Helochares conformis Hebauer 1995 21 Helochares congoensis d Orchymont 1939 Helochares congruens d Orchymont 1939 Helochares conjectus d Orchymont 1939 Helochares crenatostriatus Regimbart 1903 Helochares crenatuloides d Orchymont 1943 Helochares crenatus Regimbart 1903 Helochares crepitus Balfour Browne 1950 22 Helochares cresphontes d Orchymont 1939 Helochares crespulus d Orchymont 1939 Helochares crispus d Orchymont 1939 Helochares dalhuntyi Watts 1995 14 Helochares densepunctus Regimbart 1907 Helochares densus Sharp 1890 10 Helochares dentalus d Orchymont 1943 Helochares denudatus d Orchymont 1943 Helochares depactus d Orchymont 1939 Helochares diductus d Orchymont 1939 Helochares didymoides Balfour Browne 1947 Helochares didymus d Orchymont 1939 Helochares difficilis d Orchymont 1939 Helochares dilutus Erichson 1843 Helochares dilutus consputus Boheman 1851 23 Helochares dimorphus d Orchymont 1939 Helochares distinctus Jia amp Tang 2021 24 Helochares dollmani Balfour Browne 1950 Helochares dolus d Orchymont 1939 Helochares egregius Balfour Browne 1952 Helochares endroedyi Hebauer 1996 13 Helochares fratris Hebauer 2003 25 Helochares fulgurans Hebauer 1995 26 Helochares fuliginosus d Orchymont 1932 27 Helochares goticus Hebauer 1996 Helochares guoi Yang amp Jia 2021 24 Helochares hainanensis Dong and Bian 2021 28 Helochares hiekei Hebauer 1995 26 Helochares insolitus d Orchymont 1925 Helochares interjectus Hebauer 1998 18 Helochares iteratus Hebauer 1996 13 Helochares itylus Balfour Browne 1952 Helochares ivani Hebauer 1996 Helochares kerstinneumanni Hebauer 2009 17 Helochares knischi d Orchymont 1939 Helochares laevis Short and Giron 2018 29 Helochares lamprus d Orchymont 1940 Helochares lentus Sharp 1890 10 Helochares lepidus d Orchymont 1943 Helochares leptinus d Orchymont 1943 Helochares letus d Orchymont 1943 Helochares livianus d Orchymont 1939 Helochares lividoides Hansen and Hebauer 1988 30 Helochares lividus Forster 1771 Helochares lobatus d Orchymont 1948 Helochares lollius d Orchymont 1939 Helochares loticus Hebauer 1998 18 Helochares loweryae Watts 1995 14 Helochares luridus MacLeay 1871 Helochares lutulentus Balfour Browne 1952 Helochares maculatus Hebauer 1988 31 Helochares maculicollis Mulsant 1844 2 Helochares madli Hebauer 2002 32 Helochares marreensis Watts 1995 14 Helochares mecarus d Orchymont 1939 Helochares mediastinus d Orchymont 1939 Helochares melanophthalmus Mulsant 1844 2 Helochares mendosus Hebauer 1996 13 Helochares mentinotus Kuwert 1888 Helochares menulus d Orchymont 1943 Helochares meracus Balfour Browne 1950 Helochares mersus d Orchymont 1939 Helochares minax d Orchymont 1939 Helochares minor d Orchymont 1925 Helochares minusculus d Orchymont 1943 Helochares namcatensis Hebauer 2002 32 Helochares nebridius d Orchymont 1940 Helochares negatus Hebauer 1995 Helochares neglectus Hope 1845 Helochares nexus Short and Giron 2018 29 Helochares nigrifrons Brancsik 1893 Helochares nigripalpis Hebauer and Hendrich 1999 33 Helochares nigritulus Kuwert 1889 Helochares nigroseriatus Hebauer 1998 18 Helochares niobelus d Orchymont 1939 Helochares nipponicus Hebauer 1995 26 Helochares normatus LeConte 1861 Helochares notaticollis Regimbart 1906 Helochares notaticollis curtus Regimbart 1906 Helochares obliquus Mart Incekara and Karaca 2010 34 Helochares obscurus Muller 1776 Helochares opacus Hebauer 2009 17 Helochares pallens MacLeay 1825 Helochares parallelus Hebauer 1999 35 Helochares percyi Watts 1995 14 Helochares perminutus Hebauer 1996 13 Helochares phallicus d Orchymont 1936 Helochares politus Short and Giron 2018 29 Helochares punctatus Sharp 1869 36 Helochares rugipennis Balfour Browne 1958 Helochares salvazai d Orchymont 1919 Helochares sauteri d Orchymont 1943 Helochares schoedli Hebauer 1996 13 Helochares schwendingeri Hebauer 1995 26 Helochares scitulus Balfour Browne 1952 Helochares sechellensis Regimbart 1903 Helochares serpentinus Hebauer 1998 18 Helochares sharpi Kuwert 1890 37 Helochares silvester Hebauer 2009 17 Helochares simulator Knisch 1922 Helochares skalei Hebauer 2002 32 Helochares songi Jia and Tang 2018 Helochares steffani Hebauer 2002 32 Helochares stenius d Orchymont 1943 Helochares striatus Boheman 1851 23 Helochares strictus d Orchymont 1939 Helochares strigellus Hebauer 2002 32 Helochares structus d Orchymont 1936 Helochares sublineatus Hebauer 2002 32 Helochares subseriatus Hebauer 2009 17 Helochares subtilis d Orchymont 1936 Helochares sufflavus Balfour Browne 1952 Helochares sylvaticus Balfour Browne 1957 Helochares tamsi Balfour Browne 1947 Helochares tatei Blackburn 1896 Helochares tengchongensis Dong and Bian 2021 28 Helochares tenuistriatus Regimbart 1908 Helochares tertius Hebauer 1996 13 Helochares thurmerae Watts 1995 14 Helochares tristis MacLeay 1871 Helochares trujillo Short and Giron 2018 29 Helochares uenoi Matsui 1995 38 Helochares uhligi Hebauer 1999 35 Helochares vitalisi d Orchymont 1919 Helochares wagneri Hebauer 2002 32 Helochares wattsi Hebauer and Hendrich 1999 33 Helochares wuzhifengensis Dong and Bian 2021 28 Helochares yangae Hebauer Hendrich and Balke 1999 39 Helochares zamora Short and Giron 2018 29 References edit nbsp Wikimedia Commons has media related to Helochares a b c d e Giron Jennifer C Short Andrew Edward Z 2021 06 18 The Acidocerinae Coleoptera Hydrophilidae taxonomy classification and catalog of species ZooKeys 1045 1 236 doi 10 3897 zookeys 1045 63810 ISSN 1313 2970 PMC 8233300 PMID 34228772 a b c Mulsant E 1844 Histoire naturelle des coleopteres de France Palpicornes Vol 5 Maison a b c Short Andrew Edward Z Giron Jennifer C Toussaint Emmanuel F A 2021 Evolution and biogeography of acidocerine water scavenger beetles Coleoptera Hydrophilidae shaped by Gondwanan vicariance and Cenozoic isolation of South America Systematic Entomology 46 2 380 395 doi 10 1111 syen 12467 ISSN 1365 3113 S2CID 232264478 Balfour Browne J 1952 Contribution a l etude du peuplement de la Mauritanie Coleopteres Hydrophilides Bulletin de l Institut francais d Afrique noire 14 2 513 517 d Orchymont A 1939 L Helochares minutissimus Kuwert vrai Bulletin et Annales de la Societe Entomologique de Belgique 79 196 198 d Orchymont A 1943 Nouvelles notes sur les Helochares Hydrobaticus Coleoptera Palpi cornia Hydrophilidae Bulletin du Musee royal d Histoire naturelle de Belgique 19 20 1 12 d Orchymont A 1943 Nouvelles notes sur les Helochares Hydrobaticus Coleoptera Palpicornia Hydrophilidae Bulletin du Musee royal d Histoire naturelle de Belgique 19 20 1 12 Balfour Browne J 1947 12 01 LXXV New East African Palpicornia Annals and Magazine of Natural History 14 120 817 833 doi 10 1080 00222934708654695 ISSN 0374 5481 d Orchymont A 1943 Palpicornia Coleoptera VI Bulletin du Musee royal d histoire na turelle de Belgique 19 60 1 12 a b c Sharp David 1890 IX On some aquatic Coleoptera from Ceylon Transactions of the Royal Entomological Society of London 38 2 339 359 doi 10 1111 j 1365 2311 1890 tb03026 x ISSN 1365 2311 a b Hebauer F 2001 Beitrag zur Kenntnis der Hydrophilidae von Neuguinea Ergebnisse der zoologischen Forschungsreisen von M Balke und L Hendrich nach West Neuguinea Irian Jaya in den Jahren 1990 1998 Coleoptera Hydrophilidae Acta Coleopterologica 17 1 3 72 d Orchymont A 1939 Helochares Hydrobaticus Andreinii n sp H melanophthalmus Regimbart 1905 nec Mulsant non Reiche Redia 25 319 323 a b c d e f g Hebauer F 1996 Synopsis der afrikanischen Arten der Gattung Helochares Mulsant Coleoptera Hydrophilidae Acta Coleopterologica 12 3 38 a b c d e f Watts C H S 1995 Revision of the Australasian genera Agraphydrus Rogimbart Chasmogenus Sharp and Helochares Mulsant Coleoptera Hydrophilidae Records of the South Australian Museum 28 1 113 130 Hansen M 1999 World catalogue of insects Volume 2 Hydrophiloidea s str Coleoptera Stenstrup Apollo Books pp 414 pp Balfour Browne J 1957 Contributions a l etude de la faune entomologique du Ruanda Urun di Mission P Basilewsky 1953 CXVIII Coleoptera Hydrophilidae Annales du Musee Royal du Congo Belge Zool 58 14 25 a b c d e Hebauer F 2009 Five new species of the genus Helochares MacLeay 1871 from Gabon and the Congo Coleoptera Hydrophilidae Acta Coleopterologica 25 3 8 a b c d e Hebauer F 1998 Six new species of the genus Helochares Mulsant 1844 subgenus Hydrobaticus Mac Leay 1871 from Africa and Asia Coleoptera Hydrophilidae Acta Coleopterologica 14 41 46 Sharp D 1882 Fam Hydrophilidae In Biologia Centrali Americana Insecta Coleoptera Volume 1 Part 2 Vol 1 pp 53 80 Balfour Browne J 1952 Mission A Villiers au Togo et au Dahomey 1950 VII Coleopteres Hydrophilides Bulletin de l Institut francais d Afrique noire 14 126 139 Hebauer Franz 1995 Die Hydrophilidae und Spercheidae Namibias Unter Berucksichtigung des Materials der Namibia Expedition des Museums fur Naturkunde Berlin 1992 erganzt durch Nachweise aus fruheren Namibia und Kalahari Expeditionen Coleoptera Hydrophiloidea Hydrophilidae et Spercheidae Mitteilungen aus dem Museum fur Naturkunde in Berlin Zoologisches Museum und Institut fur Spezielle Zoologie Berlin in German 71 2 251 275 doi 10 1002 mmnz 19950710206 ISSN 1860 0743 Balfour Browne J 1950 On the aquatic Coleoptera of Northern Rhodesia Dytiscidae Gyr inidae and Palpicornia Occasional Papers of the National Museums of Southern Rhodesia 2 16 359 399 a b Boheman C H 1851 Insecta Caffrariae annis 1838 1845 a J A Wahlberg col lecta Vol 1 Coleoptera Holmiae Fritze amp Norstedt pp 625 pp a b Yang Zhenming Jia Fenglong Tang Yudan Jiang Lu 2021 12 16 Two new species of Helochares with additional faunistic records from China Coleoptera Hydrophilidae Acidocerinae ZooKeys 1078 57 83 doi 10 3897 zookeys 1078 73458 ISSN 1313 2970 PMC 8702532 PMID 35068953 Hebauer F 2003 Checklist of the Malagasy Helochares with description of a new species Coleoptera Hydrophilidae Acta Coleopterologica 19 1 67 69 a b c d Hebauer F 1995 Neues zu den Acidocerina Hansen Helocharae d Orchymont der indomalaiischen Region Coleoptera Hydrophilidae Acta Coleopterologica 11 3 3 14 Hendrich Lars Yang C M 1999 A contribution to the knowledge of the water beetle fauna of Palau Tioman peninsular Malaysia Coleoptera Noteridae Dytiscidae Hydrophilidae Hydraenidae Scirtidae Limnichidae PDF The Raffles Bulletin of Zoology 6 253 262 Archived from the original PDF on July 25 2015 a b c Dong Xue Bian Dongju 2021 03 29 Three new species and two new records of Helochares Hydrobaticus MacLeay 1871 from China Coleoptera Hydrophilidae Acidocerinae Zootaxa 4950 1 166 180 doi 10 11646 zootaxa 4950 1 9 ISSN 1175 5334 PMID 33903325 S2CID 233410003 a b c d e Short A E Z Giron J C 2018 04 09 Review of the Helochares Hydrobaticus MacLeay of the New World Coleoptera Hydrophilidae Acidocerinae Zootaxa 4407 1 29 50 doi 10 11646 zootaxa 4407 1 2 ISSN 1175 5334 PMID 29690204 Hansen Michael Hebauer Franz 1988 01 01 A new species of Helochares from Israel with a key to the European and some Near East species Coleoptera Hydrophilidae Insect Systematics amp Evolution 19 1 27 30 doi 10 1163 187631289X00023 ISSN 1876 312X Hebauer F 1988 Hydrophiloidea aus Namibia Coleoptera Hydrophilidae Spercheidae Bonner zoologische Beitrage 39 153 161 a b c d e f g Hebauer F 2002 New Hydrophilidae of the Old World Coleoptera Hydrophilidae Acta Coleopterologica 18 3 3 24 a b Hebauer Franz Hendrich Lars 1999 Two new species of Helochares from Northern Australia Coleoptera Hydrophilidae Entomological Problems 30 1 47 51 Mart Abdullah Incekara Umit Karaca Hanifi 2010 A new species and records of Hydrophiliae Coleoptera from Turkey PDF Turkish Journal of Zoology 34 297 303 doi 10 3906 zoo 0905 29 Archived from the original PDF on July 25 2015 a b Hebauer F 1999 Neue und wenig bekannte Hydrophiloidea aus dem sudlichen Afrika Co leoptera Hydrophiloidea Acta Coleopterologica 15 2 7 16 Sharp D 1869 Description of a new species of Philhydrus Entomologist s Monthly Magazine 5 240 241 Kuwert A 1890 Bestimmungs Tabellen der europaischen Coleopteren XIX Heft Hydroph ilidae I Abteilung Hydrophilini Verhandlungen des naturforschenden Vereins in Brunn 28 1889 1 121 Matsui E 1995 A new species of the genus Helochares Coleoptera Hydrophilidae from Japan with a key to the Japanese species of the subgenus Hydrobaticus Special Bulletin of the Japanese Society of Coleopterology 4 317 322 Hebauer Franz Hendrich Lars Balke Michael 1999 A contribution to the knowledge of the water beetle fauna Col Hydradephaga Hydrophiloidea and Staphylinoidea of a tropical freshwater lake Tasek Cini Pahang West Malaysia PDF The Raffles Bulletin of Zoology 47 2 333 348 Archived from the original PDF on July 25 2015 Retrieved from https en wikipedia org w index php title Helochares amp oldid 1139173901, wikipedia, wiki, book, books, library,

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