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Hector's beaked whale

Hector's beaked whale (Mesoplodon hectori), is a small mesoplodont living in the Southern Hemisphere. This whale is named after Sir James Hector, a founder of the colonial museum in Wellington, New Zealand. The species has rarely been seen in the wild.

Hector's beaked whale
Size compared to an average human
CITES Appendix II (CITES)[2]
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Infraorder: Cetacea
Family: Ziphiidae
Genus: Mesoplodon
Species:
M. hectori
Binomial name
Mesoplodon hectori
(Gray, 1871)
Hector's beaked whale range

Some data supposedly referring to this species, especially juveniles and males, turned out to be based on the misidentified specimens of Perrin's beaked whale - especially since the adult male of Hector's beaked whale was only more recently described.[3][4][5][6][7][8][9][10][11][12]

Taxonomy edit

The English taxonomist John Edward Gray first named the species Berardius hectori in 1871, based on a specimen (a 2.82 metres (9 ft 3 in) male) collected in Titahi Bay, New Zealand in January, 1866.[13] The following year, 1872, English anatomist William Henry Flower placed it in the genus Mesoplodon, while in 1873, Scottish scientist James Hector assigned the same specimen to the species M. knoxi. The species remained in the genus Mesoplodon until 1962, when Charles McCann, a vertebrate zoologist at the Dominion Museum in Wellington, argued that the species only represented a juvenile of Berardius arnuxi. Beaked whale specialist Joseph Curtis Moore (1968) and J. G. B. Ross (1970) contested this designation, arguing that M. hectori was a valid species. Adult male specimens from the 1970s and 1980s confirmed the species' specific status.[14]

Molecular Taxonomy edit

 
Illustrations of a Hector's beaked whale's skull from Volume 10 of Transactions and Proceedings of the New Zealand Institute.

With abundant and easily observable species, the use of synapomorphic characters to assign observed individuals to a particular species, is a reasonably effective approach to taxonomy, at least in its rudimentary phase.[15] However, the translation of this approach to rare species with elusive life histories, such as the Hector's beaked whales, can be problematic for many reasons. Also, the usefulness of whole or partial "voucher" specimens in private collections to assign taxonomic difference based on morphology is equally susceptible to inaccuracy, and additionally can lead to unethical acquisition methods, such as illegal hunting of a rare species.[16] Therefore, the accurate and ethical approach to the taxonomic study of rare and elusive species, such as are found within the mysterious, deep-diving family Ziphiidae, is a molecular phylogenetic taxonomic methodology.[17] This method has yielded insightful discovery in recent studies of the family Ziphiidae. Between 1975 and 1997, five beaked whales were found stranded in Southern California; based on morphology, four were identified as members of M. Hectori and one was identified as a member of Ziphius cavirostris.[3] This discovery was puzzling, because although the suspected range of Ziphius cavirostris was met by the stranding location, M. Hectori was thought to be restricted solely to the Southern Hemisphere.[3] mtDNA sequencing of the sample species revealed that all five individuals could be genetically distinguished from all other members of Ziphiidae, including the two species they were previously identified as.[18] They were subsequently classified as a new species, Perrin's beaked whale (M. Perrini).[18] More recent phylogenetic analyses of nuclear actin sequences substantiated the validity of this classification, as a distinct 34 base-pair deletion was found in the allele, screened across all species, of the two M. perrini samples in particular.[17]

Description edit

Reaching a maximum length around 4.2 m (1.9 m when born), and with an estimated weight around 1 tonne (1.032 tons), Hector's is one of the smallest of the beaked whales. It is known from only a few stranded animals and a single confirmed sighting of a juvenile off Western Australia.[1] Hector's beaked whales are dark greyish-brown dorsally, paler ventrally. A single female specimen found in Argentina was light grey dorsally and white ventrally.[19] An individual male described in the same study had several scars and teeth marks found diffuse on its back and flanks. Intra-specific male-to-male interactions are possibly the cause for such marks. Additionally, oval white scars on the ventral portion of this male specimen were likely caused from cookie-cutter sharks of the Isistius species. Another single adult male specimen had a white beak and white on the anterior portion of the head, with white, linear scars criss-crossing its body, while the juvenile seen off Western Australia had a mask covering its eyes and extending unto its melon and upper beak. The melon, which is not very prominent, slopes quite steeply to the short beak. The dorsal fin is triangular to slightly hooked, small, and rounded at the tip. The leading edge of the dorsal fin joins the body at a sharp angle.

Stomach edit

Like other members of the Mesoplodont genus, the stomach is divided into four chambers.[8] The proximal main stomach, the distal main stomach, the connecting stomachs, and the pyloric stomach are the four chambers respectively.[8][19] The first, second, and fourth chambers are pink in color and soft to touch. The third chamber is grey in color and hard externally. The first and third chamber have internal folds but the folds are longer and larger in the third chamber. The second chamber is smooth, with no folds. The first chamber connects to the esophagus and the fourth chamber connects to the small intestine. The orange gastric fluid found in the stomach has also been found in some areas of the intestines. The specimen's stomach from Argentina had many small crystalline lenses in the first and fourth stomachs. Although, other food remains were not present. The specimen's stomach also had dozens of nematodes in all the stomach chambers except the connecting chamber.[19] Histo-pathological analysis showed that these nematodes were later found to be of the Anisakis species.[20]

Dentition edit

Adult males have a pair of flattened, triangular teeth near the tip of the lower jaw. As with most other beaked whales, the teeth do not erupt in females.

In March 2016, the South Australian Museum conducted a necropsy on a beached female specimen of the species from Waitpinga beach, near Adelaide, South Australia. The specimen was found to have a pair of large fangs not seen among the species typical dentition, especially for females, who typically have none. The fangs are possibly vestigial, or atavisms of some other kind, though a definitive answer is difficult because of the dearth of knowledge about the species.[21]

Distribution and ecology edit

Hector's beaked whale has a circumpolar distribution in cool temperate Southern Hemisphere waters between about 35 and 55°S. Most records are from New Zealand, but also reports from Falkland Sound, Falkland Islands, Lottering River, South Africa, Adventure Bay, Tasmania, and Tierra del Fuego, in southern South America have been made. Supposed Northeast Pacific records in the older literature actually refer to Perrin's beaked whale.

Sightings are rare due to their deep-ocean distribution, elusive behaviour and possible low numbers. Nothing is known about the diet of this species, although it is assumed to feed on deepwater squid and fish. Because they lack functional teeth, they presumably capture most of their prey by suction.

Body scarring suggests there may be extensive fighting between males, which is common in beaked whales. Nothing is known about breeding in this species.

This species has never been hunted at all, and has not entangled itself in fishing gear. Most records of the whale have been stranded specimens on beaches, particularly in New Zealand.

Conservation edit

Hector's beaked whale is covered by the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands Region (Pacific Cetaceans MOU).[22]

Specimens edit

  1. MNZ MM001834 - 16 July 1980; Kaikoura, New Zealand

See also edit

References edit

  1. ^ a b Pitman, R.L.; Brownell Jr.; R.L. (2020). "Mesoplodon hectori". IUCN Red List of Threatened Species. 2020: e.T13248A50366525. doi:10.2305/IUCN.UK.2020-3.RLTS.T13248A50366525.en. Retrieved 19 November 2021.
  2. ^ "Appendices | CITES". cites.org. Retrieved 2022-01-14.
  3. ^ a b c Mead, James G. (1981). "First records of Mesoplodon hectori (Ziphiidae) from the Northern Hemisphere and a description of the adult male". Journal of Mammalogy. 62 (2): 430–432. doi:10.2307/1380733. JSTOR 1380733.
  4. ^ Mead, James G. (1984). "Survey of reproductive data for the beaked whales (Ziphiidae)" (PDF). International Whaling Commission Special Issue. 6: 91–96.
  5. ^ Mead, James G.; Baker, Alan N. (1987). "Notes on the rare beaked whale, Mesoplodon hectori (Gray)". Royal Society of New Zealand. 17 (3): 303–312. doi:10.1080/03036758.1987.10418163.
  6. ^ Mead, James G. (1989). "Beaked whales of the genus Mesoplodon". In Ridgway, S.H.; Harrison, R. (eds.). Handbook of marine mammals Vol.4. London: Academic Press. pp. 349–430. doi:10.2307/2403599. JSTOR 2403599.
  7. ^ Jefferson, T.A.; Leatherwood, S.; Webber, M.A. (1993). FAO species identification guide: Marine mammals of the world (PDF). United States & Rome: United States Environment Programme & Food and Agriculture Organization of the United Nations (FAO).
  8. ^ a b c Mead, James G. (1993). "The systematic importance of stomach anatomy in beaked whales" (PDF). IBI Reports. 4: 75–86.[permanent dead link]
  9. ^ Carwadine, M. (1995). Whales, dolphins and porpoises. London: HarperCollins. ISBN 0004722736.
  10. ^ Reeves, Randall R.; Leatherwood, S. (1994). Dolphins, porpoises and whales: 1994-98 Action plan for the conservation of cetaceans. Gland, Switzerland: IUCN. ISBN 2-8317-0189-9.
  11. ^ Henshaw, M.D.; Leduc, R.G.; Chivers, S.J.; Dizon, A.E. (1997). "Identification of beaked whales (family Ziphiidae) using mtDNA sequences". Marine Mammal Science. 13 (3): 487–495. doi:10.1111/j.1748-7692.1997.tb00656.x.
  12. ^ Messenger, S.L.; McQuire, J.A. (1998). "Morphology, molecules and the phylogenetics of cetaceans". Systematic Biology. 47 (1): 90–124. doi:10.1080/106351598261058. PMID 12064244.
  13. ^ Gray, J.E. (1871). "Notes on the Berardius of New Zealand". Annals and Magazine of Natural History. 8 (44): 115–117. doi:10.1080/00222937108696444. ISSN 0374-5481.
  14. ^ Mead, James G.; Baker, Alan N. (1987). "Notes on the rare beaked whale,Mesoplodon hectori(Gray)". Journal of the Royal Society of New Zealand. 17 (3): 303–312. doi:10.1080/03036758.1987.10418163. ISSN 0303-6758.
  15. ^ Yeates, David (June 2001). "Species Concepts and Phylogenetic Theory: A Debate. Quentin D. Wheeler , Rudolf Meier". The Quarterly Review of Biology. 76 (2): 234–235. doi:10.1086/393912. ISSN 0033-5770.
  16. ^ Tubbs, Philip K. (1991). "I.u.b.s. Section Of Zoological Nomenclature. Report Of Meeting, Ambsterdam, 6 September 1991". The Bulletin of Zoological Nomenclature. 48: 293–294. doi:10.5962/bhl.part.757. ISSN 0007-5167.
  17. ^ a b Dalebout, M. L.; Baker, C. S.; Mead, J. G.; Cockcroft, V. G.; Yamada, T. K. (2004-11-01). "A Comprehensive and Validated Molecular Taxonomy of Beaked Whales, Family Ziphiidae". Journal of Heredity. 95 (6): 459–473. CiteSeerX 10.1.1.507.2896. doi:10.1093/jhered/esh054. ISSN 1465-7333. PMID 15475391.
  18. ^ a b Dalebout, Merel L.; Mead, James G.; Baker, C. Scott; Baker, Alan N.; Helden, Anton L. (July 2002). "A New Species of Beaked Whale Mesoplodon perrini sp. n. (Cetacea: Ziphiidae) Discovered Through Phylogenetic Analyses of Mitochondrial DNA Sequences". Marine Mammal Science. 18 (3): 577–608. doi:10.1111/j.1748-7692.2002.tb01061.x. ISSN 0824-0469.
  19. ^ a b c Cappozzo, H. L.; Negri, M. F.; Mahler, B.; Lia, V. V.; Martinez, P.; Gianggiobe, A.; Saubidet, A. (2005). "Biological data on two Hector's beaked whales, Mesoplodon hectori , stranded in Buenos Aires province, Argentina". Latin American Journal of Aquatic Mammals. 4 (2): 113–128. doi:10.5597/lajam00076. ISSN 2236-1057.
  20. ^ Nikolov, Pavel N.; Cappozzo, H. Luis; Berón-Vera, Bárbara; Crespo, Enrique A.; Raga, J. Antonio; Fernández, Mercedes (August 2010). "Cestodes from Hector's Beaked Whale (Mesoplodon hectori) and Spectacled Porpoise (Phocoena dioptrica) from Argentinean Waters". Journal of Parasitology. 96 (4): 746–751. doi:10.1645/ge-2200.1. ISSN 0022-3395. PMID 20486735. S2CID 22848752.
  21. ^ Geggel, Laura (May 19, 2016). "Stranded, Rarely Seen Beaked Whale Has Strange Fang". LiveScience. Retrieved 2016-08-13.
  22. ^ Official webpage of the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands Region

Further reading edit

  • Baker, Alan N. (1990): Whales and dolphins of New Zealand and Australia: An identification guide. Victoria University Press, Wellington.
  • Perrin, William F.; Wursig, Bernd & Thewissen, J.G.M (eds.) (2002): Encyclopedia of Marine Mammals. Academic Press. ISBN 0-12-551340-2
  • Reeves, Randall R.; Steward, Brent S.; Clapham, Phillip J. & Owell, James A. (2002): Sea Mammals of the World. A & C Black, London. ISBN 0-7136-6334-0

External links edit

  • Hector's Beaked Whale Fact Sheet
  • Whale & Dolphin Conservation Society (WDCS)

hector, beaked, whale, mesoplodon, hectori, small, mesoplodont, living, southern, hemisphere, this, whale, named, after, james, hector, founder, colonial, museum, wellington, zealand, species, rarely, been, seen, wild, size, compared, average, humanconservatio. Hector s beaked whale Mesoplodon hectori is a small mesoplodont living in the Southern Hemisphere This whale is named after Sir James Hector a founder of the colonial museum in Wellington New Zealand The species has rarely been seen in the wild Hector s beaked whaleSize compared to an average humanConservation statusData Deficient IUCN 3 1 1 CITES Appendix II CITES 2 Scientific classificationDomain EukaryotaKingdom AnimaliaPhylum ChordataClass MammaliaOrder ArtiodactylaInfraorder CetaceaFamily ZiphiidaeGenus MesoplodonSpecies M hectoriBinomial nameMesoplodon hectori Gray 1871 Hector s beaked whale rangeSome data supposedly referring to this species especially juveniles and males turned out to be based on the misidentified specimens of Perrin s beaked whale especially since the adult male of Hector s beaked whale was only more recently described 3 4 5 6 7 8 9 10 11 12 Contents 1 Taxonomy 1 1 Molecular Taxonomy 2 Description 2 1 Stomach 2 2 Dentition 3 Distribution and ecology 4 Conservation 5 Specimens 6 See also 7 References 8 Further reading 9 External linksTaxonomy editThe English taxonomist John Edward Gray first named the species Berardius hectori in 1871 based on a specimen a 2 82 metres 9 ft 3 in male collected in Titahi Bay New Zealand in January 1866 13 The following year 1872 English anatomist William Henry Flower placed it in the genus Mesoplodon while in 1873 Scottish scientist James Hector assigned the same specimen to the species M knoxi The species remained in the genus Mesoplodon until 1962 when Charles McCann a vertebrate zoologist at the Dominion Museum in Wellington argued that the species only represented a juvenile of Berardius arnuxi Beaked whale specialist Joseph Curtis Moore 1968 and J G B Ross 1970 contested this designation arguing that M hectori was a valid species Adult male specimens from the 1970s and 1980s confirmed the species specific status 14 Molecular Taxonomy edit nbsp Illustrations of a Hector s beaked whale s skull from Volume 10 of Transactions and Proceedings of the New Zealand Institute With abundant and easily observable species the use of synapomorphic characters to assign observed individuals to a particular species is a reasonably effective approach to taxonomy at least in its rudimentary phase 15 However the translation of this approach to rare species with elusive life histories such as the Hector s beaked whales can be problematic for many reasons Also the usefulness of whole or partial voucher specimens in private collections to assign taxonomic difference based on morphology is equally susceptible to inaccuracy and additionally can lead to unethical acquisition methods such as illegal hunting of a rare species 16 Therefore the accurate and ethical approach to the taxonomic study of rare and elusive species such as are found within the mysterious deep diving family Ziphiidae is a molecular phylogenetic taxonomic methodology 17 This method has yielded insightful discovery in recent studies of the family Ziphiidae Between 1975 and 1997 five beaked whales were found stranded in Southern California based on morphology four were identified as members of M Hectori and one was identified as a member of Ziphius cavirostris 3 This discovery was puzzling because although the suspected range of Ziphius cavirostris was met by the stranding location M Hectori was thought to be restricted solely to the Southern Hemisphere 3 mtDNA sequencing of the sample species revealed that all five individuals could be genetically distinguished from all other members of Ziphiidae including the two species they were previously identified as 18 They were subsequently classified as a new species Perrin s beaked whale M Perrini 18 More recent phylogenetic analyses of nuclear actin sequences substantiated the validity of this classification as a distinct 34 base pair deletion was found in the allele screened across all species of the two M perrini samples in particular 17 Description editReaching a maximum length around 4 2 m 1 9 m when born and with an estimated weight around 1 tonne 1 032 tons Hector s is one of the smallest of the beaked whales It is known from only a few stranded animals and a single confirmed sighting of a juvenile off Western Australia 1 Hector s beaked whales are dark greyish brown dorsally paler ventrally A single female specimen found in Argentina was light grey dorsally and white ventrally 19 An individual male described in the same study had several scars and teeth marks found diffuse on its back and flanks Intra specific male to male interactions are possibly the cause for such marks Additionally oval white scars on the ventral portion of this male specimen were likely caused from cookie cutter sharks of the Isistius species Another single adult male specimen had a white beak and white on the anterior portion of the head with white linear scars criss crossing its body while the juvenile seen off Western Australia had a mask covering its eyes and extending unto its melon and upper beak The melon which is not very prominent slopes quite steeply to the short beak The dorsal fin is triangular to slightly hooked small and rounded at the tip The leading edge of the dorsal fin joins the body at a sharp angle Stomach edit Like other members of the Mesoplodont genus the stomach is divided into four chambers 8 The proximal main stomach the distal main stomach the connecting stomachs and the pyloric stomach are the four chambers respectively 8 19 The first second and fourth chambers are pink in color and soft to touch The third chamber is grey in color and hard externally The first and third chamber have internal folds but the folds are longer and larger in the third chamber The second chamber is smooth with no folds The first chamber connects to the esophagus and the fourth chamber connects to the small intestine The orange gastric fluid found in the stomach has also been found in some areas of the intestines The specimen s stomach from Argentina had many small crystalline lenses in the first and fourth stomachs Although other food remains were not present The specimen s stomach also had dozens of nematodes in all the stomach chambers except the connecting chamber 19 Histo pathological analysis showed that these nematodes were later found to be of the Anisakis species 20 Dentition edit Adult males have a pair of flattened triangular teeth near the tip of the lower jaw As with most other beaked whales the teeth do not erupt in females In March 2016 the South Australian Museum conducted a necropsy on a beached female specimen of the species from Waitpinga beach near Adelaide South Australia The specimen was found to have a pair of large fangs not seen among the species typical dentition especially for females who typically have none The fangs are possibly vestigial or atavisms of some other kind though a definitive answer is difficult because of the dearth of knowledge about the species 21 Distribution and ecology editThis section does not cite any sources Please help improve this section by adding citations to reliable sources Unsourced material may be challenged and removed April 2021 Learn how and when to remove this template message Hector s beaked whale has a circumpolar distribution in cool temperate Southern Hemisphere waters between about 35 and 55 S Most records are from New Zealand but also reports from Falkland Sound Falkland Islands Lottering River South Africa Adventure Bay Tasmania and Tierra del Fuego in southern South America have been made Supposed Northeast Pacific records in the older literature actually refer to Perrin s beaked whale Sightings are rare due to their deep ocean distribution elusive behaviour and possible low numbers Nothing is known about the diet of this species although it is assumed to feed on deepwater squid and fish Because they lack functional teeth they presumably capture most of their prey by suction Body scarring suggests there may be extensive fighting between males which is common in beaked whales Nothing is known about breeding in this species This species has never been hunted at all and has not entangled itself in fishing gear Most records of the whale have been stranded specimens on beaches particularly in New Zealand Conservation editHector s beaked whale is covered by the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands Region Pacific Cetaceans MOU 22 Specimens editThis list is incomplete you can help by adding missing items February 2012 MNZ MM001834 16 July 1980 Kaikoura New ZealandSee also edit nbsp Cetaceans portal nbsp Mammals portal nbsp Marine life portalList of cetaceansReferences edit a b Pitman R L Brownell Jr R L 2020 Mesoplodon hectori IUCN Red List of Threatened Species 2020 e T13248A50366525 doi 10 2305 IUCN UK 2020 3 RLTS T13248A50366525 en Retrieved 19 November 2021 Appendices CITES cites org Retrieved 2022 01 14 a b c Mead James G 1981 First records of Mesoplodon hectori Ziphiidae from the Northern Hemisphere and a description of the adult male Journal of Mammalogy 62 2 430 432 doi 10 2307 1380733 JSTOR 1380733 Mead James G 1984 Survey of reproductive data for the beaked whales Ziphiidae PDF International Whaling Commission Special Issue 6 91 96 Mead James G Baker Alan N 1987 Notes on the rare beaked whale Mesoplodon hectori Gray Royal Society of New Zealand 17 3 303 312 doi 10 1080 03036758 1987 10418163 Mead James G 1989 Beaked whales of the genus Mesoplodon In Ridgway S H Harrison R eds Handbook of marine mammals Vol 4 London Academic Press pp 349 430 doi 10 2307 2403599 JSTOR 2403599 Jefferson T A Leatherwood S Webber M A 1993 FAO species identification guide Marine mammals of the world PDF United States amp Rome United States Environment Programme amp Food and Agriculture Organization of the United Nations FAO a b c Mead James G 1993 The systematic importance of stomach anatomy in beaked whales PDF IBI Reports 4 75 86 permanent dead link Carwadine M 1995 Whales dolphins and porpoises London HarperCollins ISBN 0004722736 Reeves Randall R Leatherwood S 1994 Dolphins porpoises and whales 1994 98 Action plan for the conservation of cetaceans Gland Switzerland IUCN ISBN 2 8317 0189 9 Henshaw M D Leduc R G Chivers S J Dizon A E 1997 Identification of beaked whales family Ziphiidae using mtDNA sequences Marine Mammal Science 13 3 487 495 doi 10 1111 j 1748 7692 1997 tb00656 x Messenger S L McQuire J A 1998 Morphology molecules and the phylogenetics of cetaceans Systematic Biology 47 1 90 124 doi 10 1080 106351598261058 PMID 12064244 Gray J E 1871 Notes on the Berardius of New Zealand Annals and Magazine of Natural History 8 44 115 117 doi 10 1080 00222937108696444 ISSN 0374 5481 Mead James G Baker Alan N 1987 Notes on the rare beaked whale Mesoplodon hectori Gray Journal of the Royal Society of New Zealand 17 3 303 312 doi 10 1080 03036758 1987 10418163 ISSN 0303 6758 Yeates David June 2001 Species Concepts and Phylogenetic Theory A Debate Quentin D Wheeler Rudolf Meier The Quarterly Review of Biology 76 2 234 235 doi 10 1086 393912 ISSN 0033 5770 Tubbs Philip K 1991 I u b s Section Of Zoological Nomenclature Report Of Meeting Ambsterdam 6 September 1991 The Bulletin of Zoological Nomenclature 48 293 294 doi 10 5962 bhl part 757 ISSN 0007 5167 a b Dalebout M L Baker C S Mead J G Cockcroft V G Yamada T K 2004 11 01 A Comprehensive and Validated Molecular Taxonomy of Beaked Whales Family Ziphiidae Journal of Heredity 95 6 459 473 CiteSeerX 10 1 1 507 2896 doi 10 1093 jhered esh054 ISSN 1465 7333 PMID 15475391 a b Dalebout Merel L Mead James G Baker C Scott Baker Alan N Helden Anton L July 2002 A New Species of Beaked Whale Mesoplodon perrini sp n Cetacea Ziphiidae Discovered Through Phylogenetic Analyses of Mitochondrial DNA Sequences Marine Mammal Science 18 3 577 608 doi 10 1111 j 1748 7692 2002 tb01061 x ISSN 0824 0469 a b c Cappozzo H L Negri M F Mahler B Lia V V Martinez P Gianggiobe A Saubidet A 2005 Biological data on two Hector s beaked whales Mesoplodon hectori stranded in Buenos Aires province Argentina Latin American Journal of Aquatic Mammals 4 2 113 128 doi 10 5597 lajam00076 ISSN 2236 1057 Nikolov Pavel N Cappozzo H Luis Beron Vera Barbara Crespo Enrique A Raga J Antonio Fernandez Mercedes August 2010 Cestodes from Hector s Beaked Whale Mesoplodon hectori and Spectacled Porpoise Phocoena dioptrica from Argentinean Waters Journal of Parasitology 96 4 746 751 doi 10 1645 ge 2200 1 ISSN 0022 3395 PMID 20486735 S2CID 22848752 Geggel Laura May 19 2016 Stranded Rarely Seen Beaked Whale Has Strange Fang LiveScience Retrieved 2016 08 13 Official webpage of the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands RegionFurther reading editBaker Alan N 1990 Whales and dolphins of New Zealand and Australia An identification guide Victoria University Press Wellington Perrin William F Wursig Bernd amp Thewissen J G M eds 2002 Encyclopedia of Marine Mammals Academic Press ISBN 0 12 551340 2 Reeves Randall R Steward Brent S Clapham Phillip J amp Owell James A 2002 Sea Mammals of the World A amp C Black London ISBN 0 7136 6334 0External links editHector s Beaked Whale Fact Sheet Cetaceans of the World CMS Whale amp Dolphin Conservation Society WDCS Retrieved from https en wikipedia org w index php title Hector 27s beaked whale amp oldid 1195830525, wikipedia, wiki, book, books, library,

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