This page lists the genera in the family Carabidae , according to Catalogue of Life and Carabcat Database as of March 2023, excluding Tiger Beetles (which are considered to be in the family Cicindelidae).[1] [2]
Contents 1 Family Carabidae 1.1 Subfamily Anthiinae Bonelli, 1813 1.2 Subfamily Apotominae LeConte, 1853 1.3 Subfamily Brachininae Bonelli, 1810 1.4 Subfamily Broscinae Hope, 1838 1.5 Subfamily Carabinae Linnaeus, 1802 1.6 Subfamily Ctenodactylinae Laporte, 1834 1.7 Subfamily Dryptinae Bonelli, 1810 1.8 Subfamily Elaphrinae Latreille, 1802 1.9 Subfamily Gineminae Ball & Shpeley, 2002 1.10 Subfamily Harpalinae Bonelli, 1810 1.11 Subfamily Hiletinae Schiödte, 1847 1.12 Subfamily Lebiinae Bonelli, 1810 1.13 Subfamily Licininae Bonelli, 1810 1.14 Subfamily Loricerinae Bonelli, 1810 1.15 Subfamily Melaeninae Csiki, 1933 1.16 Subfamily Migadopinae Chaudoir, 1861 1.17 Subfamily Nebriinae Laporte, 1834 1.18 Subfamily Nototylinae Bänninger, 1927 1.19 Subfamily Omophroninae Bonelli, 1810 1.20 Subfamily Orthogoniinae Schaum, 1857 1.21 Subfamily Panagaeinae Bonelli, 1810 1.22 Subfamily Patrobinae Kirby, 1837 1.23 Subfamily Paussinae Latreille, 1806 1.24 Subfamily Platyninae Bonelli, 1810 1.25 Subfamily Promecognathinae LeConte, 1853 1.26 Subfamily Pseudomorphinae Hope, 1838 1.27 Subfamily Psydrinae LeConte, 1853 1.28 Subfamily Pterostichinae Bonelli, 1810 1.29 Subfamily Rhysodinae Laporte, 1840 1.30 Subfamily Scaritinae Bonelli, 1810 1.31 Subfamily Siagoninae Bonelli, 1813 1.32 Subfamily Trechinae Bonelli, 1810 1.33 Subfamily Xenaroswellianinae Erwin, 2007 1.34 Subfamily †Conjunctiinae Ponomarenko, 1977 1.35 Subfamily †Protorabinae Ponomarenko, 1977 2 References Family Carabidae edit Subfamily Anthiinae Bonelli, 1813 edit Tribe Anthiini Bonelli, 1813 Anthia Weber, 1801 Atractonotus Perroud, 1847 Baeoglossa Chaudoir, 1850 Cycloloba Chaudoir, 1850 Cypholoba Chaudoir, 1850 Eccoptoptera Chaudoir, 1878 Gonogenia Chaudoir, 1844 Netrodera Chaudoir, 1850 Tribe Helluonini Hope, 1838 Subtribe Helluonina Hope, 1838 Aenigma Newman, 1836 Ametroglossus Sloane, 1914 Dicranoglossus Chaudoir, 1872 Epimicodema Sloane, 1914 Gigadema J.Thomson, 1859 Helluapterus Sloane, 1914 Helluarchus Sloane, 1914 Helluo Bonelli, 1813 Helluodema Laporte, 1867 Helluonidius Chaudoir, 1872 Helluopapua Darlington, 1968 Helluosoma Laporte, 1867 Neohelluo Sloane, 1914 Platyhelluo Baehr, 2005 Subtribe Omphrina Jedlicka, 1941 Colfax Andrewes, 1920 Creagris Nietner, 1857 Dailodontus Reiche, 1843 Erephognathus Alluaud, 1932 Helluobrochus Reichardt, 1974 Helluomorpha Laporte, 1834 Helluomorphoides Ball, 1951 Macrocheilus Hope, 1838 Meladroma Motschulsky, 1855 Omphra Dejean, 1825 Pleuracanthus Gray, 1832 Triaenogenius Chaudoir, 1877 Tribe Physocrotaphini Chaudoir, 1863 Anguloderus Anichtchenko & Sciaky, 2017 Foveocrotaphus Anichtchenko, 2014 Helluodes Westwood, 1847 Holoponerus Fairmaire, 1883 Physocrotaphus Parry, 1849 Physoglossus Akhil & Sabu, 2020 Pogonoglossus Chaudoir, 1863 Schuelea Baehr, 2004 Subfamily Apotominae LeConte, 1853 edit Apotomus Illiger, 1807 Subfamily Brachininae Bonelli, 1810 edit Tribe Brachinini Bonelli, 1810 Aptinoderus Hubenthal, 1919 Aptinus Bonelli, 1810 Brachinulus Basilewsky, 1958 Brachinus Weber, 1801 Mastax Fischer von Waldheim, 1828 Pheropsophus Solier, 1833 Styphlodromus Basilewsky, 1959 Styphlomerus Chaudoir in Putzeys, 1875 Tribe Crepidogastrini Jeannel, 1949 Brachynillus Reitter, 1904 Crepidogaster Boheman, 1848 Crepidogastrillus Basilewsky, 1959 Crepidogastrinus Basilewsky, 1957 Crepidolomus Basilewsky, 1959 Crepidonellus Basilewsky, 1959 Subfamily Broscinae Hope, 1838 edit Tribe Broscini Hope, 1838 Subtribe Axonyina Roig-Juñent, 2000 Broscodes Bolivar y Pieltain, 1914 Rawlinsius Davidson & Ball, 1998 Subtribe Baripodina Jeannel, 1941 Baripus Dejean, 1828 Bembidiomorphum Champion, 1918 Subtribe Broscina Hope, 1838 Broscodera Lindroth, 1961 Broscosoma Rosenhauer, 1846 Broscus Panzer, 1813 Chaetobroscus Semenov, 1900 Craspedonotus Schaum, 1863 Eobroscus Kryzhanovskij, 1951 Kashmirobroscus J.Schmidt; Wrase & Sciaky, 2013 Miscodera Eschscholtz, 1830 Zacotus LeConte, 1869 Subtribe Creobiina Jeannel, 1941 Acallistus Sharp, 1886 Adotela Laporte, 1867 Anheterus Putzeys, 1868 Bountya Townsend, 1971 Brithysternum W.J.MacLeay, 1873 Cascellius Curtis, 1838 Cerotalis Laporte, 1867 Creobius Guérin-Méneville, 1838 Gnathoxys Westwood, 1842 Nothocascellius Roig-Juñent, 1995 Promecoderus Dejean, 1829 Subtribe Nothobroscina Roig-Juñent, 2000 Chylnus Sloane, 1920 Diglymma Sharp, 1886 Eurylychnus Bates, 1891 Mecodema Blanchard, 1843 Monteremita Seldon & Holwell, 2019 Nothobroscus Roig-Juñent & Ball, 1995 Oregus Putzeys, 1868 Orthoglymma Liebherr; Marris; Emberson; Syrett & Roig-Juñent, 2011 Percolestus Sloane, 1892 Percosoma Schaum, 1858 Subfamily Carabinae Linnaeus, 1802 edit Tribe Carabini Linnaeus, 1802 Subtribe Carabina Linnaeus, 1802 Aplothorax G.R.Waterhouse, 1842 Calosoma Weber, 1801 Carabus Linnaeus, 1758 Subtribe Ceroglossina Lapouge, 1927 Ceroglossus Solier, 1848 Tribe Cychrini Perty, 1830 Subtribe Cychrina Perty, 1830 Cychropsis Boileau, 1901 Cychrus Fabricius, 1794 Scaphinotus Dejean, 1826 Sphaeroderus Dejean, 1826 Subtribe Pamborina Hope, 1838 Maoripamborus Brookes, 1944 Pamborus Latreille, 1812 Tribe Ctenodactylini Laporte, 1834 Alachnothorax Liebke, 1929 Amblycoleus Chaudoir, 1872 Antipionycha Liebke, 1928 Askalaphium Liebke, 1938 Ctenodactyla Dejean, 1825 Leptotrachelon Liebke, 1928 Leptotrachelus Latreille, 1829 Oilea Liebke, 1931 Parapionycha Liebke, 1929 Pionycha Chaudoir, 1848 Plagiotelum Solier, 1849 Propionycha Liebke, 1928 Pseudometabletus Liebke, 1930 Schidonychus Klug, 1834 Teukrus Liebke, 1931 Wate Liebke, 1928 Tribe Hexagoniini G.Horn, 1881 Dinopelma Bates, 1889 Hexagonia Kirby, 1825 Omphreoides Fairmaire, 1896 Subfamily Dryptinae Bonelli, 1810 edit Tribe Dryptini Bonelli, 1810 Desera Hope, 1831 Drypta Latreille, 1797 Megadrypta Sciaky & Anichtchenko, 2020 Neodrypta Basilewsky, 1960 Nesiodrypta Jeannel, 1949 Prionodrypta Jeannel, 1949 Tribe Galeritini Kirby, 1825 Ancystroglossus Chaudoir, 1863 Eunostus Laporte, 1835 Galerita Fabricius, 1801 Trichognatha Latreille, 1829 Tribe Zuphiini Bonelli, 1810 Subtribe Dicrodontina Machado, 1992 Dicrodontus Chaudoir, 1872 Subtribe Leleupidiina Basilewsky, 1951 Colasidia Basilewsky, 1954 Gunvorita Landin, 1955 Leleupidia Basilewsky, 1951 Neoleleupidia Basilewsky, 1953 Paraleleupidia Basilewsky, 1951 Subtribe Planetina Jedlicka, 1941 Planetes W.S.MacLeay, 1825 Subtribe Zuphiina Bonelli, 1810 Acrogenys W.J.MacLeay, 1864 Agastus Schmidt-Goebel, 1846 Chaudoirella Mateu, 1982 Coarazuphium Gnaspini; Vanin & Godoy, 1998 Ildobates Español, 1966 Metaxidius Chaudoir, 1852 Metazuphium Mateu, 1992 Mischocephalus Chaudoir, 1863 Parazuphium Jeannel, 1942 Polistichus Bonelli, 1810 Pseudaptinus Laporte, 1834 Speothalpius B.Moore, 1995 Speozuphium B.Moore, 1995 Typhlozuphium Baehr, 2014 Zuphioides Ball & Shpeley, 2013 Zuphium Latreille, 1805 Subfamily Elaphrinae Latreille, 1802 edit Blethisa Bonelli, 1810 Diacheila Motschulsky, 1844 Elaphrus Fabricius, 1775 †Elaphrotites Haupt, 1856 Subfamily Gineminae Ball & Shpeley, 2002 edit Ginema Ball & Shpeley, 2002 Subfamily Harpalinae Bonelli, 1810 edit Tribe Anisodactylini Lacordaire, 1854 Allendia Noonan, 1974 Allocinopus Broun, 1903 Amphasia Newman, 1838 Anisodactylus Dejean, 1829 Anisostichus Emden, 1953 Cenogmus Sloane, 1898 Chydaeus Chaudoir, 1854 Crasodactylus Guérin-Méneville, 1847 Criniventer Emden, 1953 Diachromus Erichson, 1837 Dicheirus Mannerheim, 1843 Gaioxenus Broun, 1910 Geopinus LeConte, 1847 Gnathaphanus W.S.MacLeay, 1825 Gynandromorphus Dejean, 1829 Haplaner Chaudoir, 1878 Harpalomimetes Schauberger in Csiki, 1932 Hypharpax W.S.MacLeay, 1825 Maoriharpalus Larochelle & Larivière, 2005 Nornalupia Kataev, 2002 Notiobia Perty, 1830 Parabaris Broun, 1881 Progonochaetus G.Müller, 1938 Pseudanisotarsus Noonan, 1973 Pseudognathaphanus Schauberger, 1932 Pseudorhysopus Kataev & Wrase, 2001 Rhysopus Andrewes, 1929 Scybalicus Schaum, 1862 Triplosarus Bates, 1874 Tuiharpalus Larochelle & Larivière, 2005 Xestonotus LeConte, 1853 Tribe Harpalini Bonelli, 1810 Subtribe Amblystomina Fauvel, 1889 Amblystomus Erichson, 1837 Anomostomus LaFerté-Sénectère, 1853 Barysomus Dejean, 1829 Oosoma Nietner, 1857 Subtribe Ditomina Bonelli, 1810 Bronislavia Semenov, 1891 Carenochyrus Solsky, 1874 Carterus Dejean, 1830 Chilotomus Chaudoir, 1842 Ditomus Bonelli, 1810 Dixus Billberg, 1820 Eocarterus Stichel, 1923 Eucarterus Reitter, 1900 Graniger Motschulsky, 1864 Indocarterus Kataev & Wrase, 2018 Liochirus Tschitscherine, 1897 Machozetus Chaudoir, 1850 Odotoncarus Solier, 1835 Oedesis Motschulsky, 1850 Pachycarus Solier, 1835 Parapenthes Kataev & Wrase, 2018 Penthus Chaudoir, 1843 Phorticosomus Schaum, 1863 Proditomus Schauberger, 1934 Pseudaristus Reitter, 1900 Tschitscherinellus Csiki, 1906 Subtribe Harpalina Bonelli, 1810 Acinopus Dejean, 1821 Afromizonus Basilewsky, 1947 Allosiopelus N.Ito, 1995 Amblygnathus Dejean, 1829 Anisocnemus Chaudoir, 1843 Athrostictus Bates, 1878 Axinotoma Dejean, 1829 Aztecarpalus Ball, 1970 Bleusea Bedel, 1897 Boeomimetes Péringuey, 1896 Bradybaenus Dejean, 1829 Coleolissus Bates, 1892 Cratacanthus Dejean, 1829 Cratognathus Dejean, 1829 Daptus Fischer von Waldheim, 1823 Dioryche W.S.MacLeay, 1825 Discoderus LeConte, 1853 Dregus Motschulsky, 1864 Ectinothorax Alluaud, 1941 Eriophonus Tschitscherine, 1901 Euryderus LeConte, 1846 Geodromus Dejean, 1829 Harpalinus Jeannel, 1946 Harpaliscus Bates, 1892 Harpalobrachys Tschitscherine, 1899 Harpalodiodes Bousquet, 2002 Harpalomimus Facchini & Giachino, 2020 Harpalomorphus Péringuey, 1896 Harpalus Latreille, 1802 Harpathaumas Basilewsky, 1947 Harponixus Basilewsky, 1950 Hartonymus Casey, 1914 Heteracantha Brullé, 1835 Hyphaereon W.S.MacLeay, 1825 Indiophonus N.Ito, 1996 Kareya Andrewes, 1919 Lampetes Andrewes, 1940 Laparhetes Jeannel, 1946 Liodaptus Bates, 1889 Meroctenus Gemminger & Harold, 1868 Microderes Faldermann, 1836 Neoaulacoryssus Noonan, 1985 Neodiachipteryx Noonan, 1985 Neohyparpalus Clarke, 1981 Neophygas Noonan, 1976 Nesacinopus Tschitscherine, 1900 Nesarpalus Bedel, 1897 Nipponoharpalus Habu, 1973 Nothodaptus Maindron, 1906 Oesyperus Andrewes, 1923 Omostropus Péringuey, 1896 Ooidius Chaudoir, 1847 Ophoniscus Bates, 1892 Ophonus Dejean, 1821 Oxycentrus Chaudoir, 1854 Panagrius Andrewes, 1933 Pangus Dejean, 1821 Parasiopelus Basilewsky, 1946 Paraulacoryssus Shpeley; Hunting & Ball, 2017 Parophonus Ganglbauer, 1891 Paulianoscirtus Basilewsky, 1976 Penthophonus Reitter, 1900 Piosoma LeConte, 1847 Platymetopsis Ball & Maddison, 1987 Platymetopus Dejean, 1829 Prakasha Andrewes, 1919 Pseudodiachipteryx Burgeon, 1936 Pseudohyparpalus Basilewsky, 1946 Pseudoselenophorus Péringuey, 1896 Selenophorus Dejean, 1829 Selenotichnus Kataev, 1999 Siopelophonus Kataev, 2015 Siopelus Murray, 1859 Stenomorphus Dejean, 1831 Trichopselaphus Chaudoir, 1843 Trichotichnus A.Morawitz, 1863 Trichoxycentrus N.Ito, 2000 Typsiharpalus Tschitscherine, 1901 Tribe Pelmatellini Bates, 1882 Hakaharpalus Larochelle & Larivière, 2005 Kupeharpalus Larochelle & Larivière, 2005 Lecanomerus Chaudoir, 1850 Nemaglossa Solier, 1849 Notospeophonus B.Moore, 1962 Pelmatellus Bates, 1882 Syllectus Bates, 1878 Trachysarus Reed, 1874 Tribe Stenolophini Kirby, 1837 Acupalpus Latreille, 1829 Agonoleptus Casey, 1914 Amerinus Casey, 1884 Angionychus Klug, 1853 Anthracus Motschulsky, 1850 Batoscelis Dejean, 1836 Bradycellus Erichson, 1837 Cratosoma Jeannel, 1948 Cyptomicrus Vinson, 1939 Dicheirotrichus Jacquelin du Val, 1855 Egadyla Alluaud, 1916 Euthenarus Bates, 1874 Fuminoria Morita, 2006 Goniocellus Casey, 1914 Gugheorites Basilewsky, 1951 Haplanister B.Moore, 1996 Hemiaulax Bates, 1892 Hippoloetis Laporte, 1835 Idiomelas Tschitscherine, 1900 Kaffovatus Clarke, 1972 Kenyacus Alluaud, 1917 Kiwiharpalus Larochelle & Larivière, 2005 Lioholus Tschitscherine, 1897 Loxoncus Schmidt-Goebel, 1846 Pachytrachelus Chaudoir, 1852 Parabradycellus N.Ito, 2003 Paramecus Dejean, 1829 Philodes LeConte, 1861 Pholeodytes Britton, 1962 Pogonodaptus G.Horn, 1881 Polpochila Solier, 1849 Psychristus Andrewes, 1930 Rhabidius Basilewsky, 1948 Stenolophus Dejean, 1821 Uenanthracus Kasahara, 1994 Subfamily Hiletinae Schiödte, 1847 edit Eucamaragnathus Jeannel, 1938 Hiletus Schiödte, 1847 Subfamily Lebiinae Bonelli, 1810 edit Tribe Cyclosomini Laporte, 1834 Subtribe Corsyrina Ganglbauer Corsyra Dejean, 1825 Discoptera Semenov, 1889 Subtribe Cyclosomina Laporte, 1834 Cyclosomus Latreille, 1829 Sarothrocrepis Chaudoir, 1850 Tetragonoderus Dejean, 1829 Subtribe Graphipterina Latreille, 1802 Graphipterus Latreille, 1802 Piezia Brullé, 1834 Trichopiezia Nègre, 1955 Subtribe Masoreina Chaudoir, 1871 Anaulacus W.S.MacLeay, 1825 Atlantomasoreus Mateu, 1984 Leuropus Andrewes, 1947 Lophidius Dejean, 1831 Masoreus Dejean, 1821 Mnuphorus Chaudoir, 1873 Odontomasoreus Darlington, 1968 Somoplatodes Basilewsky, 1986 Somoplatus Dejean, 1829 Subtribe Nemotarsina Bates, 1883 Nemotarsus LeConte, 1853 Tribe Lachnophorini LeConte, 1853 Subtribe Calophaenina Jeannel, 1948 Calophaena Klug, 1821 Calophaenoidea Liebke, 1930 Subtribe Lachnophorina LeConte, 1853 Amphithasus Bates, 1871 Anchonoderus Reiche, 1843 Aporesthus Bates, 1871 Asklepia Liebke, 1938 Balligratus Moret & Ortuño, 2017 Calybe Laporte, 1834 Ega Laporte, 1834 Eucaerus LeConte, 1853 Euphorticus G.Horn, 1881 Grundmannius Basilewsky, 1965 Guatemalteca Erwin, 2004 Lachnaces Bates, 1872 Lachnophorus Dejean, 1831 Peruphorticus Erwin & Zamorano, 2014 Pseudophorticus Erwin, 2004 Selina Motschulsky, 1858 Tribe Lebiini Bonelli, 1810 Subtribe Agrina Kirby, 1837 Abaditicus Ball & Hilchie, 1983 Abrodiella Bousquet, 2002 Agra Fabricius, 1801 Allardina Basilewsky, 1963 Amelus Chaudoir, 1873 Anomotariella Baehr, 2012 Anomotarus Chaudoir, 1875 Apterodromites Mateu, 1976 Aspasiola Chaudoir, 1877 Calleida Latreille, 1824 Calleidomorpha Motschulsky, 1855 Callidadelpha Steinheil, 1875 Chaudoirina Mateu, 1955 Cryptobatis Eschscholtz, 1829 Cyanotarus Reed, 1874 Cylindronotum Putzeys, 1845 Demetrida White, 1846 Diabaticus Bates, 1878 Do Baehr, 2009 Dromidea Perroud & Montrouzier, 1864 Epikastea Liebke, 1935 Eujalmenus Bousquet, 2002 Euplatia Chaudoir, 1873 Eurycalleida Maindron, 1905 Falsodromius Mateu, 1976 Glycia Chaudoir, 1842 Hyboptera Chaudoir, 1873 Hybopteroides Erwin & Ball, 2012 Infernophilus Larson, 1969 Kteatus Liebke, 1936 Lipostratia Chaudoir, 1873 Merizomena Chaudoir, 1873 Mimodromius Chaudoir, 1873 Mimophilorhizus Mateu, 1993 Oechalius Liebke, 1935 Ogygium Liebke, 1935 Onota Chaudoir, 1873 Otoglossa Chaudoir, 1873 Paraglycia Bedel, 1904 Phacocerus Chaudoir, 1873 Philophuga Motschulsky, 1859 Phloeocarabus W.J.MacLeay, 1871 Plochionus Dejean, 1821 Pontonoa Liebke, 1935 Pseudotoglossa Mateu, 1961 Pylartesius Liebke, 1939 Rugitarus Baehr, 2009 Speotarus B.Moore, 1964 Straneotia Mateu, 1961 Tecnophilus Chaudoir, 1877 Teiresia Liebke, 1935 Thoasia Liebke, 1939 Titaresius Liebke, 1935 Trigonothops W.J.MacLeay, 1864 Valeriaaschero Erwin, 2004 Vianasia Mateu, 1955 Subtribe Apenina Ball Apenes LeConte, 1851 Cymindoidea Laporte, 1833 Habutarus Ball & Hilchie, 1983 Platytarus Fairmaire, 1850 Trymosternus Chaudoir, 1873 Subtribe Celaenephina Habu, 1982 Celaenephes Schmidt-Goebel, 1846 Subtribe Cymindidina Laporte, 1834 Afrotarus Jeannel, 1949 Assadecma Basilewsky, 1982 Cymindis Latreille, 1805 Hystrichopus Boheman, 1848 Leptosarcus Péringuey, 1896 Metaxymorphus Chaudoir, 1850 Petrimagnia Kryzhanovskij & Mikhailov, 1971 Pinacodera Schaum, 1857 Plagiopyga Boheman, 1848 Pseudomasoreus Desbrochers des Loges, 1904 Taridius Chaudoir, 1875 Subtribe Demetriadina Bates, 1886 Cylindrocranius Chaudoir, 1878 Demetrias Bonelli, 1810 Subtribe Dromiusina Bonelli, 1810 Afrodromius Basilewsky, 1958 Axinopalpus LeConte, 1846 Barrymooreana Baehr, 1997 Brachynopterus Bedel, 1898 Brigalowia Baehr, 2006 Calodromius Reitter, 1905 Carbonellia Mateu, 1968 Disciferella Kataev & Muilwijk, 2020 Dromiops Péringuey, 1898 Dromius Bonelli, 1810 Dromoceryx Schmidt-Goebel, 1846 Geoffreyella Baehr, 2012 Klepteromimus Péringuey, 1898 Mesolestes Schatzmayr, 1943 Metadromius Bedel, 1907 Microdaccus Schaum in Baudi di Selve, 1864 Microlestes Schmidt-Goebel, 1846 Microlestodes Baehr, 1987 Monnea Mateu, 1970 Negrea Mateu, 1968 Omophagus Andrewes, 1937 Oxoides Solier, 1849 Paradromius Fowler, 1887 Philorhizus Hope, 1838 Polyaulacus Chaudoir, 1878 Psammodromius Peyerimhoff, 1927 Pseudomonnea Mateu, 1983 Somalodromius Mateu, 1967 Xenodromius Bates, 1891 Zolotarevskyella Mateu, 1953 Subtribe Gallerucidiina Chaudoir, 1872 Gallerucidia Chaudoir, 1872 Lebidia A.Morawitz, 1862 Subtribe Lebiina Bonelli, 1810 Alkestis Liebke, 1939 Aristolebia Bates, 1892 Daer Semenov & Znojko, 1929 Lachnolebia Maindron, 1905 Lebia Latreille, 1802 Lebidema Motschulsky, 1864 Lebiomorphica Lorenz, 1998 Lebistina Motschulsky, 1864 Lebistinida Péringuey, 1898 Lionedya Chaudoir, 1871 Matabele Péringuey, 1896 Megalebia Mateu, 1972 Orthobasis Chaudoir, 1871 Pachylebia Jeannel, 1949 Pachylebiodes Mateu, 1972 Paulianolebia Mateu, 1972 Pseudopachylebia Mateu, 1972 Rhopalostyla Chaudoir, 1850 Scalidion Schmidt-Goebel, 1846 Setolebia Jedlicka, 1941 Sofota Jedlicka, 1951 Subtribe Lichnasthenina J.Thomson Astastus Péringuey, 1896 Australovelinda Baehr, 2012 Imasakaia Arai & Morita, 2010 Lichnasthenus J.Thomson, 1858 Mimovelindopsis Mateu, 1963 Velinda Andrewes, 1921 Velindastus Schüle & Lorenz, 2008 Velindomimus Jeannel, 1955 Velindopsis Burgeon, 1937 Subtribe Lionychina Jeannel, 1948 Apristomimus Mateu, 1969 Apristus Chaudoir, 1846 Eremolestes Maindron, 1905 Lionychus Wissmann, 1846 Lorestania Anichtchenko, 2011 Metablus Jedlicka, 1958 Omobrus Andrewes, 1930 Singiliomimus Péringuey, 1896 Syntomus Hope, 1838 Tilius Chaudoir, 1876 Trichidema Basilewsky, 1956 Subtribe Metallicina Basilewsky, 1984 Euproctinus Leng & Mutchler, 1927 Metallica Chaudoir, 1873 Pachycallida Jeannel, 1949 Parena Motschulsky, 1860 Subtribe Peliocypadina Basilewsky, 1984 Demetriola Jeannel, 1949 Peliocypas Schmidt-Goebel, 1846 Subtribe Pericalina Hope, 1838 Agonocheila Chaudoir, 1848 Allophanes Andrewes, 1939 Allophanopsis Louwerens, 1952 Amphimenes Bates, 1873 Amphimenoides Kirschenhofer, 1999 Antimerina Alluaud, 1897 Arsinoe Laporte, 1834 Auchmerus Andrewes, 1929 Bellavalentis Hunting & Yang, 2019 Brachichila Chaudoir, 1870 Brachyctis Chaudoir, 1870 Catascopellus Straneo, 1969 Catascopus Kirby, 1825 Cheilagona Baehr, 2006 Coptodera Dejean, 1825 Coptoglossus Chaudoir, 1870 Coptoptera Chaudoir, 1837 Coptopterella Basilewsky, 1961 Coptopterina Basilewsky, 1956 Crassagena Baehr, 2006 Cylindropectus Lorenz, 1998 Dolichoctis Schmidt-Goebel, 1846 Dontolobus Basilewsky, 1970 Drymatus Motschulsky, 1862 Eucheila Dejean, 1829 Eurycoleus Chaudoir, 1848 Eurydera Laporte, 1831 Formosiella Jedlicka, 1951 Gidda Andrewes, 1920 Glyphodactyla Chaudoir, 1837 Holcoderus Chaudoir, 1870 Horniulus Jedlicka, 1932 Labocephalus Chaudoir, 1848 Lelis Chaudoir, 1870 Lioptera Chaudoir, 1870 Lobodontidius Basilewsky, 1970 Lobodontulus Basilewsky, 1970 Lobodontus Chaudoir, 1842 Madecassina Jeannel, 1949 Mascarenhia Alluaud, 1933 Menarus Andrewes, 1939 Metascopus Basilewsky, 1970 Minuphloeus Darlington, 1968 Minuthodes Andrewes, 1941 Miscelus Klug, 1834 Mochtherus Schmidt-Goebel, 1846 Mormolyce Hagenbach, 1825 Mormolycina Jeannel, 1949 Neocoptodera Jeannel, 1949 Nycteis Laporte, 1835 Oreodicastes Maindron, 1905 Oxyodontus Chaudoir, 1870 Paradolichoctis Baehr, 2006 Pareurydera Jeannel, 1949 Pectinitarsus Fairmaire, 1881 Pericalus W.S.MacLeay, 1825 Philophlaeus Chaudoir, 1844 Phloeoxena Chaudoir, 1870 Pristacrus Chaudoir, 1870 Pseudoplatiella Baehr, 2012 Serrimargo Chaudoir, 1870 Sfitakantha Andrewes, 1919 Sinurus Chaudoir, 1870 Stenognathus Chaudoir, 1843 Stenotelus Bouchard, 1903 Stilboma Andrewes, 1933 Stricklandiana Bousquet, 2002 Tantillus Chaudoir, 1870 Thyreochaetus Basilewsky, 1959 Thyreopterus Dejean, 1831 Thysanotus Chaudoir, 1848 Trichocoptodera Louwerens, 1958 Xanthos Kirschenhofer, 2003 Xenitenopsis Basilewsky, 1956 Xenitenus Péringuey, 1896 Subtribe Physoderina Chaudoir, 1877 Allocota Motschulsky, 1860 Anchista Nietner, 1856 Dasiosoma Britton, 1937 Diamella Shi & Liang, 2013 Endynomena Chaudoir, 1873 Lachnoderma W.J.MacLeay, 1873 Metallanchista Shi & Liang, 2013 Orionella Jedlicka, 1964 Paraphaea Bates, 1873 Physodera Eschscholtz, 1829 Subtribe Pseudotrechina Basilewsky, 1984 Pseudotrechus Rosenhauer, 1856 Subtribe Singilina Jeannel, 1949 Singilis Rambur, 1837 Subtribe Somotrichina Mateu, 1963 Oecornis Britton, 1940 Paulianites Jeannel, 1949 Pephrica Alluaud, 1936 Somotrichus Seidlitz, 1887 Subtribe Sugimotoina Habu, 1975 Sugimotoa Habu, 1975 Subtribe Trichina Basilewsky, 1984 Trichis Klug, 1832 †Cymindoides Motschulsky, 1856 †Lebina Germar, 1813 †Miolebidia Zhang; Liu & Shangguan, 1989 †Protoscalidion L.Schaufuss, 1888 Tribe Odacanthini Laporte, 1834 Subtribe Actenonycina Bates, 1871 Actenonyx White, 1846 Subtribe Homethina Liebherr, 2016 Aeolodermus Andrewes, 1929 Cyphocoleus Chaudoir, 1877 Diplacanthogaster Liebke, 1932 Homethes Newman, 1842 Quammenis Erwin, 2000 Stenocheila Laporte, 1832 Subtribe Odacanthina Laporte, 1834 Anasis Laporte, 1867 Andrewesia Liebke, 1938 Arame Andrewes, 1919 Archicolliuris Liebke, 1931 Asios Liebke, 1933 Aulacolius Sloane, 1923 Basistichus Sloane, 1917 Clarencia Sloane, 1917 Colliuris DeGeer, 1774 Crassacantha Baehr, 1995 Cryptocolliuris Basilewsky, 1955 Deipyrodes Bousquet, 2002 Dicraspeda Chaudoir, 1863 Dobodura Darlington, 1968 Erectocolliuris Liebke, 1931 Eucolliuris Liebke, 1931 Eudalia Laporte, 1867 Gestroania Liebke, 1938 Giachinoana Baehr, 2003 Lachnothorax Motschulsky, 1862 Lasiocera Dejean, 1831 Mimocolliuris Liebke, 1933 Myrmecodemus Sloane, 1923 Neoeudalia Baehr, 2005 Odacantha Paykull, 1798 Ophionea Klug, 1821 Polydamasium Liebke, 1938 Porocara Sloane, 1917 Protocolliuris Liebke, 1931 Renneria Baehr, 1999 Smeringocera Chaudoir, 1863 Stenidia Brullé, 1834 Tricharnhemia Baehr, 2009 Subtribe Pentagonicina Bates, 1873 Parascopodes Darlington, 1968 Pentagonica Schmidt-Goebel, 1846 Scopodes Erichson, 1842 Tribe Perigonini G.Horn, 1881 Diploharpus Chaudoir, 1850 Mizotrechus Bates, 1872 Perigona Laporte, 1835 Ripogenites Basilewsky, 1954 Subfamily Licininae Bonelli, 1810 edit Tribe Chaetogenyini Emden, 1958 Camptotoma Reiche, 1843 Tribe Chlaeniini Brullé, 1834 Subtribe Callistina Laporte, 1834 Callistomimus Chaudoir, 1872 Callistus Bonelli, 1810 Subtribe Chlaeniina Brullé, 1834 Actodus Alluaud, 1915 Chlaenius Bonelli, 1810 Eccoptomenus Chaudoir, 1850 Ectenognathus Murray, 1858 Globulipalpus Sciaky & Facchini, 2019 Harpaglossus Motschulsky, 1858 Hololeius LaFerté-Sénectère, 1851 Holosoma Semenov, 1889 Mirachlaenius Facchini, 2011 Parachlaenius Kolbe, 1894 Perissostomus Alluaud, 1930 Procletodema Péringuey, 1898 Procletus Péringuey, 1896 Rhopalomelus Boheman, 1848 Sphodroschema Alluaud, 1930 Stenoodes Basilewsky, 1953 Straneomelus Sciaky & Facchini, 2019 Stuhlmannium Kolbe, 1894 Viridagonum Lassalle, 2015 †Rhopalochlaenius Zhang; Sun & Zhang, 1994 Tribe Licinini Bonelli, 1810 Subtribe Dicaelina Laporte, 1834 Dicaelus Bonelli, 1813 Diplocheila Brullé, 1835 Subtribe Dicrochilina Ball, 1992 Dicrochile Guérin-Méneville, 1846 Subtribe Lestignathina Ball, 1992 Atrotus Péringuey, 1896 Dilonchus Andrewes, 1936 Hormacrus Sloane, 1898 Lacordairia Laporte, 1867 Lestignathus Erichson, 1842 Microferonia Blackburn, 1890 Microzargus Sciaky & Facchini, 1997 Platylytron W.J.MacLeay, 1873 Siagonyx W.J.MacLeay, 1871 Zargus Wollaston, 1854 Subtribe Licinina Bonelli, 1810 Badister Clairville, 1806 Colpostoma Semenov, 1889 Derostichus Motschulsky, 1860 Eurygnathus Wollaston, 1854 Eutogeneius Solier, 1849 Licinus Latreille, 1802 Omestes Andrewes, 1933 Physolaesthus Chaudoir, 1850 Tribe Oodini LaFerté-Sénectère, 1851 Subtribe Dercylina Sloane, 1923 Dercylus Laporte, 1832 Subtribe Geobaenina Péringuey, 1896 Geobaenus Dejean, 1829 Subtribe Melanchitonina Jeannel, 1948 Dicaelindus W.S.MacLeay, 1825 Melanchiton Andrewes, 1940 Melanchrous Andrewes, 1940 Subtribe Oodina LaFerté-Sénectère, 1851 Acanthoodes Basilewsky, 1953 Acutosternus Lecordier & Girard, 1988 Adelopomorpha Heller, 1916 Anatrichis LeConte, 1853 Bamaroodes Gueorguiev, 2014 Brachyodes Jeannel, 1949 Chaetocrepis Chaudoir, 1857 Coptocarpus Chaudoir, 1857 Dercylinus Chaudoir, 1883 Evolenes LeConte, 1853 Holcocoleus Chaudoir, 1883 Hoplolenus LaFerté-Sénectère, 1851 Lobatodes Basilewsky, 1956 Lonchosternus LaFerté-Sénectère, 1851 Macroprotus Chaudoir, 1878 Megaloodes Lesne, 1896 Microodes Jeannel, 1949 Miltodes Andrewes, 1922 Nanodiodes Bousquet, 1996 Neoodes Basilewsky, 1953 Nothoodes Gueorguiev & Liang, 2020 Oodes Bonelli, 1810 Oodinus Motschulsky, 1865 Orthocerodus Basilewsky, 1946 Polychaetus Chaudoir, 1882 Prionognathus LaFerté-Sénectère, 1851 Protopidius Basilewsky, 1949 Pseudoodes Gueorguiev & Liang, 2020 Pseudosphaerodes Jeannel, 1949 Simous Chaudoir, 1882 Sphaerodinus Jeannel, 1949 Sphoerodes Chaudoir, 1883 Stenocrepis Chaudoir, 1857 Sundaoodes Gueorguiev & Liang, 2020 Systolocranius Chaudoir, 1857 Thaioodes Gueorguiev, 2014 Thryptocerus Chaudoir, 1878 Trichopalpoodes B.Gueorguiev & J.Schmidt, 2016 Subfamily Loricerinae Bonelli, 1810 edit Loricera Latreille, 1802 Subfamily Melaeninae Csiki, 1933 edit Cymbionotum Baudi di Selve, 1864 Melaenus Dejean, 1831 Subfamily Migadopinae Chaudoir, 1861 edit Tribe Amarotypini Erwin, 1985 Amarotypus Bates, 1872 Migadopiella Baehr, 2009 Tribe Migadopini Chaudoir, 1861 Antarctonomus Chaudoir, 1861 Aquilex Moret, 1989 Calathosoma Jeannel, 1938 Calyptogonia Sloane, 1920 Decogmus Sloane, 1915 Dendromigadops Baehr, 2013 Lissopterus G.R.Waterhouse, 1843 Loxomerus Chaudoir, 1842 Migadopidius Jeannel, 1938 Migadops G.R.Waterhouse, 1842 Monolobus Solier, 1849 Nebriosoma Laporte, 1867 Pseudomigadops Jeannel, 1938 Rhytidognathus Chaudoir, 1861 Stichonotus Sloane, 1910 Taenarthrus Broun, 1914 Subfamily Nebriinae Laporte, 1834 edit Tribe Cicindini Csiki, 1927 Archaeocindis Kavanaugh & Erwin, 1991 Cicindis Bruch, 1908 Tribe Nebriini Laporte, 1834 Leistus Frölich, 1799 Nebria Latreille, 1802 Nippononebria Ueno, 1955 †Archaeonebria Kavanaugh & J.Schmidt, 2019 †Ledouxnebria Deuve, 1998 Tribe Notiokasiini Kavanaugh & Nègre, 1983 Notiokasis Kavanaugh & Nègre, 1983 Tribe Notiophilini Motschulsky, 1850 Notiophilus Duméril, 1805 Tribe Opisthiini Dupuis, 1912 Opisthius Kirby, 1837 Paropisthius Casey, 1920 Tribe Pelophilini Kavanaugh, 1996 Pelophila Dejean, 1821 Subfamily Nototylinae Bänninger, 1927 edit Nototylus Gemminger & Harold, 1868 Subfamily Omophroninae Bonelli, 1810 edit Omophron Latreille, 1802 Subfamily Orthogoniinae Schaum, 1857 edit Tribe Amorphomerini Sloane, 1923 Amorphomerus Sloane, 1923 Tribe Idiomorphini Bates, 1891 Subtribe Idiomorphina Bates, 1891 Idiomorphus Chaudoir, 1846 Rathymus Dejean, 1831 Strigia Brullé, 1835 Subtribe Perochnoristhina Basilewsky, 1973 Perochnoristhus Basilewsky, 1973 Tribe Orthogoniini Schaum, 1857 Subtribe Glyptina G.Horn, 1881 Glyptus Brullé, 1837 Neoglyptus Basilewsky, 1953 Subtribe Orthogoniina Schaum, 1857 Actenoncus Chaudoir, 1872 Anoncopeucus Chaudoir, 1872 Hexachaetus Chaudoir, 1872 Neoorthogonius Tian & Deuve, 2006 Nepalorthogonius Habu, 1979 Orthogonius W.S.MacLeay, 1825 Subfamily Panagaeinae Bonelli, 1810 edit Tribe Brachygnathini Basilewsky, 1946 Brachygnathus Perty, 1830 Tribe Panagaeini Bonelli, 1810 Subtribe Bascanina Basilewsky, 1953 Bascanus Péringuey, 1896 Subtribe Panagaeina Bonelli, 1810 Adischissus Fedorenko, 2015 Calathocosmus Emden, 1928 Cintaroa Kasahara, 1989 Cintaromorpha Häckel & Anichtchenko, 2016 Coptia Brullé, 1835 Craspedophorus Hope, 1838 Dischissus Bates, 1873 Epigraphodes Basilewsky, 1967 Epigraphus Chaudoir, 1869 Euschizomerus Chaudoir, 1850 Geobius Dejean, 1831 Micrixys LeConte, 1854 Microschemus Andrewes, 1940 Panagaeus Latreille, 1802 Paregraphus Basilewsky, 1967 Peronomerus Schaum, 1854 Psecadius Alluaud, 1911 Tefflus Leach in Samouelle, 1819 Tinoderus Chaudoir, 1879 Tinognathus Chaudoir, 1879 Trichisia Motschulsky, 1865 Tribe Peleciini Chaudoir, 1880 Subtribe Agonicina Sloane, 1920 Agonica Sloane, 1920 Pseudagonica B.Moore, 1960 Subtribe Peleciina Chaudoir, 1880 Ardistomopsis Straneo & Ball, 1989 Disphericus G.R.Waterhouse, 1842 Dyschiridium Chaudoir, 1861 Eripus Dejean, 1829 Pelecium Kirby, 1819 Stricteripus Straneo & Ball, 1989 Subfamily Patrobinae Kirby, 1837 edit Tribe Lissopogonini Zamotajlov, 2000 Lissopogonus Andrewes, 1923 Tribe Patrobini Kirby, 1837 Subtribe Deltomerina Chaudoir, 1872 Deltomerus Motschulsky, 1850 Subtribe Deltomerodina Zamotajlov, 2002 Deltomerodes Deuve, 1992 Subtribe Patrobina Kirby, 1837 Apatrobus Habu & Baba, 1960 Apenetretus Kurnakov, 1960 Archipatrobus Zamotajlov, 1992 Caelopenetretus Zamotajlov & Ito, 2000 Chaetapatrobus Lafer, 1996 Chinapenetretus Kurnakov, 1963 Dimorphopatrobus Casale & Sciaky, 1994 Diplous Motschulsky, 1850 Grandipenetretus Zamotajlov & Sciaky, 1999 Himalopenetretus Zamotajlov, 2002 Indopatrobus Zamotajlov & Wrase, 2006 Ledouxius Zamotajlov, 1992 Minipenetretus Zamotajlov, 2002 Minypatrobus Ueno, 1955 Naxipenetretus Zamotajlov, 1999 Parapatrobus Zamotajlov, 1992 Parapenetretus Kurnakov, 1960 Patanitretus Zamotajlov, 2002 Patrobus Dejean, 1821 Penetretus Motschulsky, 1865 Platidiolus Chaudoir, 1878 Platypatrobus Darlington, 1938 Prodiplous Zamotajlov & Sciaky, 2006 Qiangopatrobus Zamotajlov, 2002 Quasipenetretus Zamotajlov, 2002 Tibetopenetretus Zamotajlov & Sciaky, 1999 Subfamily Paussinae Latreille, 1806 edit Tribe Metriini LeConte, 1853 Subtribe †Kryzhanovskianina Deuve, 2019 †Kryzhanovskiana Kataev & Kirejtshuk, 2019 Metrius Eschscholtz, 1829 Sinometrius Wrase & J.Schmidt, 2006 Tribe Ozaenini Hope, 1838 Subtribe Microzaenina Deuve, 2019 Microzaena Fairmaire, 1901 Subtribe Mystropomina G.Horn, 1881 Mystropomus Chaudoir, 1848 Subtribe Ozaenina Hope, 1838 Crepidozaena Deuve, 2001 Entomoantyx Ball & McCleve, 1990 Filicerozaena Deuve, 2001 Gibbozaena Deuve, 2001 Goniotropis G.R.Gray, 1831 Inflatozaena Deuve, 2001 Mimozaena Deuve, 2001 Ozaena Olivier, 1812 Pachyteles Perty, 1830 Physea Brullé, 1835 Physeomorpha Ogueta, 1963 Platycerozaena Bänninger, 1927 Proozaena Deuve, 2001 Serratozaena Deuve, 2001 Tachypeles Deuve, 2001 Tropopsis Solier, 1849 Subtribe Pseudozaenina Sloane, 1905 Anentmetus Andrewes, 1924 Dhanya Andrewes, 1919 Eustra Schmidt-Goebel, 1846 Itamus Schmidt-Goebel, 1846 Pseudozaena Laporte, 1834 Sphaerostylus Chaudoir, 1848 Tribe Paussini Latreille, 1806 Subtribe Carabidomemnina Wasmann, 1928 Carabidomemnus Kolbe, 1924 Eohomopterus Wasmann, 1919 Subtribe Cerapterina Billberg, 1820 Arthropterus W.S.MacLeay, 1838 Cerapterus Swederus, 1788 Megalopaussus Lea, 1906 Mesarthropterus Wasmann, 1926 Subtribe Heteropaussina Janssens, 1950 Heteropaussus J.Thomson, 1860 Subtribe Homopterina Wasmann, 1920 Homopterus Westwood, 1841 Subtribe Paussina Latreille, 1806 Allorhopalus Nagel, 2017 Ceratoderus Westwood, 1841 Euplatyrhopalus Desneux, 1905 Lebioderus Westwood, 1838 Leleupaussus Luna de Carvalho, 1962 Melanospilus Westwood, 1847 Paussomorphus Raffray, 1885 Paussus Linnaeus, 1775 Platyrhopalides Wasmann, 1918 Platyrhopalopsis Desneux, 1905 Platyrhopalus Westwood, 1833 Pterorhopalus Maruyama, 2011 Subtribe Pentaplatarthrina Jeannel, 1946 Hexaplatarthrus Jeannel, 1955 Pentaplatarthrus Westwood, 1833 Subtribe †Arthropteritina Luna de Carvalho, 1961 †Arthropterites Wasmann, 1926 †Cerapterites Wasmann, 1926 †Protocerapterus Wasmann, 1926 Subtribe †Eopaussina Luna de Carvalho, 1951 †Eopaussus Wasmann, 1926 Tribe Protopaussini Gestro, 1892 Protopaussus Gestro, 1892 Subfamily Platyninae Bonelli, 1810 edit Tribe Omphreini Ganglbauer, 1891 Omphreus Dejean, 1828 Tribe Platynini Bonelli, 1810 Subtribe Enoicina Basilewsky, 1985 Abacetodes Straneo, 1939 Enoicus Péringuey, 1896 Subtribe Platynina Bonelli, 1810 Achaetocephala Habu, 1975 Achaetoprothorax Habu, 1978 Aepsera Chaudoir, 1874 Agelaea Gené, 1839 Agonidium Jeannel, 1948 Agonobembix Jeannel, 1948 Agonoriascus Basilewsky, 1985 Agonorites Jeannel, 1951 Agonum Bonelli, 1810 Altagonum Darlington, 1952 Anchomenus Bonelli, 1810 Andinocolpodes Perrault, 1991 Andrewesius Andrewes, 1939 Aparupa Andrewes, 1930 Archagonum Basilewsky, 1953 Archicolpodes J.Schmidt, 2001 Arhytinus Bates, 1889 Atranodes Jedlicka, 1953 Atranus LeConte, 1847 Austroglyptolenus Roig-Juñent, 2003 Beckeria Jedlicka, 1931 Blackburnia Sharp, 1878 Bothrocolpodes Basilewsky, 1985 Bruskespar Morvan, 1998 Bruskmoal Morvan, 1998 Bryanites Valentine, 1987 Callidagonum Lorenz, 1998 Cardiomera Bassi, 1834 Catacolpodes Basilewsky, 1985 Celaenagonum Habu, 1978 Chaetagonum Burgeon, 1933 Chaetosaurus J.Schmidt, 2001 Cinctagonum Baehr, 2012 Cistelagonum Baehr, 2012 Collagonum Baehr, 1995 Colpodes W.S.MacLeay, 1825 Colpoides Jedlicka, 1931 Colpomimus Basilewsky, 1985 Colposphodrus Jedlicka, 1953 Ctenognathus Fairmaire, 1843 Cymenopterus Jeannel, 1948 Cyrtopilus Basilewsky, 1985 Dalatagonum Fedorenko, 2011 Deliaesianum Morvan, 1999 Deltocolpodes Morvan, 1992 Dendragonum Jeannel, 1948 Diacanthostylus Habu, 1978 Dicranoncoides Habu, 1978 Dicranoncus Chaudoir, 1850 Dinocolpodes J.Schmidt, 2001 Dirotus W.S.MacLeay, 1825 Dister Morvan, 2006 Dolichocolpodes Basilewsky, 1985 Dyscolus Dejean, 1831 Elliptoleus Bates, 1882 Epicolpodes Basilewsky, 1985 Eucolpodes Jeannel, 1948 Euleptus Klug, 1833 Euplynes Schmidt-Goebel, 1846 Feroniascus Jeannel, 1951 Fortagonum Darlington, 1952 Galiciotyphlotes Assmann, 1999 Gastragonum Darlington, 1952 Glaucagonum Habu, 1978 Glyptolenoides Perrault, 1991 Glyptolenus Bates, 1878 Gyrochaetostylus Habu, 1978 Habragonum Ueno, 1964 Hannaphota Landin, 1955 Haplocolpodes Jeannel, 1951 Haplopeza Boheman, 1848 Helluocolpodes Liebherr, 2005 Hemiplatynus Casey, 1920 Henvelik Morvan, 1999 Herculagonum Baehr, 2002 Hikosanoagonum Habu, 1954 Idiagonum Darlington, 1952 Idiastes Andrewes, 1931 Idiocolpodes Basilewsky, 1985 Incagonum Liebherr, 1994 Iridagonum Darlington, 1952 Ischnagonum Kasahara & Satô, 1997 Jocqueius Basilewsky, 1988 Jujiroa Ueno, 1952 Kalchdigor Morvan, 1999 Kar Morvan, 1998 Karnes Morvan, 2010 Kaszabellus Jedlicka, 1954 Kiwiplatynus Larochelle & Larivière, 2021 Klapperichella Jedlicka, 1956 Kuceraianum Morvan, 2002 Kupeplatynus Larochelle & Larivière, 2021 Laevagonum Darlington, 1952 Lassalleianum Morvan, 1999 Lepcha Andrewes, 1930 Leptagonum Kolbe, 1897 Leptocolpodes Basilewsky, 1985 Letouzeya Bruneau de Miré, 1982 Liagonum Jeannel, 1948 Liamegalonychus Basilewsky, 1963 Liocolpodes Basilewsky, 1985 Lissagonum Habu, 1978 Lithagonum Darlington, 1952 Lobocolpodes Basilewsky, 1985 Lorostema Motschulsky, 1865 Loxocrepis Eschscholtz, 1829 Lucicolpodes J.Schmidt, 2000 Maculagonum Darlington, 1952 Maoriplatynus Larochelle & Larivière, 2021 Meleagros Kirschenhofer, 1999 Mesocolpodes Basilewsky, 1985 Metacolpodes Jeannel, 1948 Mexisphodrus Barr, 1965 Montagonum Darlington, 1952 Mooreagonum Baehr, 2016 Morimotoidius Habu, 1954 Nebriagonum Darlington, 1952 Negreum Habu, 1958 Neobatenus Jeannel, 1948 Neocolpodes Jeannel, 1948 Neodendragonum Basilewsky, 1953 Neomegalonychus Jeannel, 1948 Nesiocolpodes Jeannel, 1948 Nipponagonum Habu, 1978 Notagonum Darlington, 1952 Notocolpodes Basilewsky, 1985 Notoplatynus B.Moore, 1985 Nymphagonum Habu, 1978 Olisthopus Dejean, 1828 Oncostylus Habu, 1978 Onotokiba Alluaud, 1927 Onycholabis Bates, 1873 Onypterygia Dejean, 1831 Orophicus Alluaud, 1925 Orthotrichus Peyron, 1856 Oxygonium Basilewsky, 1951 Oxypselaphus Chaudoir, 1843 Pachybatenus Basilewsky, 1973 Pachyferonia Jeannel, 1951 Paracolpodes Basilewsky, 1985 Paraliagonum Basilewsky, 1957 Paramegalonychus Basilewsky, 1953 Paranchodemus Habu, 1978 Paranchus Lindroth, 1974 Paraplatynus Baehr, 2016 Pawgammm Morvan, 1999 Platyagonum Habu, 1978 Platynus Bonelli, 1810 Plaumannium Liebke, 1939 Plicagonum Darlington, 1952 Ponapagonum Darlington, 1970 Potamagonum Darlington, 1952 Praepristus Kirschenhofer, 1999 Promecoptera Dejean, 1831 Promegalonychus Basilewsky, 1953 Prophenorites Basilewsky, 1985 Prosphodrus Britton, 1959 Protocolpodes Basilewsky, 1985 Pseudanchomenus Tarnier, 1860 Pseudobatenus Basilewsky, 1951 Pseudomegalonychus Basilewsky, 1950 Rhadine LeConte, 1846 Rupa Jedlicka, 1935 Scotagonum Habu, 1978 Sericoda Kirby, 1837 Shibataia Habu, 1978 Sinocolpodes J.Schmidt, 2001 Skoeda Morvan, 1995 Skorlagad Morvan, 1999 Skouedhirraad Morvan, 1999 Sophroferonia Alluaud, 1933 Speagonum B.Moore, 1977 Speocolpodes Barr, 1974 Speokokosia Alluaud, 1932 Sperkanhir Morvan, 2010 Stenocnemus Mannerheim, 1837 Straneoa Basilewsky, 1953 Syletor Tschitscherine, 1899 Takasagoagonum Habu, 1977 Tanystoma Motschulsky, 1845 Tarsagonum Darlington, 1952 Tetraleucus Casey, 1920 Tostkar Morvan, 1998 Trogloagonum Casale, 1982 Tuiplatynus Larochelle & Larivière, 2021 Vitagonum B.Moore, 1999 Vulcanophilus Heller, 1898 Xestagonum Habu, 1978 †Praeanchodemus J.Schmidt; Göpel & Will, 2017 Tribe Sphodrini Laporte, 1834 Subtribe Atranopsina Baehr, 1982 Amaroschema Jeannel, 1943 Amphimasoreus Piochard de la Brûlerie, 1875 Atranopsis Baehr, 1982 Broter Andrewes, 1923 Gomerina Bolivar y Pieltain, 1940 Paraeutrichopus Mateu, 1954 Platyderus Stephens, 1827 Pseudomyas Uyttenboogaart, 1929 Pseudoplatyderus Bolivar y Pieltain, 1940 Subtribe Calathina Laporte, 1834 Calathus Bonelli, 1810 Thermoscelis Putzeys, 1873 Subtribe Dolichina Brullé, 1834 Acalathus Semenov, 1889 Anchomenidius Heyden, 1880 Casaleius Sciaky & Wrase, 1998 Dolichus Bonelli, 1810 Doliodactyla Sciaky & Wrase, 1998 Morphodactyla Semenov, 1889 Xestopus Andrewes, 1937 Subtribe Pristosiina Lindroth, 1956 Pristosia Motschulsky, 1865 Subtribe Sphodrina Laporte, 1834 Calathidius Putzeys, 1873 Cephalosdrophus Lassalle & Marcilhac, 1999 Eosphodrus Casale, 1988 Eremosphodrus Semenov, 1909 Himalosphodrus Casale, 1988 Hystricosphodrus Casale & Giachino, 2004 Ifridytes Deuve & Queinnec, 1994 Laemostenus Bonelli, 1810 Licinopsis Bedel, 1899 Miquihuana Barr, 1982 Pseudotaphoxenus L.Schaufuss, 1865 Reflexisphodrus Casale, 1988 Sphodropsis Seidlitz, 1887 Sphodrus Clairville, 1806 Stenolepta Semenov, 1889 Taphoxenus Motschulsky, 1850 Subtribe Synuchina Lindroth, 1956 Nipponosynuchus Morita, 1998 Parabroscus Lindroth, 1956 Synuchus Gyllenhal, 1810 Trephionus Bates, 1883 Tribe Axinidiini Basilewsky, 1963 Axinidium Sturm, 1843 Holaxinidium Basilewsky, 1963 Metaxinidium Basilewsky, 1963 Paraxinidium Basilewsky, 1963 Tribe Dalyatini Mateu, 2002 Dalyat Mateu, 2002 Tribe Promecognathini LeConte, 1853 Promecognathus Chaudoir, 1846 Tribe †Palaeoaxinidiini McKay, 1991 †Palaeoaxinidium McKay, 1991 Adelotopus Hope, 1836 Cainogenion Notman, 1925 Cryptocephalomorpha Ritsema, 1875 Guyanemorpha Erwin, 2013 Manumorpha Erwin & Geraci, 2008 Notopseudomorpha Baehr, 1997 Paussotropus C.O.Waterhouse, 1877 Pseudomorpha Kirby, 1823 Samiriamorpha Erwin & Geraci, 2008 Sphallomorpha Westwood, 1840 Tuxtlamorpha Erwin & Geraci, 2008 Yasunimorpha Erwin & Geraci, 2008 Subfamily Psydrinae LeConte, 1853 edit Tribe Gehringiini Darlington, 1933 Subtribe Gehringiina Darlington, 1933 Gehringia Darlington, 1933 Subtribe Helenaeina Deuve, 2007 Afrogehringia Baehr; Schüle & Lorenz, 2009 Helenaea Schatzmayr & Koch, 1934 Tribe Moriomorphini Sloane, 1890 Subtribe Amblytelina Blackburn, 1892 Amblytelus Erichson, 1842 Dystrichothorax Blackburn, 1892 Epelyx Blackburn, 1892 Mecyclothorax Sharp, 1903 Meonis Laporte, 1867 Paratrichothorax Baehr, 2004 Pseudamblytelus Baehr, 2004 Raphetis B.Moore, 1963 Selenochilus Chaudoir, 1878 Trichamblytelus Baehr, 2004 Subtribe Moriomorphina Sloane, 1890 Celanida Laporte, 1867 Melisodera Westwood, 1835 Molopsida White, 1846 Moriodema Laporte, 1867 Moriomorpha Laporte, 1867 Neonomius B.Moore, 1963 Pterogmus Sloane, 1920 Rhaebolestes Sloane, 1903 Rossjoycea Liebherr, 2011 Sitaphe B.Moore, 1963 Spherita Liebherr, 2020 Tarastethus Sharp, 1883 Teraphis Laporte, 1867 Theprisa B.Moore, 1963 Trephisa B.Moore, 1963 Trichopsida Larochelle & Larivière, 2013 Subtribe Tropopterina Sloane, 1898 Pharetis Liebherr, 2020 Tropopterus Solier, 1849 Tribe Psydrini LeConte, 1853 Laccocenus Sloane, 1890 Nomius Laporte, 1835 Psydrus LeConte, 1846 Subfamily Pterostichinae Bonelli, 1810 edit Tribe Chaetodactylini Tschitscherine, 1903 Chaetodactyla Tschitscherine, 1897 Tribe Cnemalobini Germain, 1911 Cnemalobus Guérin-Méneville, 1838 Tribe Cratocerini Lacordaire, 1854 Subtribe Catapieseina Bates, 1882 Catapiesis Solier, 1835 Homalomorpha Brullé, 1837 Subtribe Cratocerina Lacordaire, 1854 Brachidius Chaudoir, 1852 Cratocerus Dejean, 1829 Oxyglychus Straneo, 1938 Subtribe Drimostomatina Chaudoir, 1873 Abacaelostus Straneo, 1952 Andrewesinulus Straneo, 1938 Apsidocnemus Alluaud, 1936 Barylaus Liebherr, 1985 Caecocaelus Straneo, 1949 Caelostomus W.S.MacLeay, 1825 Camptogenys Tschitscherine, 1899 Capabatus Csiki, 1930 Crenulostrigus Straneo, 1942 Cyrtolaus Bates, 1882 Dactyleurys Tschitscherine, 1899 Dactylinius Straneo, 1941 Diachipteryx Alluaud, 1925 Diceromerus Chaudoir, 1873 Dromistomus Jeannel, 1948 Drymonaxus Straneo, 1941 Feostoma Straneo, 1941 Hemitelestus Alluaud, 1895 Hoplizomenus Chaudoir, 1873 Leleuporites Straneo, 1960 Madapelmus Dajoz, 1985 Monodryxus Straneo, 1942 Pachycaecus Straneo, 1971 Pachyroxochus Straneo, 1942 Platyxythrius Lorenz, 1998 Stegazopteryx Will, 2004 Stomonaxellus Tschitscherine, 1901 Strigomerodes Straneo, 1939 Strigomerus Chaudoir, 1873 Trichillinus Straneo, 1938 Tribe Microcheilini Jeannel, 1948 Microcheila Brullé, 1835 Tribe Morionini Brullé, 1837 Buderes Murray, 1857 Hyperectenus Alluaud, 1935 Hyperion Laporte, 1834 Megamorio Chaudoir, 1880 Morion Latreille, 1810 Morionidius Chaudoir, 1880 Moriosomus Motschulsky, 1855 Platynodes Westwood, 1847 Stereostoma Murray, 1857 Tribe Pterostichini Bonelli, 1810 Subtribe Abacetina Chaudoir, 1873 Abacaecus Allegro & Giachino, 2020 Abacetus Dejean, 1828 Aristopus LaFerté-Sénectère, 1853 Celioinkosa Straneo, 1951 Chlaeminus Motschulsky, 1865 Colpodichius Straneo, 1952 Cosmodiscus Sloane, 1907 Cyrtomoscelis Chaudoir, 1874 Distrigidius Jeannel, 1948 Ecnomolaus Bates, 1892 Haptoderidius Straneo, 1975 Haptoderodes Straneo, 1986 Holconotus Schmidt-Goebel, 1846 Inkosa Péringuey, 1926 Mateuellus Deuve, 1990 Metabacetus Bates, 1892 Metaxellus Straneo, 1960 Metaxys Chaudoir, 1857 Novillidius Straneo, 1941 Oodinkosa Straneo, 1939 Ophonichius Straneo, 1942 Pediomorphus Chaudoir, 1878 Pioprosopus Tschitscherine, 1899 Pollicobius Vinson, 1939 Prostalomus Basilewsky, 1950 Pseudabacetus Burgeon, 1935 Pterostillichus Straneo, 1949 Rhagadillius Straneo, 1951 Tiferonia Darlington, 1962 Trachelocyphoides Straneo, 1942 Trachelocyphus Tschitscherine, 1900 Subtribe Euchroina Chaudoir, 1874 Abacillius Straneo, 1949 Abacillodes Straneo, 1988 Abaris Dejean, 1831 Apsaustodon Tschitscherine, 1901 Argutoridius Chaudoir, 1876 Blennidus Motschulsky, 1866 Blenniventer Straneo, 1986 Bothynoproctus Tschitscherine, 1900 Cedrorum Borges & A.Serrano, 1993 Cephalostichus Straneo, 1977 Cynthidia Chaudoir, 1874 Euchroa Brullé, 1835 Eumara Tschitscherine, 1901 Gastrogmus Sloane, 1915 Haplobothynus Tschitscherine, 1901 Litarthrum Sloane, 1915 Lobobrachus Sharp, 1885 Marsyas Putzeys, 1845 Meropalpus Tschitscherine, 1900 Microcephalus Dejean, 1828 Neotalus Will, 2002 Oribazus Chaudoir, 1874 Orthomus Chaudoir, 1838 Pachythecus Chaudoir, 1874 Paniestichus Will, 2011 Parorthomus B.Gueorguiev; Wrase & Farkac, 2014 Phaenaulax Tschitscherine, 1898 Prosopogmus Chaudoir, 1865 Pseudabarys Chaudoir, 1874 Setalimorphus Sloane, 1895 Setalis Laporte, 1867 Simodontus Chaudoir, 1843 Trirammatus Chaudoir, 1835 Subtribe Loxandrina Erwin & Sims, 1984 Cerabilia Laporte, 1867 Oxycrepis Reiche, 1843 Zeodera Laporte, 1867 Subtribe Metiina Straneo, 1951 Abropus G.R.Waterhouse, 1842 Antarctiola Straneo, 1977 Feroniola Tschitscherine, 1900 Kuschelinus Straneo, 1963 Metius Curtis, 1839 Subtribe Pterostichina Bonelli, 1810 Abacidus LeConte, 1863 Abacoleptus Fauvel, 1903 Abacomorphus Chaudoir, 1878 Abacophrastus Will, 2011 Abacops Tschitscherine, 1902 Abax Bonelli, 1810 Allotriopus Bates, 1882 Analoma Darlington, 1971 Apophylon B.Gueorguiev & Sciaky, 2015 Aristochroa Tschitscherine, 1898 Aristochroodes Marcilhac, 1993 Aulacopodus Britton, 1940 Basilewskya Straneo, 1948 Camptoscelis Dejean, 1828 Castelnaudia Tschitscherine, 1891 Catadromus W.S.MacLeay, 1825 Chaetauchenium Tschitscherine, 1900 Chalcochrous Chaudoir, 1838 Cophosomorpha Tschitscherine, 1891 Cratoferonia Tschitscherine, 1902 Cratogaster Blanchard, 1843 Cuneipectus Sloane, 1907 Cyclotrachelus Chaudoir, 1838 Cyrtoderus Hope, 1842 Darodilia Laporte, 1867 Delinius Westwood, 1864 Eucamptognathus Chaudoir, 1837 Eudromus Klug, 1835 Euryabax Fauvel, 1903 Euryaptus Bates, 1892 Eurypercus Jeannel, 1948 Eurystomis Chaudoir, 1878 Gastrellarius Casey, 1918 Gourlayia Britton, 1964 Harpostomus Chaudoir, 1856 Henrotiochoromus Busulini, 1958 Henrotius Jeannel, 1953 Holcaspis Chaudoir, 1865 Hybothecus Chaudoir, 1874 Leiolesticus Roux; Lassalle & Dubault, 2016 Lesticus Dejean, 1828 Licentius Jedlicka, 1939 Liopasa Tschitscherine, 1901 Lophoglossus LeConte, 1853 Loxogenius Sloane, 1907 Mecynognathus W.J.MacLeay, 1873 Megadromus Motschulsky, 1866 Molopidius Jeannel, 1942 Molopinus Jeannel, 1948 Molops Bonelli, 1810 Myas Sturm, 1826 Neoferonia Britton, 1940 Nesites Andrewes, 1939 Nirmala Andrewes, 1930 Notabax B.Moore, 1976 Notolestus Sloane, 1894 Notonomus Chaudoir, 1865 Nurus Motschulsky, 1866 Oberthueria Vuillet, 1911 Ogmophora Tschitscherine, 1896 Onawea Johns, 2007 Oscadytes Lagar Mascaro, 1975 Pachymorphus Chaudoir, 1838 Paranurus Tschitscherine, 1901 Pareuryaptus Dubault; Lassalle & Roux, 2008 Parhypates Motschulsky, 1866 Pedius Motschulsky, 1850 Percolaus Bates, 1882 Percus Bonelli, 1810 Peyrieraselus Deuve, 1981 Piesmus LeConte, 1846 Platycoelus Blanchard, 1843 Platysmodes Fauvel, 1903 Plocamostethus Britton, 1940 Poecilinus Jeannel, 1948 Poecilus Bonelli, 1810 Pseudoceneus Tschitscherine, 1891 Pterostichus Bonelli, 1810 Rhabdotus Chaudoir, 1865 Rhytiferonia Darlington, 1962 Rhytisternus Chaudoir, 1865 Sarticus Motschulsky, 1866 Secatophus Laporte, 1867 Setalidius Chaudoir, 1878 Speluncarius Reitter, 1886 Speomolops Patrizi, 1955 Sphodrosomus Perroud, 1864 Stenochoromus L.Miller, 1866 Stereocerus Kirby, 1837 Sthenocranion Tschitscherine, 1899 Stomis Clairville, 1806 Straneostichus Sciaky, 1994 Styracoderus Chaudoir, 1874 Tanythrix Schaum, 1858 Taphoxomimus Straneo, 1975 Tapinopterus Schaum, 1858 Teratotarsa Tschitscherine, 1893 Teropha Laporte, 1867 Trichosternus Chaudoir, 1865 Trigonaptus Fedorenko, 2020 Trigonotoma Dejean, 1828 Troglorites Jeannel, 1919 Tropidocerus Chaudoir, 1878 Typhlochoromus Moczarski, 1913 Wahlbergiana Bousquet, 2002 Xenion Tschitscherine, 1902 Zariquieya Jeannel, 1924 Zeopoecilus Sharp, 1886 Tribe Zabrini Bonelli, 1810 Subtribe Amarina Zimmermann, 1832 Amara Bonelli, 1810 Pseudamara Lindroth, 1968 Subtribe Zabrina Bonelli, 1810 Zabrus Clairville, 1806 Subfamily Rhysodinae Laporte, 1840 edit Tribe Clinidiini R.T. & J.R.Bell, 1978 Clinidium Kirby, 1830 Grouvellina R.T. & J.R.Bell, 1978 Rhyzodiastes Fairmaire, 1895 Tribe Dhysorini R.T. & J.R.Bell, 1978 Dhysores Grouvelle, 1903 Neodhysores R.T. & J.R.Bell, 1978 Tangarona R.T. & J.R.Bell, 1982 Tribe Leoglymmiini R.T. & J.R.Bell, 1978 Leoglymmius R.T. & J.R.Bell, 1978 Tribe Medisorini R.T. & J.R.Bell, 1987 Medisores R.T. & J.R.Bell, 1987 Tribe Omoglymmiini R.T. & J.R.Bell, 1978 Arrowina R.T. & J.R.Bell, 1978 Omoglymmius Ganglbauer, 1891 Plesioglymmius R.T. & J.R.Bell, 1978 Shyrodes Grouvelle, 1903 Srimara R.T. & J.R.Bell, 1978 Xhosores R.T. & J.R.Bell, 1978 Yamatosa R.T. & J.R.Bell, 1979 Tribe Rhysodini Laporte, 1840 Kaveinga R.T. & J.R.Bell, 1978 Kupeus R.T. & J.R.Bell, 1982 Rhysodes Germar, 1822 Tribe Sloanoglymmiini R.T. & J.R.Bell, 1991 Sloanoglymmius R.T. & J.R.Bell, 1991 Subfamily Scaritinae Bonelli, 1810 edit Tribe Clivinini Rafinesque, 1815 Subtribe Androzelmina R.T.Bell, 1998 Androzelma Dostal, 1993 Subtribe Ardistomina Putzeys, 1867 Ardistomis Putzeys, 1846 Aspidoglossa Putzeys, 1846 Kearophus Dajoz, 2004 Semiardistomis Kult, 1950 Subtribe Clivinina Rafinesque, 1815 Afrosyleter Basilewsky, 1959 Ancus Putzeys, 1867 Basilewskyana Kult, 1959 Brachypelus Putzeys, 1867 Cameroniola Baehr, 2000 Climax Putzeys, 1861 Clivina Latreille, 1802 Clivinarchus Sloane, 1896 Cryptomma Putzeys, 1846 Lachenus Putzeys, 1846 Leleuporella Basilewsky, 1956 Nannoryctes Baehr, 2000 Nyctosyles Putzeys, 1867 Orictites Andrewes, 1931 Platysphyrus Sloane, 1905 Psilidius Jeannel, 1957 Pyramoides Bousquet, 2002 Rhysocara Sloane, 1916 Rubidiclivina Baehr, 2015 Rugiluclivina Balkenohl, 1996 Semiclivina Kult, 1947 Sinesetosa Balkenohl, 1996 Syleter Andrewes, 1941 Thliboclivina Kult, 1959 Trilophidius Jeannel, 1957 Trilophus Andrewes, 1927 Trogloclivina Deuve, 2003 Whiteheadiana Perrault, 1994 Subtribe Forcipatorina Bänninger, 1938 Camptidius Putzeys, 1867 Camptodontus Dejean, 1826 Forcipator Maindron, 1904 Kultianella Perrault, 1994 Mesus Chevrolat, 1858 Obadius Burmeister, 1875 Oxygnathopsis Louwerens, 1953 Oxygnathus Dejean, 1826 Scolyptus Putzeys, 1861 Stratiotes Putzeys, 1846 Subtribe Reicheiina Jeannel, 1957 Alpiodytes Jeannel, 1957 Asioreicheia Bulirsch & Magrini, 2014 Catalanodytes Sciaky, 1989 Dalmatoreicheia Magrini & Bulirsch, 2005 Dimorphoreicheia Magrini; Fancello & Leo, 2002 Galicioreicheia Felix & Bulirsch, 2015 Guiodytes Tian, 2013 Gymnetoreicheia Magrini; Fancello & Casale, 2019 Iberodytes Jeannel, 1949 Ichnusodytes Magrini; Fancello & Onnis, 2019 Italodytes G.Müller, 1938 Kenyoreicheia Bulirsch & Magrini, 2007 Laoreicheia Balkenohl, 2005 Madagascareicheia Magrini & Bulirsch, 2009 Malagasyreicheia Magrini; Bulirsch & Fancello, 2021 Orientoreicheia Bulirsch & Hurka, 1994 Oxydrepanus Putzeys, 1867 Parareicheia Jeannel, 1957 Reicheadella Reitter, 1913 Reicheia Saulcy, 1862 Reicheidius Jeannel, 1957 Sikelioreicheia Magrini & Fancello, 2019 Spelaeodytes L.Miller, 1863 Typhloreicheia Holdhaus, 1924 Subtribe Schizogeniina Dostal, 2017 Baehrogenius Dostal, 2017 Coryza Putzeys, 1867 Halocoryza Alluaud, 1919 Lophocoryza Alluaud, 1941 Paracoryza Basilewsky, 1952 Psammocoryza Hogan, 2006 Schizogenius Putzeys, 1846 Subtribe Sparostesina Dostal, 2017 Bohemaniella Bousquet, 2002 Pseudoclivina Kult, 1947 Sparostes Putzeys, 1867 Tribe Corintascarini Basilewsky, 1973 Corintascaris Basilewsky, 1952 Tribe Dyschiriini Kolbe, 1880 Akephorus LeConte, 1852 Caledyschirius Bulirsch, 2010 Clivinopsis Bedel, 1895 Cribrodyschirius Bruneau de Miré, 1952 Dyschirius Bonelli, 1810 Neodyschirius Kult, 1954 Reicheiodes Ganglbauer, 1891 Setodyschirius Fedorenko, 1996 Striganoviella Fedorenko, 2012 Torretassoa Schatzmayr & Koch, 1933 †Dyschiriomimus Iablokoff-Khnzorian, 1960 Tribe Salcediini Alluaud, 1930 Subtribe Salcediina Alluaud, 1930 Salcedia Fairmaire, 1899 Subtribe Solenogenyina R.T.Bell, 1998 Holoprizus Putzeys, 1867 Solenogenys Westwood, 1859 Tribe Scaritini Bonelli, 1810 Subtribe Carenina W.J.MacLeay, 1887 Carenidium Chaudoir, 1868 Carenum Bonelli, 1813 Epilectus Blackburn, 1888 Euryscaphus W.J.MacLeay, 1865 Laccopterum W.J.MacLeay, 1878 Monocentrum Chaudoir, 1868 Mouhotia Laporte, 1862 Neocarenum Laporte, 1867 Neoscaphus Sloane, 1888 Philoscaphus W.J.MacLeay, 1871 Scaraphites Westwood, 1842 Trichocarenum Blackburn, 1892 Subtribe Pasimachina Putzeys, 1867 Pasimachus Bonelli, 1813 Subtribe Scapterina Putzeys, 1867 Acanthoscelis Dejean, 1825 Oxylobus Chaudoir, 1855 Parathlibops Basilewsky, 1958 Passalidius Chaudoir, 1863 Scapterus Dejean, 1826 Steganomma W.J.MacLeay, 1887 Thlibops Putzeys, 1867 Subtribe Scaritina Bonelli, 1810 Anomophaenus Fauvel, 1882 Antilliscaris Lorenz, 1998 Baenningeria Reichardt, 1976 Coptolobus Chaudoir, 1857 Crepidopterus Chaudoir, 1855 Cryptoscaphus Chaudoir, 1855 Dinoscaris Alluaud, 1902 Distichus Motschulsky, 1858 Dyscaris Bänninger, 1940 Dyscherinus Jeannel, 1955 Dyscherus Chaudoir, 1855 Geoscaptus Chaudoir, 1855 Glyptogrus Bates, 1875 Gnaphon Andrewes, 1920 Haplogaster Chaudoir, 1879 Haplotrachelus Chaudoir, 1855 Macromorphus Chaudoir, 1857 Madascaris Bänninger, 1938 Mamboicus Bates, 1886 Mecynoscaris Alluaud, 1930 Menigius Chaudoir, 1879 Neochryopus Bänninger, 1932 Ochyropus Schiödte, 1847 Pachyodontus Chaudoir, 1879 Paradyscherus Basilewsky, 1971 Pilades Heyne, 1895 Prodyscherodes Jeannel, 1955 Prodyscherus Jeannel, 1946 Scarites Fabricius, 1775 Storthodontus Chaudoir, 1855 Tapinoscaris Jeannel, 1946 Tibioscarites Bänninger, 1929 Tonkinoscaris Bänninger, 1956 Typhloscaris Kuntzen, 1914 Subfamily Siagoninae Bonelli, 1813 edit Tribe Enceladini G.Horn, 1881 Enceladus Bonelli, 1813 Tribe Siagonini Bonelli, 1813 Luperca Laporte, 1840 Siagona Latreille, 1804 Subfamily Trechinae Bonelli, 1810 edit Tribe Bembidarenini Maddison et al., 2019 Andinodontis Erwin & Toledano, 2010 Argentinatachoides Sallanave; Erwin & Roig-Juñent, 2008 Bembidarenas Erwin, 1972 Tasmanitachoides Erwin, 1972 Tribe Bembidiini Stephens, 1827 Subtribe Anillina Jeannel, 1937 Afranillus Giachino, 2015 Afrodipnus Giachino, 2015 Angustanillus Baehr & Main, 2016 Anillaspis Casey, 1918 Anillinus Casey, 1918 Anillodes Jeannel, 1963 Anillopsidius Coiffait, 1969 Anillopsis Jeannel, 1937 Anillotarsus Mateu, 1980 Anillus Jacquelin du Val, 1851 Argiloborus Jeannel, 1937 Austranillus Giachino, 2005 Bafutyphlus Bruneau de Miré, 1986 Bhutanillus M.E.Schmid, 1975 Bulirschia Giachino, 2008 Bylibaraphanus Giachino; Eberhard & Perina, 2021 Caeconannus Jeannel, 1963 Caecoparvus Jeannel, 1937 Carayonites Bruneau de Miré, 1986 Cryptocharidius M.Etonti & Mateu, 1992 Cryptorites Jeannel, 1950 Dicropterus Ehlers, 1883 Elgonotyphlus Sciaky & Zaballos, 1993 Erwinanillus Giachino; Eberhard & Perina, 2021 Externanillus Baehr & Main, 2016 Geocharidius Jeannel, 1963 Geocharis Ehlers, 1883 Gilesdytes Giachino; Eberhard & Perina, 2021 Gracilanillus Baehr & Main, 2016 Gregorydytes Giachino; Eberhard & Perina, 2021 Hesperanillus Baehr & Main, 2016 Honduranillus Zaballos, 1997 Hygranillus B.Moore, 1980 Hypodipnites Jeannel, 1963 Hypotyphlus Jeannel, 1937 Iason Giachino & Vailati, 2011 Iberanillus Español, 1971 Illaphanus W.J.MacLeay, 1865 Kimberleytyphlus Giachino; Eberhard & Perina, 2021 Leleupanillus Basilewsky, 1976 Magnanillus Baehr, 2017 Malagasydipnus Giachino, 2008 Malagasytyphlus Giachino, 2008 Megastylulus Giachino & Sciaky, 2003 Mexanillus Vigna Taglianti, 1973 Microdipnites Jeannel, 1957 Microdipnodes Basilewsky, 1960 Microdipnus Jeannel, 1937 Microtyphlus Linder, 1863 Mystroceridius Reichardt, 1972 Neodipnus Jeannel, 1957 Neoillaphanus Giachino; Eberhard & Perina, 2021 Nesamblyops Jeannel, 1937 Nothanillus Jeannel, 1962 Orthotyphlus Mateu & Zaballos, 1998 Paranillopsis Cicchino & Roig-Juñent, 2001 Paranillus Jeannel, 1949 Parillaphanus Baehr, 2018 Parvocaecus Coiffait, 1956 Pelocharis Jeannel, 1960 Pelodiaetodes B.Moore, 1980 Pelodiaetus Jeannel, 1937 Pelonomites Jeannel, 1963 Perucharidius Mateu & M.Etonti, 2002 Pilbaradytes Giachino; Eberhard & Perina, 2021 Pilbaranillus Baehr & Main, 2016 Pilbaraphanus Giachino; Eberhard & Perina, 2021 Prioniomus Jeannel, 1937 Pseudanillus Bedel, 1896 Pseudillaphanus Giachino, 2005 Rhegmatobius Jeannel, 1937 Scotodipnus Schaum, 1860 Selenodipnus Jeannel, 1963 Serranillus Barr, 1995 Serratotyphlus Giachino, 2008 Stylulus L.Schaufuss, 1882 Tasmanillus Giachino, 2005 Typhlocharis Dieck, 1869 Typhlonesiotes Jeannel, 1937 Winklerites Jeannel, 1937 Zapotecanillus Sokolov, 2013 Zeanillus Jeannel, 1937 Zoianillus Sciaky, 1994 Subtribe Bembidiina Stephens, 1827 Amerizus Chaudoir, 1868 Asaphidion Gozis, 1886 Bembidion Latreille, 1802 Caecidium Ueno, 1971 Lionepha Casey, 1918 Ocys Stephens, 1828 Orzolina Machado, 1987 Sinechostictus Motschulsky, 1864 Subtribe Horologionina Jeannel, 1949 Horologion Valentine, 1932 Subtribe Lovriciina Giachino; B.Gueorguiev & Vailati, 2011 Lovricia Pretner, 1979 Neolovricia Lakota; Jalzic & J.Moravec, 2009 Paralovricia Giachino; B.Gueorguiev & Vailati, 2011 Subtribe Tachyina Motschulsky, 1862 Aenigmatachys Baehr, 2020 Afrotachys Basilewsky, 1958 Anomotachys Jeannel, 1946 Costitachys Erwin, 1974 Elaphropus Motschulsky, 1839 Kiwitachys Larochelle & Larivière, 2007 Liotachys Bates, 1871 Lymnastis Motschulsky, 1862 Meotachys Erwin, 1974 Micratopus Casey, 1914 Moirainpa Erwin, 1984 Nothoderis Boyd & Erwin, 2016 Pericompsus LeConte, 1852 Polyderidius Jeannel, 1962 Porotachys Netolitzky, 1914 Setitachys Baehr, 2016 Stigmatachys Boyd & Erwin, 2016 Straneoites Basilewsky, 1947 Tachys Dejean, 1821 Tachysbembix Erwin, 2004 Tachyta Kirby, 1837 Tachyxysta Boyd & Erwin, 2016 Subtribe Xystosomina Erwin, 1994 Erwiniana Paulsen & A.Smith, 2003 Geballusa Erwin, 1994 Gouleta Erwin, 1994 Inpa Erwin, 1978 Mioptachys Bates, 1882 Philipis Erwin, 1994 Xystosomus Schaum, 1863 Tribe Pogonini Laporte, 1834 Bedeliolus Semenov, 1900 Cardiaderus Dejean, 1828 Diodercarus Lutshnik, 1931 Diplochaetus Chaudoir, 1872 Ochtozetus Chaudoir, 1872 Olegius Komarov, 1996 Pogonistes Chaudoir, 1872 Pogonopsis Bedel, 1898 Pogonus Dejean, 1821 Sirdenus Dejean, 1828 Syrdenoidius Baehr & Hudson, 2001 Thalassotrechus Van Dyke, 1918 Tribe Sinozolini Deuve, 1997 Chaltenia Roig-Juñent & Cicchino, 2001 Phrypeus Casey, 1924 Sinozolus Deuve, 1997 Tribe Trechini Bonelli, 1810 Subtribe Aepina Fowler, 1887 Aepopsis Jeannel, 1922 Aepus Leach in Samouelle, 1819 Kenodactylus Broun, 1909 Kiwitrechus Larochelle & Larivière, 2007 Maoritrechus Brookes, 1932 Neanops Britton, 1962 Oarotrechus Townsend, 2010 Temnostega Enderlein, 1905 Thalassobius Solier, 1849 Waiputrechus Townsend, 2010 Subtribe Perileptina Sloane, 1903 Apoplotrechus Alluaud, 1915 Neoblemus Jeannel, 1923 Perileptus Schaum, 1860 Subtribe Trechina Bonelli, 1810 Accoella Ueno, 1990 Acheroniotes Lohaj & Lakota, 2010 Adriaphaenops Noesske, 1928 Aepiblemus Belousov & Kabak, 1993 Agonotrechus Jeannel, 1923 Agostinia Jeannel, 1928 Alanorites Belousov, 1998 Albanotrechus Casale & V.B.Gueorguiev, 1994 Allegrettia Jeannel, 1928 Allotrechiama Ueno, 1970 Ameroduvalius Valentine, 1952 Anchotrechus Jeannel, 1927 Andinorites Mateu & Belles, 1980 Andinotrechus Mateu, 1981 Anillidius Jeannel, 1928 Anophthalmus Sturm, 1844 Aphaenopidius G.Müller, 1913 Aphaenops Bonvouloir, 1862 Aphaenopsis G.Müller, 1913 Apocimmerites Belousov, 1998 Apoduvalius Jeannel, 1953 Aputrechisibus Trezzi, 2007 Arctaphaenops Meixner, 1925 Aspidaphaenops Ueno, 2006 Austrotrechus B.Moore, 1972 Awatrechus Ueno, 1955 Balazucellus Deuve, 2001 Bathytrechus Ueno, 2005 Beronaphaenops B.V.Gueorguiev, 2012 Birmaphaenops Deuve, 2017 Blemus Dejean, 1821 Boldoriella Jeannel, 1928 Boreaphaenops Ueno, 2002 Bothynotrechus B.Moore, 1972 Casaleaphaenops Tian; Huang & Ma, 2021 Cathaiaphaenops Deuve, 1996 Caucasaphaenops Belousov, 1999 Caucasorites Belousov & Zamotajlov, 1997 Chaetoduvalius Jeannel, 1928 Chaetotrechiama Ueno, 1982 Chiapadytes Vigna Taglianti, 1977 Chu Tian & He, 2020 Cimbrodytes Piva, 2021 Cimmerites Jeannel, 1928 Cimmeritodes Deuve, 1996 Ckacus Tian & Huang, 2021 Columbitrechus Mateu, 1982 Coreoblemus Ueno, 1969 Croatotrechus Casale & Jalzic, 1999 Dabatrechus Ueno, 2004 Dactylotrechus Belousov & Kabak, 2003 Daiconotrechus Ueno, 1971 Dalmataphaenops Monguzzi, 1993 Darlingtonea Valentine, 1952 Derossiella Queinnec, 2008 Deuveaphaenops Tian & Huang, 2017 Dianotrechus Tian, 2016 Doderotrechus Vigna Taglianti, 1968 Dongoblemus Deuve & Tian, 2016 Dongodytes Deuve, 1993 Dracotrechus Ueno, 2010 Duvalioblemus Deuve, 1995 Duvaliomimus Jeannel, 1928 Duvaliopsis Jeannel, 1928 Duvalius Delarouzée, 1859 Eocnides Jeannel, 1954 Epaphiopsis Ueno, 1953 Epaphiotrechus Deuve & Kavanaugh, 2016 Epaphius Leach in Samouelle, 1819 Erebotrechus Britton, 1964 Escolatrechus Mateu, 2003 Eutrechopsis B.Moore, 1972 Eutrechus B.Moore, 1972 Geotrechus Jeannel, 1919 Giraffaphaenops Deuve, 2002 Glabroduvalius Vrbica; Curcic; Antic & Curcic, 2013 Goedetrechus B.Moore, 1972 Gotoblemus Ueno, 1970 Graciliblemus Deuve & Tian, 2016 Guatemalatrechus Trezzi, 2003 Guiaphaenops Deuve, 2002 Guizhaphaenops Vigna Taglianti, 1997 Guizhaphaenopsoides Tian; Huang & Li, 2021 Gulaphaenops Ueno, 1987 Himalaphaenops Ueno, 1980 Himalotrechus Belousov; Kabak & J.Schmidt, 2019 Homaloderodes Jeannel, 1962 Huoyanodytes Tian & Huang, 2016 Iberotrechus Jeannel, 1920 Iga Ueno, 1953 Incatrechus Mateu & Belles, 1982 Inotrechus Dolzhanski & Ljovushkin, 1989 Ishidatrechus Ueno, 1956 Ishikawatrechus Habu, 1950 Italaphaenops Ghidini, 1964 Iyotrechus Ueno & Naito, 2009 Jalzicaphaenops Lohaj & Lakota, 2010 Jeannelius Kurnakov, 1959 Jiangxiaphaenops Ueno & Clarke, 2007 Jiulongotrechus Tian; Huang & Wang, 2015 Junaphaenops Ueno, 1997 Junnanotrechus Ueno & Yin, 1993 Kettlotrechus Townsend, 2010 Kosswigia Jeannel, 1947 Kozlovites Jeannel, 1935 Kupetrechus Larochelle & Larivière, 2007 Kurasawatrechus Yoshida & Nomura, 1952 Kusumia Ueno, 1952 Lanxangaphaenops Deuve, 2012 Laoblemus Ueno, 2006 Laosaphaenops Deuve, 2000 Lessinodytes Vigna Taglianti, 1982 Libotrechus Ueno, 1998 Luoxiaotrechus Tian & Yin, 2013 Luyatrechus M.Etonti & Mateu, 2000 Luzonotrechus Ueno, 1979 Mamesdytes Trezzi, 2003 Masuzoa Ueno, 1960 Masuzonoblemus Ueno, 1989 Mayaphaenops Vigna Taglianti, 1977 Meganophthalmus Kurnakov, 1959 Mexaphaenops Bolivar y Pieltain, 1942 Mexitrechus Barr, 1982 Miaotrechus Tian; Chen & Ma, 2020 Microblemus Ueno, 2007 Mimanillus B.Moore, 1972 Mimotrechus B.Moore, 1972 Minimaphaenops Deuve, 2000 Minosaphaenops Queinnec, 2008 Minutotrechus Deuve & Kavanaugh, 2016 Nannotrechus Winkler, 1926 Nanotrechiama Belousov & Kabak, 2018 Neaphaenops Jeannel, 1920 Nelsonites Valentine, 1952 Neotrechus G.Müller, 1913 Nesaeoduvalius Casale & Giachino, 2016 Nesiotrechus Ueno, 1995 Nipponaphaenops Ueno, 1971 Nototrechus B.Moore, 1972 Nunbergites Pawlowski & Stachowiak, 1991 Omalodera Blanchard, 1842 Oodinotrechus Ueno, 1998 Oroblemites Ueno & Pawlowski, 1981 Oroblemus Ueno & Yoshida, 1966 Orotrechus G.Müller, 1913 Oxytrechus Jeannel, 1927 Panaphaenops Tian; Huang & Ma, 2021 Paracimmerites Belousov, 1998 Paraphaenops Jeannel, 1916 Paratrechiotes Ueno, 1995 Paratrechus Jeannel, 1920 Petraphaenops Delic; Kapla & Colla, 2019 Pheggomisetes Knirsch, 1923 Pilosaphaenops Deuve & Tian, 2008 Plesioaphaenops Deuve & Tian, 2011 Pogonoschema Jeannel, 1927 Pontodytes Casale & Giachino, 1989 Porocimmerites Belousov, 1998 Protrechiama Belousov & Kabak, 2003 Pseudanophthalmus Jeannel, 1920 Pseudaphaenops Winkler, 1912 Pseudocnides Jeannel, 1927 Pseudotrechisibus Mateu & Belles, 1982 Puertrechus Belousov & Kabak, 2014 Putzeysius Jeannel, 1962 Qianaphaenops Ueno, 2000 Qianotrechus Ueno, 2000 Queinnectrechus Deuve, 1992 Rakantrechus Ueno, 1951 Ryugadous Habu, 1950 Sardaphaenops Cerruti & Henrot, 1956 Satotrechus Ueno, 2006 Sbordoniella Vigna Taglianti, 1980 Scotoplanetes Absolon, 1913 Scototrechus Britton, 1962 Shenaphaenops Ueno, 1999 Shennongotrechus Ueno, 2004 Shenoblemus Tian & Fang, 2020 Shilinotrechus Ueno, 2003 Shiqianaphaenops Tian, 2016 Shuangheaphaenops Tian, 2017 Shuaphaenops Ueno, 1999 Sichuanotrechus Deuve, 2005 Sidublemus Tian & Yin, 2013 Sinaphaenops Ueno & Wang, 1991 Sinocimmerites Deuve, 2007 Sinotrechiama Ueno, 2000 Sinotroglodytes Deuve, 1996 Sloanella Jeannel, 1927 Speotrechus Jeannel, 1922 Stevensius Jeannel, 1923 Stygiotrechus Ueno, 1958 Subilsia Español, 1967 Sumatrechus Deuve, 2005 Superbotrechus Deuve & Tian, 2009 Suzuka Ueno, 1956 Taiwanotrechus Ueno, 1987 Taniatrechus Belousov & Dolzhanski, 1994 Tasmanorites Jeannel, 1927 Tasmanotrechus B.Moore, 1972 Taurocimmerites Belousov, 1998 Thalassoduvalius Ueno, 1956 Tianeotrechus Tian & Tang, 2016 Tianzhuaphaenops Zhao & Tian, 2016 Tienmutrechus Suenson, 1957 Tonkinaphaenops Deuve, 2013 Toshiaphaenops Ueno, 1999 Trechepaphiama Deuve & Kavanaugh, 2016 Trechepaphiopsis Deuve & Kavanaugh, 2016 Trechiama Jeannel, 1927 Trechiamiotes Deuve, 1998 Trechicomimus Mateu & Nègre, 1972 Trechiella Jeannel, 1927 Trechimorphus Jeannel, 1927 Trechinotus Jeannel, 1962 Trechiotes Jeannel, 1954 Trechisibus Motschulsky, 1862 Trechistus B.Moore, 1972 Trechoblemus Ganglbauer, 1891 Trechus Clairville, 1806 Trichaphaenops Jeannel, 1916 Troglocimmerites Ljovuschkin, 1970 Tropidotrechus Jeannel, 1927 Typhlotrechus G.Müller, 1913 Uenoaphaenops Tian & He, 2020 Uenoites Belousov & Kabak, 2016 Uenotrechus Deuve & Tian, 1999 Ushijimaella Ueno, 1980 Velebitaphaenops Casale & Jalzic, 2012 Velesaphaenops S.Curcic et al., 2018 Vietotrechus Ueno, 1995 Wanhuaphaenops Tian & Wang, 2016 Wanoblemus Tian & Fang, 2016 Wulongoblemus Ueno, 2007 Xenotrechus Barr & Krekeler, 1967 Xiangxius Tian; Huang & Li, 2021 Xuedytes Tian & Huang, 2017 Yalongaphaenops Belousov & Kabak, 2021 Yamautidius Ueno, 1957 Yanzaphaenops Ueno, 2010 Yunotrechus Tian & Huang, 2014 Zhijinaphaenops Ueno & Ran, 2002 Subtribe Trechodina Jeannel, 1926 Amblystogenium Enderlein, 1905 Canarobius Machado, 1987 Cnides Motschulsky, 1862 Cothresia Jeannel, 1964 Cyphotrechodes Jeannel, 1926 Eotrechodes Ueno; Lafer & Sundukov, 1995 Himalotrechodes Ueno, 1981 Iberotrechodes Faille et al., 2021 Pachydesus Motschulsky, 1864 Pachytrechodes Jeannel, 1960 Paratrechodes Jeannel, 1926 Scaurotrechodes Geginat, 2006 Spelaeovulcania Machado, 1987 Sporades Fauvel, 1882 Thalassophilus Wollaston, 1854 Trechobembix Jeannel, 1926 Trechodes Blackburn, 1901 Trechosia Jeannel, 1926 Trechosiella Jeannel, 1960 †Antarctotrechus Ashworth & Erwin, 2016 †Trechinites Heer, 1862 †Trechoides Motschulsky, 1856 Tribe Zolini Sharp, 1886 Subtribe Merizodontina Sloane, 1920 Idacarabus Lea, 1910 Maungazolus Larochelle & Larivière, 2017 Merizodus Solier, 1849 Percodermus Sloane, 1920 Pseudoopterus Csiki, 1928 Pterocyrtus Sloane, 1920 Sloaneana Csiki, 1933 Synteratus Broun, 1909 Thayerella Baehr, 2016 Subtribe Zolina Sharp, 1886 Oopterus Guérin-Méneville, 1841 Zolus Sharp, 1886 Xenaroswelliana Erwin, 2007 Subfamily †Conjunctiinae Ponomarenko, 1977 edit †Conjunctia Ponomarenko, 1977 Subfamily †Protorabinae Ponomarenko, 1977 edit †Aethocarabus Ren; Lu & Guo, 1995 †Atrirabus Hong & Wang, 1990 †Cordorabus Ponomarenko, 1977 †Cretorabus Ponomarenko, 1977 †Denudirabus Ren; Lu & Guo, 1995 †Lirabus Hong, 1992 †Lithorabus Ponomarenko, 1977 †Magnirabus Hong & Wang, 1990 †Mesorabus Ponomarenko, 1977 †Nebrorabus Ponomarenko, 1989 †Ovrabites Ponomarenko, 1977 †Penecupes Ren, 1995 †Protorabus Ponomarenko, 1977 Not Assigned to Subfamily †Agatoides Motschulsky, 1856 †Amphoxyne Bode, 1953 †Carabites Heer, 1852 †Cavicarabus Hong, 1991 †Conexicoxa Lin, 1986 †Cymatopterus Lomnicki, 1894 †Fangshania Hong, 1981 †Glenopterus Heer, 1847 †Hebeicarabus Hong, 1983 †Megacarabus Hong, 1983 †Meileyingia Hong, 1987 †Miocarabus Hong, 1983 †Neothanes Scudder, 1890 †Procarabus Oppenheim, 1888 †Prosynactus Bode, 1953 †Shanwangicarabus Hong, 1985 †Sinis Heer, 1862 †Sinocalosoma Hong & Wang, 1986 †Sinocaralosoma Hong, 1984 †Sunocarabus Hong, 1987 †Tauredon Handlirsch, 1910 †Wuchangicarabus Hong, 1991 †Xishanocarabus Hong, 1984 †Yunnanocarabus Lin, 1977 References edit ^ Lorenz, Wolfgang (2021). "Carabcat Database". doi :10.48580/dfqf-3dk . Retrieved 2023-03-04 . ^ "Carabidae Latreille, 1802". Catalogue of Life . Retrieved 2023-03-04 .
list, ground, beetle, genera, this, page, lists, genera, family, carabidae, according, catalogue, life, carabcat, database, march, 2023, excluding, tiger, beetles, which, considered, family, cicindelidae, contents, family, carabidae, subfamily, anthiinae, bone. This page lists the genera in the family Carabidae according to Catalogue of Life and Carabcat Database as of March 2023 excluding Tiger Beetles which are considered to be in the family Cicindelidae 1 2 Contents 1 Family Carabidae 1 1 Subfamily Anthiinae Bonelli 1813 1 2 Subfamily Apotominae LeConte 1853 1 3 Subfamily Brachininae Bonelli 1810 1 4 Subfamily Broscinae Hope 1838 1 5 Subfamily Carabinae Linnaeus 1802 1 6 Subfamily Ctenodactylinae Laporte 1834 1 7 Subfamily Dryptinae Bonelli 1810 1 8 Subfamily Elaphrinae Latreille 1802 1 9 Subfamily Gineminae Ball amp Shpeley 2002 1 10 Subfamily Harpalinae Bonelli 1810 1 11 Subfamily Hiletinae Schiodte 1847 1 12 Subfamily Lebiinae Bonelli 1810 1 13 Subfamily Licininae Bonelli 1810 1 14 Subfamily Loricerinae Bonelli 1810 1 15 Subfamily Melaeninae Csiki 1933 1 16 Subfamily Migadopinae Chaudoir 1861 1 17 Subfamily Nebriinae Laporte 1834 1 18 Subfamily Nototylinae Banninger 1927 1 19 Subfamily Omophroninae Bonelli 1810 1 20 Subfamily Orthogoniinae Schaum 1857 1 21 Subfamily Panagaeinae Bonelli 1810 1 22 Subfamily Patrobinae Kirby 1837 1 23 Subfamily Paussinae Latreille 1806 1 24 Subfamily Platyninae Bonelli 1810 1 25 Subfamily Promecognathinae LeConte 1853 1 26 Subfamily Pseudomorphinae Hope 1838 1 27 Subfamily Psydrinae LeConte 1853 1 28 Subfamily Pterostichinae Bonelli 1810 1 29 Subfamily Rhysodinae Laporte 1840 1 30 Subfamily Scaritinae Bonelli 1810 1 31 Subfamily Siagoninae Bonelli 1813 1 32 Subfamily Trechinae Bonelli 1810 1 33 Subfamily Xenaroswellianinae Erwin 2007 1 34 Subfamily Conjunctiinae Ponomarenko 1977 1 35 Subfamily Protorabinae Ponomarenko 1977 2 ReferencesFamily Carabidae editSubfamily Anthiinae Bonelli 1813 edit Tribe Anthiini Bonelli 1813Anthia Weber 1801 Atractonotus Perroud 1847 Baeoglossa Chaudoir 1850 Cycloloba Chaudoir 1850 Cypholoba Chaudoir 1850 Eccoptoptera Chaudoir 1878 Gonogenia Chaudoir 1844 Netrodera Chaudoir 1850 dd Tribe Helluonini Hope 1838Subtribe Helluonina Hope 1838Aenigma Newman 1836 Ametroglossus Sloane 1914 Dicranoglossus Chaudoir 1872 Epimicodema Sloane 1914 Gigadema J Thomson 1859 Helluapterus Sloane 1914 Helluarchus Sloane 1914 Helluo Bonelli 1813 Helluodema Laporte 1867 Helluonidius Chaudoir 1872 Helluopapua Darlington 1968 Helluosoma Laporte 1867 Neohelluo Sloane 1914 Platyhelluo Baehr 2005 dd Subtribe Omphrina Jedlicka 1941Colfax Andrewes 1920 Creagris Nietner 1857 Dailodontus Reiche 1843 Erephognathus Alluaud 1932 Helluobrochus Reichardt 1974 Helluomorpha Laporte 1834 Helluomorphoides Ball 1951 Macrocheilus Hope 1838 Meladroma Motschulsky 1855 Omphra Dejean 1825 Pleuracanthus Gray 1832 Triaenogenius Chaudoir 1877 dd dd Tribe Physocrotaphini Chaudoir 1863Anguloderus Anichtchenko amp Sciaky 2017 Foveocrotaphus Anichtchenko 2014 Helluodes Westwood 1847 Holoponerus Fairmaire 1883 Physocrotaphus Parry 1849 Physoglossus Akhil amp Sabu 2020 Pogonoglossus Chaudoir 1863 Schuelea Baehr 2004 dd Subfamily Apotominae LeConte 1853 edit Apotomus Illiger 1807Subfamily Brachininae Bonelli 1810 edit Tribe Brachinini Bonelli 1810Aptinoderus Hubenthal 1919 Aptinus Bonelli 1810 Brachinulus Basilewsky 1958 Brachinus Weber 1801 Mastax Fischer von Waldheim 1828 Pheropsophus Solier 1833 Styphlodromus Basilewsky 1959 Styphlomerus Chaudoir in Putzeys 1875 dd Tribe Crepidogastrini Jeannel 1949Brachynillus Reitter 1904 Crepidogaster Boheman 1848 Crepidogastrillus Basilewsky 1959 Crepidogastrinus Basilewsky 1957 Crepidolomus Basilewsky 1959 Crepidonellus Basilewsky 1959 dd Subfamily Broscinae Hope 1838 edit Tribe Broscini Hope 1838Subtribe Axonyina Roig Junent 2000Broscodes Bolivar y Pieltain 1914 Rawlinsius Davidson amp Ball 1998 dd Subtribe Baripodina Jeannel 1941Baripus Dejean 1828 Bembidiomorphum Champion 1918 dd Subtribe Broscina Hope 1838Broscodera Lindroth 1961 Broscosoma Rosenhauer 1846 Broscus Panzer 1813 Chaetobroscus Semenov 1900 Craspedonotus Schaum 1863 Eobroscus Kryzhanovskij 1951 Kashmirobroscus J Schmidt Wrase amp Sciaky 2013 Miscodera Eschscholtz 1830 Zacotus LeConte 1869 dd Subtribe Creobiina Jeannel 1941Acallistus Sharp 1886 Adotela Laporte 1867 Anheterus Putzeys 1868 Bountya Townsend 1971 Brithysternum W J MacLeay 1873 Cascellius Curtis 1838 Cerotalis Laporte 1867 Creobius Guerin Meneville 1838 Gnathoxys Westwood 1842 Nothocascellius Roig Junent 1995 Promecoderus Dejean 1829 dd Subtribe Nothobroscina Roig Junent 2000Chylnus Sloane 1920 Diglymma Sharp 1886 Eurylychnus Bates 1891 Mecodema Blanchard 1843 Monteremita Seldon amp Holwell 2019 Nothobroscus Roig Junent amp Ball 1995 Oregus Putzeys 1868 Orthoglymma Liebherr Marris Emberson Syrett amp Roig Junent 2011 Percolestus Sloane 1892 Percosoma Schaum 1858 dd dd Subfamily Carabinae Linnaeus 1802 edit Tribe Carabini Linnaeus 1802Subtribe Carabina Linnaeus 1802Aplothorax G R Waterhouse 1842 Calosoma Weber 1801 Carabus Linnaeus 1758 dd Subtribe Ceroglossina Lapouge 1927Ceroglossus Solier 1848 dd dd Tribe Cychrini Perty 1830Subtribe Cychrina Perty 1830Cychropsis Boileau 1901 Cychrus Fabricius 1794 Scaphinotus Dejean 1826 Sphaeroderus Dejean 1826 dd Subtribe Pamborina Hope 1838Maoripamborus Brookes 1944 Pamborus Latreille 1812 dd dd Subfamily Ctenodactylinae Laporte 1834 edit Tribe Ctenodactylini Laporte 1834Alachnothorax Liebke 1929 Amblycoleus Chaudoir 1872 Antipionycha Liebke 1928 Askalaphium Liebke 1938 Ctenodactyla Dejean 1825 Leptotrachelon Liebke 1928 Leptotrachelus Latreille 1829 Oilea Liebke 1931 Parapionycha Liebke 1929 Pionycha Chaudoir 1848 Plagiotelum Solier 1849 Propionycha Liebke 1928 Pseudometabletus Liebke 1930 Schidonychus Klug 1834 Teukrus Liebke 1931 Wate Liebke 1928 dd Tribe Hexagoniini G Horn 1881Dinopelma Bates 1889 Hexagonia Kirby 1825 Omphreoides Fairmaire 1896 dd Subfamily Dryptinae Bonelli 1810 edit Tribe Dryptini Bonelli 1810Desera Hope 1831 Drypta Latreille 1797 Megadrypta Sciaky amp Anichtchenko 2020 Neodrypta Basilewsky 1960 Nesiodrypta Jeannel 1949 Prionodrypta Jeannel 1949 dd Tribe Galeritini Kirby 1825Ancystroglossus Chaudoir 1863 Eunostus Laporte 1835 Galerita Fabricius 1801 Trichognatha Latreille 1829 dd Tribe Zuphiini Bonelli 1810Subtribe Dicrodontina Machado 1992Dicrodontus Chaudoir 1872 dd Subtribe Leleupidiina Basilewsky 1951Colasidia Basilewsky 1954 Gunvorita Landin 1955 Leleupidia Basilewsky 1951 Neoleleupidia Basilewsky 1953 Paraleleupidia Basilewsky 1951 dd Subtribe Planetina Jedlicka 1941Planetes W S MacLeay 1825 dd Subtribe Zuphiina Bonelli 1810Acrogenys W J MacLeay 1864 Agastus Schmidt Goebel 1846 Chaudoirella Mateu 1982 Coarazuphium Gnaspini Vanin amp Godoy 1998 Ildobates Espanol 1966 Metaxidius Chaudoir 1852 Metazuphium Mateu 1992 Mischocephalus Chaudoir 1863 Parazuphium Jeannel 1942 Polistichus Bonelli 1810 Pseudaptinus Laporte 1834 Speothalpius B Moore 1995 Speozuphium B Moore 1995 Typhlozuphium Baehr 2014 Zuphioides Ball amp Shpeley 2013 Zuphium Latreille 1805 dd dd Subfamily Elaphrinae Latreille 1802 edit Blethisa Bonelli 1810 Diacheila Motschulsky 1844 Elaphrus Fabricius 1775 Elaphrotites Haupt 1856Subfamily Gineminae Ball amp Shpeley 2002 edit Ginema Ball amp Shpeley 2002Subfamily Harpalinae Bonelli 1810 edit Tribe Anisodactylini Lacordaire 1854Allendia Noonan 1974 Allocinopus Broun 1903 Amphasia Newman 1838 Anisodactylus Dejean 1829 Anisostichus Emden 1953 Cenogmus Sloane 1898 Chydaeus Chaudoir 1854 Crasodactylus Guerin Meneville 1847 Criniventer Emden 1953 Diachromus Erichson 1837 Dicheirus Mannerheim 1843 Gaioxenus Broun 1910 Geopinus LeConte 1847 Gnathaphanus W S MacLeay 1825 Gynandromorphus Dejean 1829 Haplaner Chaudoir 1878 Harpalomimetes Schauberger in Csiki 1932 Hypharpax W S MacLeay 1825 Maoriharpalus Larochelle amp Lariviere 2005 Nornalupia Kataev 2002 Notiobia Perty 1830 Parabaris Broun 1881 Progonochaetus G Muller 1938 Pseudanisotarsus Noonan 1973 Pseudognathaphanus Schauberger 1932 Pseudorhysopus Kataev amp Wrase 2001 Rhysopus Andrewes 1929 Scybalicus Schaum 1862 Triplosarus Bates 1874 Tuiharpalus Larochelle amp Lariviere 2005 Xestonotus LeConte 1853 dd Tribe Harpalini Bonelli 1810Subtribe Amblystomina Fauvel 1889Amblystomus Erichson 1837 Anomostomus LaFerte Senectere 1853 Barysomus Dejean 1829 Oosoma Nietner 1857 dd Subtribe Ditomina Bonelli 1810Bronislavia Semenov 1891 Carenochyrus Solsky 1874 Carterus Dejean 1830 Chilotomus Chaudoir 1842 Ditomus Bonelli 1810 Dixus Billberg 1820 Eocarterus Stichel 1923 Eucarterus Reitter 1900 Graniger Motschulsky 1864 Indocarterus Kataev amp Wrase 2018 Liochirus Tschitscherine 1897 Machozetus Chaudoir 1850 Odotoncarus Solier 1835 Oedesis Motschulsky 1850 Pachycarus Solier 1835 Parapenthes Kataev amp Wrase 2018 Penthus Chaudoir 1843 Phorticosomus Schaum 1863 Proditomus Schauberger 1934 Pseudaristus Reitter 1900 Tschitscherinellus Csiki 1906 dd Subtribe Harpalina Bonelli 1810Acinopus Dejean 1821 Afromizonus Basilewsky 1947 Allosiopelus N Ito 1995 Amblygnathus Dejean 1829 Anisocnemus Chaudoir 1843 Athrostictus Bates 1878 Axinotoma Dejean 1829 Aztecarpalus Ball 1970 Bleusea Bedel 1897 Boeomimetes Peringuey 1896 Bradybaenus Dejean 1829 Coleolissus Bates 1892 Cratacanthus Dejean 1829 Cratognathus Dejean 1829 Daptus Fischer von Waldheim 1823 Dioryche W S MacLeay 1825 Discoderus LeConte 1853 Dregus Motschulsky 1864 Ectinothorax Alluaud 1941 Eriophonus Tschitscherine 1901 Euryderus LeConte 1846 Geodromus Dejean 1829 Harpalinus Jeannel 1946 Harpaliscus Bates 1892 Harpalobrachys Tschitscherine 1899 Harpalodiodes Bousquet 2002 Harpalomimus Facchini amp Giachino 2020 Harpalomorphus Peringuey 1896 Harpalus Latreille 1802 Harpathaumas Basilewsky 1947 Harponixus Basilewsky 1950 Hartonymus Casey 1914 Heteracantha Brulle 1835 Hyphaereon W S MacLeay 1825 Indiophonus N Ito 1996 Kareya Andrewes 1919 Lampetes Andrewes 1940 Laparhetes Jeannel 1946 Liodaptus Bates 1889 Meroctenus Gemminger amp Harold 1868 Microderes Faldermann 1836 Neoaulacoryssus Noonan 1985 Neodiachipteryx Noonan 1985 Neohyparpalus Clarke 1981 Neophygas Noonan 1976 Nesacinopus Tschitscherine 1900 Nesarpalus Bedel 1897 Nipponoharpalus Habu 1973 Nothodaptus Maindron 1906 Oesyperus Andrewes 1923 Omostropus Peringuey 1896 Ooidius Chaudoir 1847 Ophoniscus Bates 1892 Ophonus Dejean 1821 Oxycentrus Chaudoir 1854 Panagrius Andrewes 1933 Pangus Dejean 1821 Parasiopelus Basilewsky 1946 Paraulacoryssus Shpeley Hunting amp Ball 2017 Parophonus Ganglbauer 1891 Paulianoscirtus Basilewsky 1976 Penthophonus Reitter 1900 Piosoma LeConte 1847 Platymetopsis Ball amp Maddison 1987 Platymetopus Dejean 1829 Prakasha Andrewes 1919 Pseudodiachipteryx Burgeon 1936 Pseudohyparpalus Basilewsky 1946 Pseudoselenophorus Peringuey 1896 Selenophorus Dejean 1829 Selenotichnus Kataev 1999 Siopelophonus Kataev 2015 Siopelus Murray 1859 Stenomorphus Dejean 1831 Trichopselaphus Chaudoir 1843 Trichotichnus A Morawitz 1863 Trichoxycentrus N Ito 2000 Typsiharpalus Tschitscherine 1901 dd dd Tribe Pelmatellini Bates 1882Hakaharpalus Larochelle amp Lariviere 2005 Kupeharpalus Larochelle amp Lariviere 2005 Lecanomerus Chaudoir 1850 Nemaglossa Solier 1849 Notospeophonus B Moore 1962 Pelmatellus Bates 1882 Syllectus Bates 1878 Trachysarus Reed 1874 dd Tribe Stenolophini Kirby 1837Acupalpus Latreille 1829 Agonoleptus Casey 1914 Amerinus Casey 1884 Angionychus Klug 1853 Anthracus Motschulsky 1850 Batoscelis Dejean 1836 Bradycellus Erichson 1837 Cratosoma Jeannel 1948 Cyptomicrus Vinson 1939 Dicheirotrichus Jacquelin du Val 1855 Egadyla Alluaud 1916 Euthenarus Bates 1874 Fuminoria Morita 2006 Goniocellus Casey 1914 Gugheorites Basilewsky 1951 Haplanister B Moore 1996 Hemiaulax Bates 1892 Hippoloetis Laporte 1835 Idiomelas Tschitscherine 1900 Kaffovatus Clarke 1972 Kenyacus Alluaud 1917 Kiwiharpalus Larochelle amp Lariviere 2005 Lioholus Tschitscherine 1897 Loxoncus Schmidt Goebel 1846 Pachytrachelus Chaudoir 1852 Parabradycellus N Ito 2003 Paramecus Dejean 1829 Philodes LeConte 1861 Pholeodytes Britton 1962 Pogonodaptus G Horn 1881 Polpochila Solier 1849 Psychristus Andrewes 1930 Rhabidius Basilewsky 1948 Stenolophus Dejean 1821 Uenanthracus Kasahara 1994 dd Subfamily Hiletinae Schiodte 1847 edit Eucamaragnathus Jeannel 1938 Hiletus Schiodte 1847Subfamily Lebiinae Bonelli 1810 edit Tribe Cyclosomini Laporte 1834Subtribe Corsyrina GanglbauerCorsyra Dejean 1825 Discoptera Semenov 1889 dd Subtribe Cyclosomina Laporte 1834Cyclosomus Latreille 1829 Sarothrocrepis Chaudoir 1850 Tetragonoderus Dejean 1829 dd Subtribe Graphipterina Latreille 1802Graphipterus Latreille 1802 Piezia Brulle 1834 Trichopiezia Negre 1955 dd Subtribe Masoreina Chaudoir 1871Anaulacus W S MacLeay 1825 Atlantomasoreus Mateu 1984 Leuropus Andrewes 1947 Lophidius Dejean 1831 Masoreus Dejean 1821 Mnuphorus Chaudoir 1873 Odontomasoreus Darlington 1968 Somoplatodes Basilewsky 1986 Somoplatus Dejean 1829 dd Subtribe Nemotarsina Bates 1883Nemotarsus LeConte 1853 dd dd Tribe Lachnophorini LeConte 1853Subtribe Calophaenina Jeannel 1948Calophaena Klug 1821 Calophaenoidea Liebke 1930 dd Subtribe Lachnophorina LeConte 1853Amphithasus Bates 1871 Anchonoderus Reiche 1843 Aporesthus Bates 1871 Asklepia Liebke 1938 Balligratus Moret amp Ortuno 2017 Calybe Laporte 1834 Ega Laporte 1834 Eucaerus LeConte 1853 Euphorticus G Horn 1881 Grundmannius Basilewsky 1965 Guatemalteca Erwin 2004 Lachnaces Bates 1872 Lachnophorus Dejean 1831 Peruphorticus Erwin amp Zamorano 2014 Pseudophorticus Erwin 2004 Selina Motschulsky 1858 dd dd Tribe Lebiini Bonelli 1810Subtribe Agrina Kirby 1837Abaditicus Ball amp Hilchie 1983 Abrodiella Bousquet 2002 Agra Fabricius 1801 Allardina Basilewsky 1963 Amelus Chaudoir 1873 Anomotariella Baehr 2012 Anomotarus Chaudoir 1875 Apterodromites Mateu 1976 Aspasiola Chaudoir 1877 Calleida Latreille 1824 Calleidomorpha Motschulsky 1855 Callidadelpha Steinheil 1875 Chaudoirina Mateu 1955 Cryptobatis Eschscholtz 1829 Cyanotarus Reed 1874 Cylindronotum Putzeys 1845 Demetrida White 1846 Diabaticus Bates 1878 Do Baehr 2009 Dromidea Perroud amp Montrouzier 1864 Epikastea Liebke 1935 Eujalmenus Bousquet 2002 Euplatia Chaudoir 1873 Eurycalleida Maindron 1905 Falsodromius Mateu 1976 Glycia Chaudoir 1842 Hyboptera Chaudoir 1873 Hybopteroides Erwin amp Ball 2012 Infernophilus Larson 1969 Kteatus Liebke 1936 Lipostratia Chaudoir 1873 Merizomena Chaudoir 1873 Mimodromius Chaudoir 1873 Mimophilorhizus Mateu 1993 Oechalius Liebke 1935 Ogygium Liebke 1935 Onota Chaudoir 1873 Otoglossa Chaudoir 1873 Paraglycia Bedel 1904 Phacocerus Chaudoir 1873 Philophuga Motschulsky 1859 Phloeocarabus W J MacLeay 1871 Plochionus Dejean 1821 Pontonoa Liebke 1935 Pseudotoglossa Mateu 1961 Pylartesius Liebke 1939 Rugitarus Baehr 2009 Speotarus B Moore 1964 Straneotia Mateu 1961 Tecnophilus Chaudoir 1877 Teiresia Liebke 1935 Thoasia Liebke 1939 Titaresius Liebke 1935 Trigonothops W J MacLeay 1864 Valeriaaschero Erwin 2004 Vianasia Mateu 1955 dd Subtribe Apenina BallApenes LeConte 1851 Cymindoidea Laporte 1833 Habutarus Ball amp Hilchie 1983 Platytarus Fairmaire 1850 Trymosternus Chaudoir 1873 dd Subtribe Celaenephina Habu 1982Celaenephes Schmidt Goebel 1846 dd Subtribe Cymindidina Laporte 1834Afrotarus Jeannel 1949 Assadecma Basilewsky 1982 Cymindis Latreille 1805 Hystrichopus Boheman 1848 Leptosarcus Peringuey 1896 Metaxymorphus Chaudoir 1850 Petrimagnia Kryzhanovskij amp Mikhailov 1971 Pinacodera Schaum 1857 Plagiopyga Boheman 1848 Pseudomasoreus Desbrochers des Loges 1904 Taridius Chaudoir 1875 dd Subtribe Demetriadina Bates 1886Cylindrocranius Chaudoir 1878 Demetrias Bonelli 1810 dd Subtribe Dromiusina Bonelli 1810Afrodromius Basilewsky 1958 Axinopalpus LeConte 1846 Barrymooreana Baehr 1997 Brachynopterus Bedel 1898 Brigalowia Baehr 2006 Calodromius Reitter 1905 Carbonellia Mateu 1968 Disciferella Kataev amp Muilwijk 2020 Dromiops Peringuey 1898 Dromius Bonelli 1810 Dromoceryx Schmidt Goebel 1846 Geoffreyella Baehr 2012 Klepteromimus Peringuey 1898 Mesolestes Schatzmayr 1943 Metadromius Bedel 1907 Microdaccus Schaum in Baudi di Selve 1864 Microlestes Schmidt Goebel 1846 Microlestodes Baehr 1987 Monnea Mateu 1970 Negrea Mateu 1968 Omophagus Andrewes 1937 Oxoides Solier 1849 Paradromius Fowler 1887 Philorhizus Hope 1838 Polyaulacus Chaudoir 1878 Psammodromius Peyerimhoff 1927 Pseudomonnea Mateu 1983 Somalodromius Mateu 1967 Xenodromius Bates 1891 Zolotarevskyella Mateu 1953 dd Subtribe Gallerucidiina Chaudoir 1872Gallerucidia Chaudoir 1872 Lebidia A Morawitz 1862 dd Subtribe Lebiina Bonelli 1810Alkestis Liebke 1939 Aristolebia Bates 1892 Daer Semenov amp Znojko 1929 Lachnolebia Maindron 1905 Lebia Latreille 1802 Lebidema Motschulsky 1864 Lebiomorphica Lorenz 1998 Lebistina Motschulsky 1864 Lebistinida Peringuey 1898 Lionedya Chaudoir 1871 Matabele Peringuey 1896 Megalebia Mateu 1972 Orthobasis Chaudoir 1871 Pachylebia Jeannel 1949 Pachylebiodes Mateu 1972 Paulianolebia Mateu 1972 Pseudopachylebia Mateu 1972 Rhopalostyla Chaudoir 1850 Scalidion Schmidt Goebel 1846 Setolebia Jedlicka 1941 Sofota Jedlicka 1951 dd Subtribe Lichnasthenina J ThomsonAstastus Peringuey 1896 Australovelinda Baehr 2012 Imasakaia Arai amp Morita 2010 Lichnasthenus J Thomson 1858 Mimovelindopsis Mateu 1963 Velinda Andrewes 1921 Velindastus Schule amp Lorenz 2008 Velindomimus Jeannel 1955 Velindopsis Burgeon 1937 dd Subtribe Lionychina Jeannel 1948Apristomimus Mateu 1969 Apristus Chaudoir 1846 Eremolestes Maindron 1905 Lionychus Wissmann 1846 Lorestania Anichtchenko 2011 Metablus Jedlicka 1958 Omobrus Andrewes 1930 Singiliomimus Peringuey 1896 Syntomus Hope 1838 Tilius Chaudoir 1876 Trichidema Basilewsky 1956 dd Subtribe Metallicina Basilewsky 1984Euproctinus Leng amp Mutchler 1927 Metallica Chaudoir 1873 Pachycallida Jeannel 1949 Parena Motschulsky 1860 dd Subtribe Peliocypadina Basilewsky 1984Demetriola Jeannel 1949 Peliocypas Schmidt Goebel 1846 dd Subtribe Pericalina Hope 1838Agonocheila Chaudoir 1848 Allophanes Andrewes 1939 Allophanopsis Louwerens 1952 Amphimenes Bates 1873 Amphimenoides Kirschenhofer 1999 Antimerina Alluaud 1897 Arsinoe Laporte 1834 Auchmerus Andrewes 1929 Bellavalentis Hunting amp Yang 2019 Brachichila Chaudoir 1870 Brachyctis Chaudoir 1870 Catascopellus Straneo 1969 Catascopus Kirby 1825 Cheilagona Baehr 2006 Coptodera Dejean 1825 Coptoglossus Chaudoir 1870 Coptoptera Chaudoir 1837 Coptopterella Basilewsky 1961 Coptopterina Basilewsky 1956 Crassagena Baehr 2006 Cylindropectus Lorenz 1998 Dolichoctis Schmidt Goebel 1846 Dontolobus Basilewsky 1970 Drymatus Motschulsky 1862 Eucheila Dejean 1829 Eurycoleus Chaudoir 1848 Eurydera Laporte 1831 Formosiella Jedlicka 1951 Gidda Andrewes 1920 Glyphodactyla Chaudoir 1837 Holcoderus Chaudoir 1870 Horniulus Jedlicka 1932 Labocephalus Chaudoir 1848 Lelis Chaudoir 1870 Lioptera Chaudoir 1870 Lobodontidius Basilewsky 1970 Lobodontulus Basilewsky 1970 Lobodontus Chaudoir 1842 Madecassina Jeannel 1949 Mascarenhia Alluaud 1933 Menarus Andrewes 1939 Metascopus Basilewsky 1970 Minuphloeus Darlington 1968 Minuthodes Andrewes 1941 Miscelus Klug 1834 Mochtherus Schmidt Goebel 1846 Mormolyce Hagenbach 1825 Mormolycina Jeannel 1949 Neocoptodera Jeannel 1949 Nycteis Laporte 1835 Oreodicastes Maindron 1905 Oxyodontus Chaudoir 1870 Paradolichoctis Baehr 2006 Pareurydera Jeannel 1949 Pectinitarsus Fairmaire 1881 Pericalus W S MacLeay 1825 Philophlaeus Chaudoir 1844 Phloeoxena Chaudoir 1870 Pristacrus Chaudoir 1870 Pseudoplatiella Baehr 2012 Serrimargo Chaudoir 1870 Sfitakantha Andrewes 1919 Sinurus Chaudoir 1870 Stenognathus Chaudoir 1843 Stenotelus Bouchard 1903 Stilboma Andrewes 1933 Stricklandiana Bousquet 2002 Tantillus Chaudoir 1870 Thyreochaetus Basilewsky 1959 Thyreopterus Dejean 1831 Thysanotus Chaudoir 1848 Trichocoptodera Louwerens 1958 Xanthos Kirschenhofer 2003 Xenitenopsis Basilewsky 1956 Xenitenus Peringuey 1896 dd Subtribe Physoderina Chaudoir 1877Allocota Motschulsky 1860 Anchista Nietner 1856 Dasiosoma Britton 1937 Diamella Shi amp Liang 2013 Endynomena Chaudoir 1873 Lachnoderma W J MacLeay 1873 Metallanchista Shi amp Liang 2013 Orionella Jedlicka 1964 Paraphaea Bates 1873 Physodera Eschscholtz 1829 dd Subtribe Pseudotrechina Basilewsky 1984Pseudotrechus Rosenhauer 1856 dd Subtribe Singilina Jeannel 1949Singilis Rambur 1837 dd Subtribe Somotrichina Mateu 1963Oecornis Britton 1940 Paulianites Jeannel 1949 Pephrica Alluaud 1936 Somotrichus Seidlitz 1887 dd Subtribe Sugimotoina Habu 1975Sugimotoa Habu 1975 dd Subtribe Trichina Basilewsky 1984Trichis Klug 1832 dd Cymindoides Motschulsky 1856 Lebina Germar 1813 Miolebidia Zhang Liu amp Shangguan 1989 Protoscalidion L Schaufuss 1888 dd Tribe Odacanthini Laporte 1834Subtribe Actenonycina Bates 1871Actenonyx White 1846 dd Subtribe Homethina Liebherr 2016Aeolodermus Andrewes 1929 Cyphocoleus Chaudoir 1877 Diplacanthogaster Liebke 1932 Homethes Newman 1842 Quammenis Erwin 2000 Stenocheila Laporte 1832 dd Subtribe Odacanthina Laporte 1834Anasis Laporte 1867 Andrewesia Liebke 1938 Arame Andrewes 1919 Archicolliuris Liebke 1931 Asios Liebke 1933 Aulacolius Sloane 1923 Basistichus Sloane 1917 Clarencia Sloane 1917 Colliuris DeGeer 1774 Crassacantha Baehr 1995 Cryptocolliuris Basilewsky 1955 Deipyrodes Bousquet 2002 Dicraspeda Chaudoir 1863 Dobodura Darlington 1968 Erectocolliuris Liebke 1931 Eucolliuris Liebke 1931 Eudalia Laporte 1867 Gestroania Liebke 1938 Giachinoana Baehr 2003 Lachnothorax Motschulsky 1862 Lasiocera Dejean 1831 Mimocolliuris Liebke 1933 Myrmecodemus Sloane 1923 Neoeudalia Baehr 2005 Odacantha Paykull 1798 Ophionea Klug 1821 Polydamasium Liebke 1938 Porocara Sloane 1917 Protocolliuris Liebke 1931 Renneria Baehr 1999 Smeringocera Chaudoir 1863 Stenidia Brulle 1834 Tricharnhemia Baehr 2009 dd Subtribe Pentagonicina Bates 1873Parascopodes Darlington 1968 Pentagonica Schmidt Goebel 1846 Scopodes Erichson 1842 dd dd Tribe Perigonini G Horn 1881Diploharpus Chaudoir 1850 Mizotrechus Bates 1872 Perigona Laporte 1835 Ripogenites Basilewsky 1954 dd Subfamily Licininae Bonelli 1810 edit Tribe Chaetogenyini Emden 1958Camptotoma Reiche 1843 dd Tribe Chlaeniini Brulle 1834Subtribe Callistina Laporte 1834Callistomimus Chaudoir 1872 Callistus Bonelli 1810 dd Subtribe Chlaeniina Brulle 1834Actodus Alluaud 1915 Chlaenius Bonelli 1810 Eccoptomenus Chaudoir 1850 Ectenognathus Murray 1858 Globulipalpus Sciaky amp Facchini 2019 Harpaglossus Motschulsky 1858 Hololeius LaFerte Senectere 1851 Holosoma Semenov 1889 Mirachlaenius Facchini 2011 Parachlaenius Kolbe 1894 Perissostomus Alluaud 1930 Procletodema Peringuey 1898 Procletus Peringuey 1896 Rhopalomelus Boheman 1848 Sphodroschema Alluaud 1930 Stenoodes Basilewsky 1953 Straneomelus Sciaky amp Facchini 2019 Stuhlmannium Kolbe 1894 Viridagonum Lassalle 2015 Rhopalochlaenius Zhang Sun amp Zhang 1994 dd dd Tribe Licinini Bonelli 1810Subtribe Dicaelina Laporte 1834Dicaelus Bonelli 1813 Diplocheila Brulle 1835 dd Subtribe Dicrochilina Ball 1992Dicrochile Guerin Meneville 1846 dd Subtribe Lestignathina Ball 1992Atrotus Peringuey 1896 Dilonchus Andrewes 1936 Hormacrus Sloane 1898 Lacordairia Laporte 1867 Lestignathus Erichson 1842 Microferonia Blackburn 1890 Microzargus Sciaky amp Facchini 1997 Platylytron W J MacLeay 1873 Siagonyx W J MacLeay 1871 Zargus Wollaston 1854 dd Subtribe Licinina Bonelli 1810Badister Clairville 1806 Colpostoma Semenov 1889 Derostichus Motschulsky 1860 Eurygnathus Wollaston 1854 Eutogeneius Solier 1849 Licinus Latreille 1802 Omestes Andrewes 1933 Physolaesthus Chaudoir 1850 dd dd Tribe Oodini LaFerte Senectere 1851Subtribe Dercylina Sloane 1923Dercylus Laporte 1832 dd Subtribe Geobaenina Peringuey 1896Geobaenus Dejean 1829 dd Subtribe Melanchitonina Jeannel 1948Dicaelindus W S MacLeay 1825 Melanchiton Andrewes 1940 Melanchrous Andrewes 1940 dd Subtribe Oodina LaFerte Senectere 1851Acanthoodes Basilewsky 1953 Acutosternus Lecordier amp Girard 1988 Adelopomorpha Heller 1916 Anatrichis LeConte 1853 Bamaroodes Gueorguiev 2014 Brachyodes Jeannel 1949 Chaetocrepis Chaudoir 1857 Coptocarpus Chaudoir 1857 Dercylinus Chaudoir 1883 Evolenes LeConte 1853 Holcocoleus Chaudoir 1883 Hoplolenus LaFerte Senectere 1851 Lobatodes Basilewsky 1956 Lonchosternus LaFerte Senectere 1851 Macroprotus Chaudoir 1878 Megaloodes Lesne 1896 Microodes Jeannel 1949 Miltodes Andrewes 1922 Nanodiodes Bousquet 1996 Neoodes Basilewsky 1953 Nothoodes Gueorguiev amp Liang 2020 Oodes Bonelli 1810 Oodinus Motschulsky 1865 Orthocerodus Basilewsky 1946 Polychaetus Chaudoir 1882 Prionognathus LaFerte Senectere 1851 Protopidius Basilewsky 1949 Pseudoodes Gueorguiev amp Liang 2020 Pseudosphaerodes Jeannel 1949 Simous Chaudoir 1882 Sphaerodinus Jeannel 1949 Sphoerodes Chaudoir 1883 Stenocrepis Chaudoir 1857 Sundaoodes Gueorguiev amp Liang 2020 Systolocranius Chaudoir 1857 Thaioodes Gueorguiev 2014 Thryptocerus Chaudoir 1878 Trichopalpoodes B Gueorguiev amp J Schmidt 2016 dd dd Subfamily Loricerinae Bonelli 1810 edit Loricera Latreille 1802Subfamily Melaeninae Csiki 1933 edit Cymbionotum Baudi di Selve 1864 Melaenus Dejean 1831Subfamily Migadopinae Chaudoir 1861 edit Tribe Amarotypini Erwin 1985Amarotypus Bates 1872 Migadopiella Baehr 2009 dd Tribe Migadopini Chaudoir 1861Antarctonomus Chaudoir 1861 Aquilex Moret 1989 Calathosoma Jeannel 1938 Calyptogonia Sloane 1920 Decogmus Sloane 1915 Dendromigadops Baehr 2013 Lissopterus G R Waterhouse 1843 Loxomerus Chaudoir 1842 Migadopidius Jeannel 1938 Migadops G R Waterhouse 1842 Monolobus Solier 1849 Nebriosoma Laporte 1867 Pseudomigadops Jeannel 1938 Rhytidognathus Chaudoir 1861 Stichonotus Sloane 1910 Taenarthrus Broun 1914 dd Subfamily Nebriinae Laporte 1834 edit Tribe Cicindini Csiki 1927Archaeocindis Kavanaugh amp Erwin 1991 Cicindis Bruch 1908 dd Tribe Nebriini Laporte 1834Leistus Frolich 1799 Nebria Latreille 1802 Nippononebria Ueno 1955 Archaeonebria Kavanaugh amp J Schmidt 2019 Ledouxnebria Deuve 1998 dd Tribe Notiokasiini Kavanaugh amp Negre 1983Notiokasis Kavanaugh amp Negre 1983 dd Tribe Notiophilini Motschulsky 1850Notiophilus Dumeril 1805 dd Tribe Opisthiini Dupuis 1912Opisthius Kirby 1837 Paropisthius Casey 1920 dd Tribe Pelophilini Kavanaugh 1996Pelophila Dejean 1821 dd Subfamily Nototylinae Banninger 1927 edit Nototylus Gemminger amp Harold 1868Subfamily Omophroninae Bonelli 1810 edit Omophron Latreille 1802Subfamily Orthogoniinae Schaum 1857 edit Tribe Amorphomerini Sloane 1923Amorphomerus Sloane 1923 dd Tribe Idiomorphini Bates 1891Subtribe Idiomorphina Bates 1891Idiomorphus Chaudoir 1846 Rathymus Dejean 1831 Strigia Brulle 1835 dd Subtribe Perochnoristhina Basilewsky 1973Perochnoristhus Basilewsky 1973 dd dd Tribe Orthogoniini Schaum 1857Subtribe Glyptina G Horn 1881Glyptus Brulle 1837 Neoglyptus Basilewsky 1953 dd Subtribe Orthogoniina Schaum 1857Actenoncus Chaudoir 1872 Anoncopeucus Chaudoir 1872 Hexachaetus Chaudoir 1872 Neoorthogonius Tian amp Deuve 2006 Nepalorthogonius Habu 1979 Orthogonius W S MacLeay 1825 dd dd Subfamily Panagaeinae Bonelli 1810 edit Tribe Brachygnathini Basilewsky 1946Brachygnathus Perty 1830 dd Tribe Panagaeini Bonelli 1810Subtribe Bascanina Basilewsky 1953Bascanus Peringuey 1896 dd Subtribe Panagaeina Bonelli 1810Adischissus Fedorenko 2015 Calathocosmus Emden 1928 Cintaroa Kasahara 1989 Cintaromorpha Hackel amp Anichtchenko 2016 Coptia Brulle 1835 Craspedophorus Hope 1838 Dischissus Bates 1873 Epigraphodes Basilewsky 1967 Epigraphus Chaudoir 1869 Euschizomerus Chaudoir 1850 Geobius Dejean 1831 Micrixys LeConte 1854 Microschemus Andrewes 1940 Panagaeus Latreille 1802 Paregraphus Basilewsky 1967 Peronomerus Schaum 1854 Psecadius Alluaud 1911 Tefflus Leach in Samouelle 1819 Tinoderus Chaudoir 1879 Tinognathus Chaudoir 1879 Trichisia Motschulsky 1865 dd dd Tribe Peleciini Chaudoir 1880Subtribe Agonicina Sloane 1920Agonica Sloane 1920 Pseudagonica B Moore 1960 dd Subtribe Peleciina Chaudoir 1880Ardistomopsis Straneo amp Ball 1989 Disphericus G R Waterhouse 1842 Dyschiridium Chaudoir 1861 Eripus Dejean 1829 Pelecium Kirby 1819 Stricteripus Straneo amp Ball 1989 dd dd Subfamily Patrobinae Kirby 1837 edit Tribe Lissopogonini Zamotajlov 2000Lissopogonus Andrewes 1923 dd Tribe Patrobini Kirby 1837Subtribe Deltomerina Chaudoir 1872Deltomerus Motschulsky 1850 dd Subtribe Deltomerodina Zamotajlov 2002Deltomerodes Deuve 1992 dd Subtribe Patrobina Kirby 1837Apatrobus Habu amp Baba 1960 Apenetretus Kurnakov 1960 Archipatrobus Zamotajlov 1992 Caelopenetretus Zamotajlov amp Ito 2000 Chaetapatrobus Lafer 1996 Chinapenetretus Kurnakov 1963 Dimorphopatrobus Casale amp Sciaky 1994 Diplous Motschulsky 1850 Grandipenetretus Zamotajlov amp Sciaky 1999 Himalopenetretus Zamotajlov 2002 Indopatrobus Zamotajlov amp Wrase 2006 Ledouxius Zamotajlov 1992 Minipenetretus Zamotajlov 2002 Minypatrobus Ueno 1955 Naxipenetretus Zamotajlov 1999 Parapatrobus Zamotajlov 1992 Parapenetretus Kurnakov 1960 Patanitretus Zamotajlov 2002 Patrobus Dejean 1821 Penetretus Motschulsky 1865 Platidiolus Chaudoir 1878 Platypatrobus Darlington 1938 Prodiplous Zamotajlov amp Sciaky 2006 Qiangopatrobus Zamotajlov 2002 Quasipenetretus Zamotajlov 2002 Tibetopenetretus Zamotajlov amp Sciaky 1999 dd dd Subfamily Paussinae Latreille 1806 edit Tribe Metriini LeConte 1853Subtribe Kryzhanovskianina Deuve 2019 Kryzhanovskiana Kataev amp Kirejtshuk 2019 dd Metrius Eschscholtz 1829 Sinometrius Wrase amp J Schmidt 2006 dd Tribe Ozaenini Hope 1838Subtribe Microzaenina Deuve 2019Microzaena Fairmaire 1901 dd Subtribe Mystropomina G Horn 1881Mystropomus Chaudoir 1848 dd Subtribe Ozaenina Hope 1838Crepidozaena Deuve 2001 Entomoantyx Ball amp McCleve 1990 Filicerozaena Deuve 2001 Gibbozaena Deuve 2001 Goniotropis G R Gray 1831 Inflatozaena Deuve 2001 Mimozaena Deuve 2001 Ozaena Olivier 1812 Pachyteles Perty 1830 Physea Brulle 1835 Physeomorpha Ogueta 1963 Platycerozaena Banninger 1927 Proozaena Deuve 2001 Serratozaena Deuve 2001 Tachypeles Deuve 2001 Tropopsis Solier 1849 dd Subtribe Pseudozaenina Sloane 1905Anentmetus Andrewes 1924 Dhanya Andrewes 1919 Eustra Schmidt Goebel 1846 Itamus Schmidt Goebel 1846 Pseudozaena Laporte 1834 Sphaerostylus Chaudoir 1848 dd dd Tribe Paussini Latreille 1806Subtribe Carabidomemnina Wasmann 1928Carabidomemnus Kolbe 1924 Eohomopterus Wasmann 1919 dd Subtribe Cerapterina Billberg 1820Arthropterus W S MacLeay 1838 Cerapterus Swederus 1788 Megalopaussus Lea 1906 Mesarthropterus Wasmann 1926 dd Subtribe Heteropaussina Janssens 1950Heteropaussus J Thomson 1860 dd Subtribe Homopterina Wasmann 1920Homopterus Westwood 1841 dd Subtribe Paussina Latreille 1806Allorhopalus Nagel 2017 Ceratoderus Westwood 1841 Euplatyrhopalus Desneux 1905 Lebioderus Westwood 1838 Leleupaussus Luna de Carvalho 1962 Melanospilus Westwood 1847 Paussomorphus Raffray 1885 Paussus Linnaeus 1775 Platyrhopalides Wasmann 1918 Platyrhopalopsis Desneux 1905 Platyrhopalus Westwood 1833 Pterorhopalus Maruyama 2011 dd Subtribe Pentaplatarthrina Jeannel 1946Hexaplatarthrus Jeannel 1955 Pentaplatarthrus Westwood 1833 dd Subtribe Arthropteritina Luna de Carvalho 1961 Arthropterites Wasmann 1926 Cerapterites Wasmann 1926 Protocerapterus Wasmann 1926 dd Subtribe Eopaussina Luna de Carvalho 1951 Eopaussus Wasmann 1926 dd dd Tribe Protopaussini Gestro 1892Protopaussus Gestro 1892 dd Subfamily Platyninae Bonelli 1810 edit Tribe Omphreini Ganglbauer 1891Omphreus Dejean 1828 dd Tribe Platynini Bonelli 1810Subtribe Enoicina Basilewsky 1985Abacetodes Straneo 1939 Enoicus Peringuey 1896 dd Subtribe Platynina Bonelli 1810Achaetocephala Habu 1975 Achaetoprothorax Habu 1978 Aepsera Chaudoir 1874 Agelaea Gene 1839 Agonidium Jeannel 1948 Agonobembix Jeannel 1948 Agonoriascus Basilewsky 1985 Agonorites Jeannel 1951 Agonum Bonelli 1810 Altagonum Darlington 1952 Anchomenus Bonelli 1810 Andinocolpodes Perrault 1991 Andrewesius Andrewes 1939 Aparupa Andrewes 1930 Archagonum Basilewsky 1953 Archicolpodes J Schmidt 2001 Arhytinus Bates 1889 Atranodes Jedlicka 1953 Atranus LeConte 1847 Austroglyptolenus Roig Junent 2003 Beckeria Jedlicka 1931 Blackburnia Sharp 1878 Bothrocolpodes Basilewsky 1985 Bruskespar Morvan 1998 Bruskmoal Morvan 1998 Bryanites Valentine 1987 Callidagonum Lorenz 1998 Cardiomera Bassi 1834 Catacolpodes Basilewsky 1985 Celaenagonum Habu 1978 Chaetagonum Burgeon 1933 Chaetosaurus J Schmidt 2001 Cinctagonum Baehr 2012 Cistelagonum Baehr 2012 Collagonum Baehr 1995 Colpodes W S MacLeay 1825 Colpoides Jedlicka 1931 Colpomimus Basilewsky 1985 Colposphodrus Jedlicka 1953 Ctenognathus Fairmaire 1843 Cymenopterus Jeannel 1948 Cyrtopilus Basilewsky 1985 Dalatagonum Fedorenko 2011 Deliaesianum Morvan 1999 Deltocolpodes Morvan 1992 Dendragonum Jeannel 1948 Diacanthostylus Habu 1978 Dicranoncoides Habu 1978 Dicranoncus Chaudoir 1850 Dinocolpodes J Schmidt 2001 Dirotus W S MacLeay 1825 Dister Morvan 2006 Dolichocolpodes Basilewsky 1985 Dyscolus Dejean 1831 Elliptoleus Bates 1882 Epicolpodes Basilewsky 1985 Eucolpodes Jeannel 1948 Euleptus Klug 1833 Euplynes Schmidt Goebel 1846 Feroniascus Jeannel 1951 Fortagonum Darlington 1952 Galiciotyphlotes Assmann 1999 Gastragonum Darlington 1952 Glaucagonum Habu 1978 Glyptolenoides Perrault 1991 Glyptolenus Bates 1878 Gyrochaetostylus Habu 1978 Habragonum Ueno 1964 Hannaphota Landin 1955 Haplocolpodes Jeannel 1951 Haplopeza Boheman 1848 Helluocolpodes Liebherr 2005 Hemiplatynus Casey 1920 Henvelik Morvan 1999 Herculagonum Baehr 2002 Hikosanoagonum Habu 1954 Idiagonum Darlington 1952 Idiastes Andrewes 1931 Idiocolpodes Basilewsky 1985 Incagonum Liebherr 1994 Iridagonum Darlington 1952 Ischnagonum Kasahara amp Sato 1997 Jocqueius Basilewsky 1988 Jujiroa Ueno 1952 Kalchdigor Morvan 1999 Kar Morvan 1998 Karnes Morvan 2010 Kaszabellus Jedlicka 1954 Kiwiplatynus Larochelle amp Lariviere 2021 Klapperichella Jedlicka 1956 Kuceraianum Morvan 2002 Kupeplatynus Larochelle amp Lariviere 2021 Laevagonum Darlington 1952 Lassalleianum Morvan 1999 Lepcha Andrewes 1930 Leptagonum Kolbe 1897 Leptocolpodes Basilewsky 1985 Letouzeya Bruneau de Mire 1982 Liagonum Jeannel 1948 Liamegalonychus Basilewsky 1963 Liocolpodes Basilewsky 1985 Lissagonum Habu 1978 Lithagonum Darlington 1952 Lobocolpodes Basilewsky 1985 Lorostema Motschulsky 1865 Loxocrepis Eschscholtz 1829 Lucicolpodes J Schmidt 2000 Maculagonum Darlington 1952 Maoriplatynus Larochelle amp Lariviere 2021 Meleagros Kirschenhofer 1999 Mesocolpodes Basilewsky 1985 Metacolpodes Jeannel 1948 Mexisphodrus Barr 1965 Montagonum Darlington 1952 Mooreagonum Baehr 2016 Morimotoidius Habu 1954 Nebriagonum Darlington 1952 Negreum Habu 1958 Neobatenus Jeannel 1948 Neocolpodes Jeannel 1948 Neodendragonum Basilewsky 1953 Neomegalonychus Jeannel 1948 Nesiocolpodes Jeannel 1948 Nipponagonum Habu 1978 Notagonum Darlington 1952 Notocolpodes Basilewsky 1985 Notoplatynus B Moore 1985 Nymphagonum Habu 1978 Olisthopus Dejean 1828 Oncostylus Habu 1978 Onotokiba Alluaud 1927 Onycholabis Bates 1873 Onypterygia Dejean 1831 Orophicus Alluaud 1925 Orthotrichus Peyron 1856 Oxygonium Basilewsky 1951 Oxypselaphus Chaudoir 1843 Pachybatenus Basilewsky 1973 Pachyferonia Jeannel 1951 Paracolpodes Basilewsky 1985 Paraliagonum Basilewsky 1957 Paramegalonychus Basilewsky 1953 Paranchodemus Habu 1978 Paranchus Lindroth 1974 Paraplatynus Baehr 2016 Pawgammm Morvan 1999 Platyagonum Habu 1978 Platynus Bonelli 1810 Plaumannium Liebke 1939 Plicagonum Darlington 1952 Ponapagonum Darlington 1970 Potamagonum Darlington 1952 Praepristus Kirschenhofer 1999 Promecoptera Dejean 1831 Promegalonychus Basilewsky 1953 Prophenorites Basilewsky 1985 Prosphodrus Britton 1959 Protocolpodes Basilewsky 1985 Pseudanchomenus Tarnier 1860 Pseudobatenus Basilewsky 1951 Pseudomegalonychus Basilewsky 1950 Rhadine LeConte 1846 Rupa Jedlicka 1935 Scotagonum Habu 1978 Sericoda Kirby 1837 Shibataia Habu 1978 Sinocolpodes J Schmidt 2001 Skoeda Morvan 1995 Skorlagad Morvan 1999 Skouedhirraad Morvan 1999 Sophroferonia Alluaud 1933 Speagonum B Moore 1977 Speocolpodes Barr 1974 Speokokosia Alluaud 1932 Sperkanhir Morvan 2010 Stenocnemus Mannerheim 1837 Straneoa Basilewsky 1953 Syletor Tschitscherine 1899 Takasagoagonum Habu 1977 Tanystoma Motschulsky 1845 Tarsagonum Darlington 1952 Tetraleucus Casey 1920 Tostkar Morvan 1998 Trogloagonum Casale 1982 Tuiplatynus Larochelle amp Lariviere 2021 Vitagonum B Moore 1999 Vulcanophilus Heller 1898 Xestagonum Habu 1978 Praeanchodemus J Schmidt Gopel amp Will 2017 dd dd Tribe Sphodrini Laporte 1834Subtribe Atranopsina Baehr 1982Amaroschema Jeannel 1943 Amphimasoreus Piochard de la Brulerie 1875 Atranopsis Baehr 1982 Broter Andrewes 1923 Gomerina Bolivar y Pieltain 1940 Paraeutrichopus Mateu 1954 Platyderus Stephens 1827 Pseudomyas Uyttenboogaart 1929 Pseudoplatyderus Bolivar y Pieltain 1940 dd Subtribe Calathina Laporte 1834Calathus Bonelli 1810 Thermoscelis Putzeys 1873 dd Subtribe Dolichina Brulle 1834Acalathus Semenov 1889 Anchomenidius Heyden 1880 Casaleius Sciaky amp Wrase 1998 Dolichus Bonelli 1810 Doliodactyla Sciaky amp Wrase 1998 Morphodactyla Semenov 1889 Xestopus Andrewes 1937 dd Subtribe Pristosiina Lindroth 1956Pristosia Motschulsky 1865 dd Subtribe Sphodrina Laporte 1834Calathidius Putzeys 1873 Cephalosdrophus Lassalle amp Marcilhac 1999 Eosphodrus Casale 1988 Eremosphodrus Semenov 1909 Himalosphodrus Casale 1988 Hystricosphodrus Casale amp Giachino 2004 Ifridytes Deuve amp Queinnec 1994 Laemostenus Bonelli 1810 Licinopsis Bedel 1899 Miquihuana Barr 1982 Pseudotaphoxenus L Schaufuss 1865 Reflexisphodrus Casale 1988 Sphodropsis Seidlitz 1887 Sphodrus Clairville 1806 Stenolepta Semenov 1889 Taphoxenus Motschulsky 1850 dd Subtribe Synuchina Lindroth 1956Nipponosynuchus Morita 1998 Parabroscus Lindroth 1956 Synuchus Gyllenhal 1810 Trephionus Bates 1883 dd dd Subfamily Promecognathinae LeConte 1853 edit Tribe Axinidiini Basilewsky 1963Axinidium Sturm 1843 Holaxinidium Basilewsky 1963 Metaxinidium Basilewsky 1963 Paraxinidium Basilewsky 1963 dd Tribe Dalyatini Mateu 2002Dalyat Mateu 2002 dd Tribe Promecognathini LeConte 1853Promecognathus Chaudoir 1846 dd Tribe Palaeoaxinidiini McKay 1991 Palaeoaxinidium McKay 1991 dd Subfamily Pseudomorphinae Hope 1838 edit Adelotopus Hope 1836 Cainogenion Notman 1925 Cryptocephalomorpha Ritsema 1875 Guyanemorpha Erwin 2013 Manumorpha Erwin amp Geraci 2008 Notopseudomorpha Baehr 1997 Paussotropus C O Waterhouse 1877 Pseudomorpha Kirby 1823 Samiriamorpha Erwin amp Geraci 2008 Sphallomorpha Westwood 1840 Tuxtlamorpha Erwin amp Geraci 2008 Yasunimorpha Erwin amp Geraci 2008Subfamily Psydrinae LeConte 1853 edit Tribe Gehringiini Darlington 1933Subtribe Gehringiina Darlington 1933Gehringia Darlington 1933 dd Subtribe Helenaeina Deuve 2007Afrogehringia Baehr Schule amp Lorenz 2009 Helenaea Schatzmayr amp Koch 1934 dd dd Tribe Moriomorphini Sloane 1890Subtribe Amblytelina Blackburn 1892Amblytelus Erichson 1842 Dystrichothorax Blackburn 1892 Epelyx Blackburn 1892 Mecyclothorax Sharp 1903 Meonis Laporte 1867 Paratrichothorax Baehr 2004 Pseudamblytelus Baehr 2004 Raphetis B Moore 1963 Selenochilus Chaudoir 1878 Trichamblytelus Baehr 2004 dd Subtribe Moriomorphina Sloane 1890Celanida Laporte 1867 Melisodera Westwood 1835 Molopsida White 1846 Moriodema Laporte 1867 Moriomorpha Laporte 1867 Neonomius B Moore 1963 Pterogmus Sloane 1920 Rhaebolestes Sloane 1903 Rossjoycea Liebherr 2011 Sitaphe B Moore 1963 Spherita Liebherr 2020 Tarastethus Sharp 1883 Teraphis Laporte 1867 Theprisa B Moore 1963 Trephisa B Moore 1963 Trichopsida Larochelle amp Lariviere 2013 dd Subtribe Tropopterina Sloane 1898Pharetis Liebherr 2020 Tropopterus Solier 1849 dd dd Tribe Psydrini LeConte 1853Laccocenus Sloane 1890 Nomius Laporte 1835 Psydrus LeConte 1846 dd Subfamily Pterostichinae Bonelli 1810 edit Tribe Chaetodactylini Tschitscherine 1903Chaetodactyla Tschitscherine 1897 dd Tribe Cnemalobini Germain 1911Cnemalobus Guerin Meneville 1838 dd Tribe Cratocerini Lacordaire 1854Subtribe Catapieseina Bates 1882Catapiesis Solier 1835 Homalomorpha Brulle 1837 dd Subtribe Cratocerina Lacordaire 1854Brachidius Chaudoir 1852 Cratocerus Dejean 1829 Oxyglychus Straneo 1938 dd Subtribe Drimostomatina Chaudoir 1873Abacaelostus Straneo 1952 Andrewesinulus Straneo 1938 Apsidocnemus Alluaud 1936 Barylaus Liebherr 1985 Caecocaelus Straneo 1949 Caelostomus W S MacLeay 1825 Camptogenys Tschitscherine 1899 Capabatus Csiki 1930 Crenulostrigus Straneo 1942 Cyrtolaus Bates 1882 Dactyleurys Tschitscherine 1899 Dactylinius Straneo 1941 Diachipteryx Alluaud 1925 Diceromerus Chaudoir 1873 Dromistomus Jeannel 1948 Drymonaxus Straneo 1941 Feostoma Straneo 1941 Hemitelestus Alluaud 1895 Hoplizomenus Chaudoir 1873 Leleuporites Straneo 1960 Madapelmus Dajoz 1985 Monodryxus Straneo 1942 Pachycaecus Straneo 1971 Pachyroxochus Straneo 1942 Platyxythrius Lorenz 1998 Stegazopteryx Will 2004 Stomonaxellus Tschitscherine 1901 Strigomerodes Straneo 1939 Strigomerus Chaudoir 1873 Trichillinus Straneo 1938 dd dd Tribe Microcheilini Jeannel 1948Microcheila Brulle 1835 dd Tribe Morionini Brulle 1837Buderes Murray 1857 Hyperectenus Alluaud 1935 Hyperion Laporte 1834 Megamorio Chaudoir 1880 Morion Latreille 1810 Morionidius Chaudoir 1880 Moriosomus Motschulsky 1855 Platynodes Westwood 1847 Stereostoma Murray 1857 dd Tribe Pterostichini Bonelli 1810Subtribe Abacetina Chaudoir 1873Abacaecus Allegro amp Giachino 2020 Abacetus Dejean 1828 Aristopus LaFerte Senectere 1853 Celioinkosa Straneo 1951 Chlaeminus Motschulsky 1865 Colpodichius Straneo 1952 Cosmodiscus Sloane 1907 Cyrtomoscelis Chaudoir 1874 Distrigidius Jeannel 1948 Ecnomolaus Bates 1892 Haptoderidius Straneo 1975 Haptoderodes Straneo 1986 Holconotus Schmidt Goebel 1846 Inkosa Peringuey 1926 Mateuellus Deuve 1990 Metabacetus Bates 1892 Metaxellus Straneo 1960 Metaxys Chaudoir 1857 Novillidius Straneo 1941 Oodinkosa Straneo 1939 Ophonichius Straneo 1942 Pediomorphus Chaudoir 1878 Pioprosopus Tschitscherine 1899 Pollicobius Vinson 1939 Prostalomus Basilewsky 1950 Pseudabacetus Burgeon 1935 Pterostillichus Straneo 1949 Rhagadillius Straneo 1951 Tiferonia Darlington 1962 Trachelocyphoides Straneo 1942 Trachelocyphus Tschitscherine 1900 dd Subtribe Euchroina Chaudoir 1874Abacillius Straneo 1949 Abacillodes Straneo 1988 Abaris Dejean 1831 Apsaustodon Tschitscherine 1901 Argutoridius Chaudoir 1876 Blennidus Motschulsky 1866 Blenniventer Straneo 1986 Bothynoproctus Tschitscherine 1900 Cedrorum Borges amp A Serrano 1993 Cephalostichus Straneo 1977 Cynthidia Chaudoir 1874 Euchroa Brulle 1835 Eumara Tschitscherine 1901 Gastrogmus Sloane 1915 Haplobothynus Tschitscherine 1901 Litarthrum Sloane 1915 Lobobrachus Sharp 1885 Marsyas Putzeys 1845 Meropalpus Tschitscherine 1900 Microcephalus Dejean 1828 Neotalus Will 2002 Oribazus Chaudoir 1874 Orthomus Chaudoir 1838 Pachythecus Chaudoir 1874 Paniestichus Will 2011 Parorthomus B Gueorguiev Wrase amp Farkac 2014 Phaenaulax Tschitscherine 1898 Prosopogmus Chaudoir 1865 Pseudabarys Chaudoir 1874 Setalimorphus Sloane 1895 Setalis Laporte 1867 Simodontus Chaudoir 1843 Trirammatus Chaudoir 1835 dd Subtribe Loxandrina Erwin amp Sims 1984Cerabilia Laporte 1867 Oxycrepis Reiche 1843 Zeodera Laporte 1867 dd Subtribe Metiina Straneo 1951Abropus G R Waterhouse 1842 Antarctiola Straneo 1977 Feroniola Tschitscherine 1900 Kuschelinus Straneo 1963 Metius Curtis 1839 dd Subtribe Pterostichina Bonelli 1810Abacidus LeConte 1863 Abacoleptus Fauvel 1903 Abacomorphus Chaudoir 1878 Abacophrastus Will 2011 Abacops Tschitscherine 1902 Abax Bonelli 1810 Allotriopus Bates 1882 Analoma Darlington 1971 Apophylon B Gueorguiev amp Sciaky 2015 Aristochroa Tschitscherine 1898 Aristochroodes Marcilhac 1993 Aulacopodus Britton 1940 Basilewskya Straneo 1948 Camptoscelis Dejean 1828 Castelnaudia Tschitscherine 1891 Catadromus W S MacLeay 1825 Chaetauchenium Tschitscherine 1900 Chalcochrous Chaudoir 1838 Cophosomorpha Tschitscherine 1891 Cratoferonia Tschitscherine 1902 Cratogaster Blanchard 1843 Cuneipectus Sloane 1907 Cyclotrachelus Chaudoir 1838 Cyrtoderus Hope 1842 Darodilia Laporte 1867 Delinius Westwood 1864 Eucamptognathus Chaudoir 1837 Eudromus Klug 1835 Euryabax Fauvel 1903 Euryaptus Bates 1892 Eurypercus Jeannel 1948 Eurystomis Chaudoir 1878 Gastrellarius Casey 1918 Gourlayia Britton 1964 Harpostomus Chaudoir 1856 Henrotiochoromus Busulini 1958 Henrotius Jeannel 1953 Holcaspis Chaudoir 1865 Hybothecus Chaudoir 1874 Leiolesticus Roux Lassalle amp Dubault 2016 Lesticus Dejean 1828 Licentius Jedlicka 1939 Liopasa Tschitscherine 1901 Lophoglossus LeConte 1853 Loxogenius Sloane 1907 Mecynognathus W J MacLeay 1873 Megadromus Motschulsky 1866 Molopidius Jeannel 1942 Molopinus Jeannel 1948 Molops Bonelli 1810 Myas Sturm 1826 Neoferonia Britton 1940 Nesites Andrewes 1939 Nirmala Andrewes 1930 Notabax B Moore 1976 Notolestus Sloane 1894 Notonomus Chaudoir 1865 Nurus Motschulsky 1866 Oberthueria Vuillet 1911 Ogmophora Tschitscherine 1896 Onawea Johns 2007 Oscadytes Lagar Mascaro 1975 Pachymorphus Chaudoir 1838 Paranurus Tschitscherine 1901 Pareuryaptus Dubault Lassalle amp Roux 2008 Parhypates Motschulsky 1866 Pedius Motschulsky 1850 Percolaus Bates 1882 Percus Bonelli 1810 Peyrieraselus Deuve 1981 Piesmus LeConte 1846 Platycoelus Blanchard 1843 Platysmodes Fauvel 1903 Plocamostethus Britton 1940 Poecilinus Jeannel 1948 Poecilus Bonelli 1810 Pseudoceneus Tschitscherine 1891 Pterostichus Bonelli 1810 Rhabdotus Chaudoir 1865 Rhytiferonia Darlington 1962 Rhytisternus Chaudoir 1865 Sarticus Motschulsky 1866 Secatophus Laporte 1867 Setalidius Chaudoir 1878 Speluncarius Reitter 1886 Speomolops Patrizi 1955 Sphodrosomus Perroud 1864 Stenochoromus L Miller 1866 Stereocerus Kirby 1837 Sthenocranion Tschitscherine 1899 Stomis Clairville 1806 Straneostichus Sciaky 1994 Styracoderus Chaudoir 1874 Tanythrix Schaum 1858 Taphoxomimus Straneo 1975 Tapinopterus Schaum 1858 Teratotarsa Tschitscherine 1893 Teropha Laporte 1867 Trichosternus Chaudoir 1865 Trigonaptus Fedorenko 2020 Trigonotoma Dejean 1828 Troglorites Jeannel 1919 Tropidocerus Chaudoir 1878 Typhlochoromus Moczarski 1913 Wahlbergiana Bousquet 2002 Xenion Tschitscherine 1902 Zariquieya Jeannel 1924 Zeopoecilus Sharp 1886 dd dd Tribe Zabrini Bonelli 1810Subtribe Amarina Zimmermann 1832Amara Bonelli 1810 Pseudamara Lindroth 1968 dd Subtribe Zabrina Bonelli 1810Zabrus Clairville 1806 dd dd Subfamily Rhysodinae Laporte 1840 edit Tribe Clinidiini R T amp J R Bell 1978Clinidium Kirby 1830 Grouvellina R T amp J R Bell 1978 Rhyzodiastes Fairmaire 1895 dd Tribe Dhysorini R T amp J R Bell 1978Dhysores Grouvelle 1903 Neodhysores R T amp J R Bell 1978 Tangarona R T amp J R Bell 1982 dd Tribe Leoglymmiini R T amp J R Bell 1978Leoglymmius R T amp J R Bell 1978 dd Tribe Medisorini R T amp J R Bell 1987Medisores R T amp J R Bell 1987 dd Tribe Omoglymmiini R T amp J R Bell 1978Arrowina R T amp J R Bell 1978 Omoglymmius Ganglbauer 1891 Plesioglymmius R T amp J R Bell 1978 Shyrodes Grouvelle 1903 Srimara R T amp J R Bell 1978 Xhosores R T amp J R Bell 1978 Yamatosa R T amp J R Bell 1979 dd Tribe Rhysodini Laporte 1840Kaveinga R T amp J R Bell 1978 Kupeus R T amp J R Bell 1982 Rhysodes Germar 1822 dd Tribe Sloanoglymmiini R T amp J R Bell 1991Sloanoglymmius R T amp J R Bell 1991 dd Subfamily Scaritinae Bonelli 1810 edit Tribe Clivinini Rafinesque 1815Subtribe Androzelmina R T Bell 1998Androzelma Dostal 1993 dd Subtribe Ardistomina Putzeys 1867Ardistomis Putzeys 1846 Aspidoglossa Putzeys 1846 Kearophus Dajoz 2004 Semiardistomis Kult 1950 dd Subtribe Clivinina Rafinesque 1815Afrosyleter Basilewsky 1959 Ancus Putzeys 1867 Basilewskyana Kult 1959 Brachypelus Putzeys 1867 Cameroniola Baehr 2000 Climax Putzeys 1861 Clivina Latreille 1802 Clivinarchus Sloane 1896 Cryptomma Putzeys 1846 Lachenus Putzeys 1846 Leleuporella Basilewsky 1956 Nannoryctes Baehr 2000 Nyctosyles Putzeys 1867 Orictites Andrewes 1931 Platysphyrus Sloane 1905 Psilidius Jeannel 1957 Pyramoides Bousquet 2002 Rhysocara Sloane 1916 Rubidiclivina Baehr 2015 Rugiluclivina Balkenohl 1996 Semiclivina Kult 1947 Sinesetosa Balkenohl 1996 Syleter Andrewes 1941 Thliboclivina Kult 1959 Trilophidius Jeannel 1957 Trilophus Andrewes 1927 Trogloclivina Deuve 2003 Whiteheadiana Perrault 1994 dd Subtribe Forcipatorina Banninger 1938Camptidius Putzeys 1867 Camptodontus Dejean 1826 Forcipator Maindron 1904 Kultianella Perrault 1994 Mesus Chevrolat 1858 Obadius Burmeister 1875 Oxygnathopsis Louwerens 1953 Oxygnathus Dejean 1826 Scolyptus Putzeys 1861 Stratiotes Putzeys 1846 dd Subtribe Reicheiina Jeannel 1957Alpiodytes Jeannel 1957 Asioreicheia Bulirsch amp Magrini 2014 Catalanodytes Sciaky 1989 Dalmatoreicheia Magrini amp Bulirsch 2005 Dimorphoreicheia Magrini Fancello amp Leo 2002 Galicioreicheia Felix amp Bulirsch 2015 Guiodytes Tian 2013 Gymnetoreicheia Magrini Fancello amp Casale 2019 Iberodytes Jeannel 1949 Ichnusodytes Magrini Fancello amp Onnis 2019 Italodytes G Muller 1938 Kenyoreicheia Bulirsch amp Magrini 2007 Laoreicheia Balkenohl 2005 Madagascareicheia Magrini amp Bulirsch 2009 Malagasyreicheia Magrini Bulirsch amp Fancello 2021 Orientoreicheia Bulirsch amp Hurka 1994 Oxydrepanus Putzeys 1867 Parareicheia Jeannel 1957 Reicheadella Reitter 1913 Reicheia Saulcy 1862 Reicheidius Jeannel 1957 Sikelioreicheia Magrini amp Fancello 2019 Spelaeodytes L Miller 1863 Typhloreicheia Holdhaus 1924 dd Subtribe Schizogeniina Dostal 2017Baehrogenius Dostal 2017 Coryza Putzeys 1867 Halocoryza Alluaud 1919 Lophocoryza Alluaud 1941 Paracoryza Basilewsky 1952 Psammocoryza Hogan 2006 Schizogenius Putzeys 1846 dd Subtribe Sparostesina Dostal 2017Bohemaniella Bousquet 2002 Pseudoclivina Kult 1947 Sparostes Putzeys 1867 dd dd Tribe Corintascarini Basilewsky 1973Corintascaris Basilewsky 1952 dd Tribe Dyschiriini Kolbe 1880Akephorus LeConte 1852 Caledyschirius Bulirsch 2010 Clivinopsis Bedel 1895 Cribrodyschirius Bruneau de Mire 1952 Dyschirius Bonelli 1810 Neodyschirius Kult 1954 Reicheiodes Ganglbauer 1891 Setodyschirius Fedorenko 1996 Striganoviella Fedorenko 2012 Torretassoa Schatzmayr amp Koch 1933 Dyschiriomimus Iablokoff Khnzorian 1960 dd Tribe Salcediini Alluaud 1930Subtribe Salcediina Alluaud 1930Salcedia Fairmaire 1899 dd Subtribe Solenogenyina R T Bell 1998Holoprizus Putzeys 1867 Solenogenys Westwood 1859 dd dd Tribe Scaritini Bonelli 1810Subtribe Carenina W J MacLeay 1887Carenidium Chaudoir 1868 Carenum Bonelli 1813 Epilectus Blackburn 1888 Euryscaphus W J MacLeay 1865 Laccopterum W J MacLeay 1878 Monocentrum Chaudoir 1868 Mouhotia Laporte 1862 Neocarenum Laporte 1867 Neoscaphus Sloane 1888 Philoscaphus W J MacLeay 1871 Scaraphites Westwood 1842 Trichocarenum Blackburn 1892 dd Subtribe Pasimachina Putzeys 1867Pasimachus Bonelli 1813 dd Subtribe Scapterina Putzeys 1867Acanthoscelis Dejean 1825 Oxylobus Chaudoir 1855 Parathlibops Basilewsky 1958 Passalidius Chaudoir 1863 Scapterus Dejean 1826 Steganomma W J MacLeay 1887 Thlibops Putzeys 1867 dd Subtribe Scaritina Bonelli 1810Anomophaenus Fauvel 1882 Antilliscaris Lorenz 1998 Baenningeria Reichardt 1976 Coptolobus Chaudoir 1857 Crepidopterus Chaudoir 1855 Cryptoscaphus Chaudoir 1855 Dinoscaris Alluaud 1902 Distichus Motschulsky 1858 Dyscaris Banninger 1940 Dyscherinus Jeannel 1955 Dyscherus Chaudoir 1855 Geoscaptus Chaudoir 1855 Glyptogrus Bates 1875 Gnaphon Andrewes 1920 Haplogaster Chaudoir 1879 Haplotrachelus Chaudoir 1855 Macromorphus Chaudoir 1857 Madascaris Banninger 1938 Mamboicus Bates 1886 Mecynoscaris Alluaud 1930 Menigius Chaudoir 1879 Neochryopus Banninger 1932 Ochyropus Schiodte 1847 Pachyodontus Chaudoir 1879 Paradyscherus Basilewsky 1971 Pilades Heyne 1895 Prodyscherodes Jeannel 1955 Prodyscherus Jeannel 1946 Scarites Fabricius 1775 Storthodontus Chaudoir 1855 Tapinoscaris Jeannel 1946 Tibioscarites Banninger 1929 Tonkinoscaris Banninger 1956 Typhloscaris Kuntzen 1914 dd dd Subfamily Siagoninae Bonelli 1813 edit Tribe Enceladini G Horn 1881Enceladus Bonelli 1813 dd Tribe Siagonini Bonelli 1813Luperca Laporte 1840 Siagona Latreille 1804 dd Subfamily Trechinae Bonelli 1810 edit Tribe Bembidarenini Maddison et al 2019Andinodontis Erwin amp Toledano 2010 Argentinatachoides Sallanave Erwin amp Roig Junent 2008 Bembidarenas Erwin 1972 Tasmanitachoides Erwin 1972 dd Tribe Bembidiini Stephens 1827Subtribe Anillina Jeannel 1937Afranillus Giachino 2015 Afrodipnus Giachino 2015 Angustanillus Baehr amp Main 2016 Anillaspis Casey 1918 Anillinus Casey 1918 Anillodes Jeannel 1963 Anillopsidius Coiffait 1969 Anillopsis Jeannel 1937 Anillotarsus Mateu 1980 Anillus Jacquelin du Val 1851 Argiloborus Jeannel 1937 Austranillus Giachino 2005 Bafutyphlus Bruneau de Mire 1986 Bhutanillus M E Schmid 1975 Bulirschia Giachino 2008 Bylibaraphanus Giachino Eberhard amp Perina 2021 Caeconannus Jeannel 1963 Caecoparvus Jeannel 1937 Carayonites Bruneau de Mire 1986 Cryptocharidius M Etonti amp Mateu 1992 Cryptorites Jeannel 1950 Dicropterus Ehlers 1883 Elgonotyphlus Sciaky amp Zaballos 1993 Erwinanillus Giachino Eberhard amp Perina 2021 Externanillus Baehr amp Main 2016 Geocharidius Jeannel 1963 Geocharis Ehlers 1883 Gilesdytes Giachino Eberhard amp Perina 2021 Gracilanillus Baehr amp Main 2016 Gregorydytes Giachino Eberhard amp Perina 2021 Hesperanillus Baehr amp Main 2016 Honduranillus Zaballos 1997 Hygranillus B Moore 1980 Hypodipnites Jeannel 1963 Hypotyphlus Jeannel 1937 Iason Giachino amp Vailati 2011 Iberanillus Espanol 1971 Illaphanus W J MacLeay 1865 Kimberleytyphlus Giachino Eberhard amp Perina 2021 Leleupanillus Basilewsky 1976 Magnanillus Baehr 2017 Malagasydipnus Giachino 2008 Malagasytyphlus Giachino 2008 Megastylulus Giachino amp Sciaky 2003 Mexanillus Vigna Taglianti 1973 Microdipnites Jeannel 1957 Microdipnodes Basilewsky 1960 Microdipnus Jeannel 1937 Microtyphlus Linder 1863 Mystroceridius Reichardt 1972 Neodipnus Jeannel 1957 Neoillaphanus Giachino Eberhard amp Perina 2021 Nesamblyops Jeannel 1937 Nothanillus Jeannel 1962 Orthotyphlus Mateu amp Zaballos 1998 Paranillopsis Cicchino amp Roig Junent 2001 Paranillus Jeannel 1949 Parillaphanus Baehr 2018 Parvocaecus Coiffait 1956 Pelocharis Jeannel 1960 Pelodiaetodes B Moore 1980 Pelodiaetus Jeannel 1937 Pelonomites Jeannel 1963 Perucharidius Mateu amp M Etonti 2002 Pilbaradytes Giachino Eberhard amp Perina 2021 Pilbaranillus Baehr amp Main 2016 Pilbaraphanus Giachino Eberhard amp Perina 2021 Prioniomus Jeannel 1937 Pseudanillus Bedel 1896 Pseudillaphanus Giachino 2005 Rhegmatobius Jeannel 1937 Scotodipnus Schaum 1860 Selenodipnus Jeannel 1963 Serranillus Barr 1995 Serratotyphlus Giachino 2008 Stylulus L Schaufuss 1882 Tasmanillus Giachino 2005 Typhlocharis Dieck 1869 Typhlonesiotes Jeannel 1937 Winklerites Jeannel 1937 Zapotecanillus Sokolov 2013 Zeanillus Jeannel 1937 Zoianillus Sciaky 1994 dd Subtribe Bembidiina Stephens 1827Amerizus Chaudoir 1868 Asaphidion Gozis 1886 Bembidion Latreille 1802 Caecidium Ueno 1971 Lionepha Casey 1918 Ocys Stephens 1828 Orzolina Machado 1987 Sinechostictus Motschulsky 1864 dd Subtribe Horologionina Jeannel 1949Horologion Valentine 1932 dd Subtribe Lovriciina Giachino B Gueorguiev amp Vailati 2011Lovricia Pretner 1979 Neolovricia Lakota Jalzic amp J Moravec 2009 Paralovricia Giachino B Gueorguiev amp Vailati 2011 dd Subtribe Tachyina Motschulsky 1862Aenigmatachys Baehr 2020 Afrotachys Basilewsky 1958 Anomotachys Jeannel 1946 Costitachys Erwin 1974 Elaphropus Motschulsky 1839 Kiwitachys Larochelle amp Lariviere 2007 Liotachys Bates 1871 Lymnastis Motschulsky 1862 Meotachys Erwin 1974 Micratopus Casey 1914 Moirainpa Erwin 1984 Nothoderis Boyd amp Erwin 2016 Pericompsus LeConte 1852 Polyderidius Jeannel 1962 Porotachys Netolitzky 1914 Setitachys Baehr 2016 Stigmatachys Boyd amp Erwin 2016 Straneoites Basilewsky 1947 Tachys Dejean 1821 Tachysbembix Erwin 2004 Tachyta Kirby 1837 Tachyxysta Boyd amp Erwin 2016 dd Subtribe Xystosomina Erwin 1994Erwiniana Paulsen amp A Smith 2003 Geballusa Erwin 1994 Gouleta Erwin 1994 Inpa Erwin 1978 Mioptachys Bates 1882 Philipis Erwin 1994 Xystosomus Schaum 1863 dd dd Tribe Pogonini Laporte 1834Bedeliolus Semenov 1900 Cardiaderus Dejean 1828 Diodercarus Lutshnik 1931 Diplochaetus Chaudoir 1872 Ochtozetus Chaudoir 1872 Olegius Komarov 1996 Pogonistes Chaudoir 1872 Pogonopsis Bedel 1898 Pogonus Dejean 1821 Sirdenus Dejean 1828 Syrdenoidius Baehr amp Hudson 2001 Thalassotrechus Van Dyke 1918 dd Tribe Sinozolini Deuve 1997Chaltenia Roig Junent amp Cicchino 2001 Phrypeus Casey 1924 Sinozolus Deuve 1997 dd Tribe Trechini Bonelli 1810Subtribe Aepina Fowler 1887Aepopsis Jeannel 1922 Aepus Leach in Samouelle 1819 Kenodactylus Broun 1909 Kiwitrechus Larochelle amp Lariviere 2007 Maoritrechus Brookes 1932 Neanops Britton 1962 Oarotrechus Townsend 2010 Temnostega Enderlein 1905 Thalassobius Solier 1849 Waiputrechus Townsend 2010 dd Subtribe Perileptina Sloane 1903Apoplotrechus Alluaud 1915 Neoblemus Jeannel 1923 Perileptus Schaum 1860 dd Subtribe Trechina Bonelli 1810Accoella Ueno 1990 Acheroniotes Lohaj amp Lakota 2010 Adriaphaenops Noesske 1928 Aepiblemus Belousov amp Kabak 1993 Agonotrechus Jeannel 1923 Agostinia Jeannel 1928 Alanorites Belousov 1998 Albanotrechus Casale amp V B Gueorguiev 1994 Allegrettia Jeannel 1928 Allotrechiama Ueno 1970 Ameroduvalius Valentine 1952 Anchotrechus Jeannel 1927 Andinorites Mateu amp Belles 1980 Andinotrechus Mateu 1981 Anillidius Jeannel 1928 Anophthalmus Sturm 1844 Aphaenopidius G Muller 1913 Aphaenops Bonvouloir 1862 Aphaenopsis G Muller 1913 Apocimmerites Belousov 1998 Apoduvalius Jeannel 1953 Aputrechisibus Trezzi 2007 Arctaphaenops Meixner 1925 Aspidaphaenops Ueno 2006 Austrotrechus B Moore 1972 Awatrechus Ueno 1955 Balazucellus Deuve 2001 Bathytrechus Ueno 2005 Beronaphaenops B V Gueorguiev 2012 Birmaphaenops Deuve 2017 Blemus Dejean 1821 Boldoriella Jeannel 1928 Boreaphaenops Ueno 2002 Bothynotrechus B Moore 1972 Casaleaphaenops Tian Huang amp Ma 2021 Cathaiaphaenops Deuve 1996 Caucasaphaenops Belousov 1999 Caucasorites Belousov amp Zamotajlov 1997 Chaetoduvalius Jeannel 1928 Chaetotrechiama Ueno 1982 Chiapadytes Vigna Taglianti 1977 Chu Tian amp He 2020 Cimbrodytes Piva 2021 Cimmerites Jeannel 1928 Cimmeritodes Deuve 1996 Ckacus Tian amp Huang 2021 Columbitrechus Mateu 1982 Coreoblemus Ueno 1969 Croatotrechus Casale amp Jalzic 1999 Dabatrechus Ueno 2004 Dactylotrechus Belousov amp Kabak 2003 Daiconotrechus Ueno 1971 Dalmataphaenops Monguzzi 1993 Darlingtonea Valentine 1952 Derossiella Queinnec 2008 Deuveaphaenops Tian amp Huang 2017 Dianotrechus Tian 2016 Doderotrechus Vigna Taglianti 1968 Dongoblemus Deuve amp Tian 2016 Dongodytes Deuve 1993 Dracotrechus Ueno 2010 Duvalioblemus Deuve 1995 Duvaliomimus Jeannel 1928 Duvaliopsis Jeannel 1928 Duvalius Delarouzee 1859 Eocnides Jeannel 1954 Epaphiopsis Ueno 1953 Epaphiotrechus Deuve amp Kavanaugh 2016 Epaphius Leach in Samouelle 1819 Erebotrechus Britton 1964 Escolatrechus Mateu 2003 Eutrechopsis B Moore 1972 Eutrechus B Moore 1972 Geotrechus Jeannel 1919 Giraffaphaenops Deuve 2002 Glabroduvalius Vrbica Curcic Antic amp Curcic 2013 Goedetrechus B Moore 1972 Gotoblemus Ueno 1970 Graciliblemus Deuve amp Tian 2016 Guatemalatrechus Trezzi 2003 Guiaphaenops Deuve 2002 Guizhaphaenops Vigna Taglianti 1997 Guizhaphaenopsoides Tian Huang amp Li 2021 Gulaphaenops Ueno 1987 Himalaphaenops Ueno 1980 Himalotrechus Belousov Kabak amp J Schmidt 2019 Homaloderodes Jeannel 1962 Huoyanodytes Tian amp Huang 2016 Iberotrechus Jeannel 1920 Iga Ueno 1953 Incatrechus Mateu amp Belles 1982 Inotrechus Dolzhanski amp Ljovushkin 1989 Ishidatrechus Ueno 1956 Ishikawatrechus Habu 1950 Italaphaenops Ghidini 1964 Iyotrechus Ueno amp Naito 2009 Jalzicaphaenops Lohaj amp Lakota 2010 Jeannelius Kurnakov 1959 Jiangxiaphaenops Ueno amp Clarke 2007 Jiulongotrechus Tian Huang amp Wang 2015 Junaphaenops Ueno 1997 Junnanotrechus Ueno amp Yin 1993 Kettlotrechus Townsend 2010 Kosswigia Jeannel 1947 Kozlovites Jeannel 1935 Kupetrechus Larochelle amp Lariviere 2007 Kurasawatrechus Yoshida amp Nomura 1952 Kusumia Ueno 1952 Lanxangaphaenops Deuve 2012 Laoblemus Ueno 2006 Laosaphaenops Deuve 2000 Lessinodytes Vigna Taglianti 1982 Libotrechus Ueno 1998 Luoxiaotrechus Tian amp Yin 2013 Luyatrechus M Etonti amp Mateu 2000 Luzonotrechus Ueno 1979 Mamesdytes Trezzi 2003 Masuzoa Ueno 1960 Masuzonoblemus Ueno 1989 Mayaphaenops Vigna Taglianti 1977 Meganophthalmus Kurnakov 1959 Mexaphaenops Bolivar y Pieltain 1942 Mexitrechus Barr 1982 Miaotrechus Tian Chen amp Ma 2020 Microblemus Ueno 2007 Mimanillus B Moore 1972 Mimotrechus B Moore 1972 Minimaphaenops Deuve 2000 Minosaphaenops Queinnec 2008 Minutotrechus Deuve amp Kavanaugh 2016 Nannotrechus Winkler 1926 Nanotrechiama Belousov amp Kabak 2018 Neaphaenops Jeannel 1920 Nelsonites Valentine 1952 Neotrechus G Muller 1913 Nesaeoduvalius Casale amp Giachino 2016 Nesiotrechus Ueno 1995 Nipponaphaenops Ueno 1971 Nototrechus B Moore 1972 Nunbergites Pawlowski amp Stachowiak 1991 Omalodera Blanchard 1842 Oodinotrechus Ueno 1998 Oroblemites Ueno amp Pawlowski 1981 Oroblemus Ueno amp Yoshida 1966 Orotrechus G Muller 1913 Oxytrechus Jeannel 1927 Panaphaenops Tian Huang amp Ma 2021 Paracimmerites Belousov 1998 Paraphaenops Jeannel 1916 Paratrechiotes Ueno 1995 Paratrechus Jeannel 1920 Petraphaenops Delic Kapla amp Colla 2019 Pheggomisetes Knirsch 1923 Pilosaphaenops Deuve amp Tian 2008 Plesioaphaenops Deuve amp Tian 2011 Pogonoschema Jeannel 1927 Pontodytes Casale amp Giachino 1989 Porocimmerites Belousov 1998 Protrechiama Belousov amp Kabak 2003 Pseudanophthalmus Jeannel 1920 Pseudaphaenops Winkler 1912 Pseudocnides Jeannel 1927 Pseudotrechisibus Mateu amp Belles 1982 Puertrechus Belousov amp Kabak 2014 Putzeysius Jeannel 1962 Qianaphaenops Ueno 2000 Qianotrechus Ueno 2000 Queinnectrechus Deuve 1992 Rakantrechus Ueno 1951 Ryugadous Habu 1950 Sardaphaenops Cerruti amp Henrot 1956 Satotrechus Ueno 2006 Sbordoniella Vigna Taglianti 1980 Scotoplanetes Absolon 1913 Scototrechus Britton 1962 Shenaphaenops Ueno 1999 Shennongotrechus Ueno 2004 Shenoblemus Tian amp Fang 2020 Shilinotrechus Ueno 2003 Shiqianaphaenops Tian 2016 Shuangheaphaenops Tian 2017 Shuaphaenops Ueno 1999 Sichuanotrechus Deuve 2005 Sidublemus Tian amp Yin 2013 Sinaphaenops Ueno amp Wang 1991 Sinocimmerites Deuve 2007 Sinotrechiama Ueno 2000 Sinotroglodytes Deuve 1996 Sloanella Jeannel 1927 Speotrechus Jeannel 1922 Stevensius Jeannel 1923 Stygiotrechus Ueno 1958 Subilsia Espanol 1967 Sumatrechus Deuve 2005 Superbotrechus Deuve amp Tian 2009 Suzuka Ueno 1956 Taiwanotrechus Ueno 1987 Taniatrechus Belousov amp Dolzhanski 1994 Tasmanorites Jeannel 1927 Tasmanotrechus B Moore 1972 Taurocimmerites Belousov 1998 Thalassoduvalius Ueno 1956 Tianeotrechus Tian amp Tang 2016 Tianzhuaphaenops Zhao amp Tian 2016 Tienmutrechus Suenson 1957 Tonkinaphaenops Deuve 2013 Toshiaphaenops Ueno 1999 Trechepaphiama Deuve amp Kavanaugh 2016 Trechepaphiopsis Deuve amp Kavanaugh 2016 Trechiama Jeannel 1927 Trechiamiotes Deuve 1998 Trechicomimus Mateu amp Negre 1972 Trechiella Jeannel 1927 Trechimorphus Jeannel 1927 Trechinotus Jeannel 1962 Trechiotes Jeannel 1954 Trechisibus Motschulsky 1862 Trechistus B Moore 1972 Trechoblemus Ganglbauer 1891 Trechus Clairville 1806 Trichaphaenops Jeannel 1916 Troglocimmerites Ljovuschkin 1970 Tropidotrechus Jeannel 1927 Typhlotrechus G Muller 1913 Uenoaphaenops Tian amp He 2020 Uenoites Belousov amp Kabak 2016 Uenotrechus Deuve amp Tian 1999 Ushijimaella Ueno 1980 Velebitaphaenops Casale amp Jalzic 2012 Velesaphaenops S Curcic et al 2018 Vietotrechus Ueno 1995 Wanhuaphaenops Tian amp Wang 2016 Wanoblemus Tian amp Fang 2016 Wulongoblemus Ueno 2007 Xenotrechus Barr amp Krekeler 1967 Xiangxius Tian Huang amp Li 2021 Xuedytes Tian amp Huang 2017 Yalongaphaenops Belousov amp Kabak 2021 Yamautidius Ueno 1957 Yanzaphaenops Ueno 2010 Yunotrechus Tian amp Huang 2014 Zhijinaphaenops Ueno amp Ran 2002 dd Subtribe Trechodina Jeannel 1926Amblystogenium Enderlein 1905 Canarobius Machado 1987 Cnides Motschulsky 1862 Cothresia Jeannel 1964 Cyphotrechodes Jeannel 1926 Eotrechodes Ueno Lafer amp Sundukov 1995 Himalotrechodes Ueno 1981 Iberotrechodes Faille et al 2021 Pachydesus Motschulsky 1864 Pachytrechodes Jeannel 1960 Paratrechodes Jeannel 1926 Scaurotrechodes Geginat 2006 Spelaeovulcania Machado 1987 Sporades Fauvel 1882 Thalassophilus Wollaston 1854 Trechobembix Jeannel 1926 Trechodes Blackburn 1901 Trechosia Jeannel 1926 Trechosiella Jeannel 1960 dd Antarctotrechus Ashworth amp Erwin 2016 Trechinites Heer 1862 Trechoides Motschulsky 1856 dd Tribe Zolini Sharp 1886Subtribe Merizodontina Sloane 1920Idacarabus Lea 1910 Maungazolus Larochelle amp Lariviere 2017 Merizodus Solier 1849 Percodermus Sloane 1920 Pseudoopterus Csiki 1928 Pterocyrtus Sloane 1920 Sloaneana Csiki 1933 Synteratus Broun 1909 Thayerella Baehr 2016 dd Subtribe Zolina Sharp 1886Oopterus Guerin Meneville 1841 Zolus Sharp 1886 dd dd Subfamily Xenaroswellianinae Erwin 2007 edit Xenaroswelliana Erwin 2007Subfamily Conjunctiinae Ponomarenko 1977 edit Conjunctia Ponomarenko 1977Subfamily Protorabinae Ponomarenko 1977 edit Aethocarabus Ren Lu amp Guo 1995 Atrirabus Hong amp Wang 1990 Cordorabus Ponomarenko 1977 Cretorabus Ponomarenko 1977 Denudirabus Ren Lu amp Guo 1995 Lirabus Hong 1992 Lithorabus Ponomarenko 1977 Magnirabus Hong amp Wang 1990 Mesorabus Ponomarenko 1977 Nebrorabus Ponomarenko 1989 Ovrabites Ponomarenko 1977 Penecupes Ren 1995 Protorabus Ponomarenko 1977 Not Assigned to Subfamily Agatoides Motschulsky 1856 Amphoxyne Bode 1953 Carabites Heer 1852 Cavicarabus Hong 1991 Conexicoxa Lin 1986 Cymatopterus Lomnicki 1894 Fangshania Hong 1981 Glenopterus Heer 1847 Hebeicarabus Hong 1983 Megacarabus Hong 1983 Meileyingia Hong 1987 Miocarabus Hong 1983 Neothanes Scudder 1890 Procarabus Oppenheim 1888 Prosynactus Bode 1953 Shanwangicarabus Hong 1985 Sinis Heer 1862 Sinocalosoma Hong amp Wang 1986 Sinocaralosoma Hong 1984 Sunocarabus Hong 1987 Tauredon Handlirsch 1910 Wuchangicarabus Hong 1991 Xishanocarabus Hong 1984 Yunnanocarabus Lin 1977References edit Lorenz Wolfgang 2021 Carabcat Database doi 10 48580 dfqf 3dk Retrieved 2023 03 04 Carabidae Latreille 1802 Catalogue of Life Retrieved 2023 03 04 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