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Dryandra subg. Hemiclidia

Dryandra subg. Hemiclidia is an obsolete plant taxon that encompassed material that is now included in Banksia. Published at genus rank as Hemiclidia by Robert Brown in 1830, it was set aside by George Bentham in 1870, but reinstated at subgenus rank by Alex George in 1996. In 2007, all Dryandra species were transferred into Banksia at series rank, and the infrageneric Dryandra taxa, including D. subg. Hemiclidia, were set aside.

Banksia falcata as figured in Plate 1455 of Edwards's Botanical Register (Volume 17, 1831). This is the type species of Hemiclidia under the name Hemiclidia baxteri, and the type species of Dryandra subg. Hemiclidia under the name Dryandra falcata.

According to Brown edit

In 1810, Robert Brown published the genus Dryandra in his On the Proteaceae of Jussieu. Thirteen species were published, including Dryandra falcata (now Banksia falcata), but no infrageneric arrangement was proffered.[1] Twenty years later, Brown published a further eleven species and the first infrageneric arrangement in his Supplementum primum prodromi florae Novae Hollandiae. By this time, Brown had observed the tendency in D. falcata for one of the two ovules in each follicle to abort, thereafter developing into a winglike appendage to the seed separator. On this basis, he transferred the species into a monotypic genus, which he named Hemiclidia,[2] from the Greek hemi ("half") and perhaps kleidos ("barrier", "means of closing").[3]

Hemiclidia was retained in the 1856 arrangement of Carl Meissner,[4] but discarded by George Bentham in his 1870 revision of the genus. Bentham correctly observed that the abortion of seeds occurs in other Dryandra species, and is a diagnostically unimportant character:

"as far as I can understand the characters given, the difference in the fruit upon which the genus Hemiclidia was founded is merely the result of the abortion of one ovule, which occurs occasionally or perhaps constantly in one or two other species of Dryandra."[5]

According to George edit

In 1996, Alex George published a thorough revision of Dryandra in preparation of his monograph on that genus for the Flora of Australia series. He segregated D. falcata and the newly published D. glauca (now Banksia glaucifolia) into a subgenus based on their unusual follicles, which are small, hairy and pliable. Needing a name for the subgenus, George noted that Hemiclidia was available, and had an appropriate type species in D. falcata. He therefore demoted Hemiclidia to subgenus rank as Dryandra subg. Hemiclidia (R.Br.) A.S.George.[6] Its placement and circumscription in George's taxonomic arrangement of Dryandra may be summarised as follows:[6][7]

Genus Dryandra (now Banksia ser. Dryandra)
Subgenus Dryandra (24 series, 91 species, 39 subspecies, 23 varieties)
Subgenus Hemiclidia
D. falcata (now Banksia falcata)
D. glauca (now Banksia glaucifolia)
Subgenus Diplophragma (1 species)

This arrangement remained current until 2007, when Austin Mast and Kevin Thiele transferred Dryandra into Banksia. They were not yet ready, however, to tender an infrageneric arrangement encompassing Dryandra, so as an interim measure they transferred Dryandra into Banksia at series rank. This minimised the nomenclatural disruption of the transfer, but also caused George's rich infrageneric arrangement to be set aside. Thus under the interim arrangements implemented by Mast and Thiele, D. subg. Hemiclidia has been set aside.[8]

References edit

  1. ^ Brown, Robert (1810). "On the Proteaceae of Jussieu". Transactions of the Linnean Society of London. 10: 15–226. doi:10.1111/j.1096-3642.1810.tb00013.x.
  2. ^ Brown, Robert (1830). "Dryandra". Supplementum Primum Prodromi Florae Novae Hollandiae. London: Taylor. pp. 37–40.
  3. ^ Cavanagh, Tony; Pieroni, Margaret (2006). The Dryandras. Melbourne: Australian Plants Society (SGAP Victoria); Perth: Wildflower Society of Western Australia. ISBN 1-876473-54-1.
  4. ^ Meissner, Carl (1856). "XLV. Hemiclidia". In de Candolle, A. P. (ed.). Prodromus systematis naturalis regni vegetabilis. Vol. XIV. Paris: Sumptibus Sociorum Treuttel et Wurtz. pp. 481–482.
  5. ^ Bentham, George (1870). "Dryandra". Flora Australiensis. Vol. V. London: L. Reeve & Co. pp. 562–584.
  6. ^ a b George, Alex S. (1996). "New taxa and a new infragenetic classification in Dryandra R.Br. (Proteaceae: Grevilleoideae)". Nuytsia. 10 (3): 313–408. doi:10.58828/nuy00235. S2CID 92008567.
  7. ^ George, Alex S. (1999). "Banksia". In Wilson, Annette (ed.). Flora of Australia Volume 17B: Proteaceae 3: Hakea to Dryandra. Collingwood, Victoria: CSIRO Publishing / Australian Biological Resources Study. pp. 175–251. ISBN 0-643-06454-0.
  8. ^ Mast, Austin R.; Thiele, Kevin (2007). "The transfer of Dryandra R.Br. to Banksia L.f. (Proteaceae)". Australian Systematic Botany. 20: 63–71. doi:10.1071/SB06016.

External links edit

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Dryandra subg Hemiclidia is an obsolete plant taxon that encompassed material that is now included in Banksia Published at genus rank as Hemiclidia by Robert Brown in 1830 it was set aside by George Bentham in 1870 but reinstated at subgenus rank by Alex George in 1996 In 2007 all Dryandra species were transferred into Banksia at series rank and the infrageneric Dryandra taxa including D subg Hemiclidia were set aside Banksia falcata as figured in Plate 1455 of Edwards s Botanical Register Volume 17 1831 This is the type species of Hemiclidia under the name Hemiclidia baxteri and the type species of Dryandra subg Hemiclidia under the name Dryandra falcata Contents 1 According to Brown 2 According to George 3 References 4 External linksAccording to Brown editIn 1810 Robert Brown published the genus Dryandra in his On the Proteaceae of Jussieu Thirteen species were published including Dryandra falcata now Banksia falcata but no infrageneric arrangement was proffered 1 Twenty years later Brown published a further eleven species and the first infrageneric arrangement in his Supplementum primum prodromi florae Novae Hollandiae By this time Brown had observed the tendency in D falcata for one of the two ovules in each follicle to abort thereafter developing into a winglike appendage to the seed separator On this basis he transferred the species into a monotypic genus which he named Hemiclidia 2 from the Greek hemi half and perhaps kleidos barrier means of closing 3 Hemiclidia was retained in the 1856 arrangement of Carl Meissner 4 but discarded by George Bentham in his 1870 revision of the genus Bentham correctly observed that the abortion of seeds occurs in other Dryandra species and is a diagnostically unimportant character as far as I can understand the characters given the difference in the fruit upon which the genus Hemiclidia was founded is merely the result of the abortion of one ovule which occurs occasionally or perhaps constantly in one or two other species of Dryandra 5 According to George editIn 1996 Alex George published a thorough revision of Dryandra in preparation of his monograph on that genus for the Flora of Australia series He segregated D falcata and the newly published D glauca now Banksia glaucifolia into a subgenus based on their unusual follicles which are small hairy and pliable Needing a name for the subgenus George noted that Hemiclidia was available and had an appropriate type species in D falcata He therefore demoted Hemiclidia to subgenus rank as Dryandra subg Hemiclidia R Br A S George 6 Its placement and circumscription in George s taxonomic arrangement of Dryandra may be summarised as follows 6 7 Genus Dryandra now Banksia ser Dryandra Subgenus Dryandra 24 series 91 species 39 subspecies 23 varieties Subgenus HemiclidiaD falcata now Banksia falcata D glauca now Banksia glaucifolia dd Subgenus Diplophragma 1 species dd This arrangement remained current until 2007 when Austin Mast and Kevin Thiele transferred Dryandra into Banksia They were not yet ready however to tender an infrageneric arrangement encompassing Dryandra so as an interim measure they transferred Dryandra into Banksia at series rank This minimised the nomenclatural disruption of the transfer but also caused George s rich infrageneric arrangement to be set aside Thus under the interim arrangements implemented by Mast and Thiele D subg Hemiclidia has been set aside 8 References edit Brown Robert 1810 On the Proteaceae of Jussieu Transactions of the Linnean Society of London 10 15 226 doi 10 1111 j 1096 3642 1810 tb00013 x Brown Robert 1830 Dryandra Supplementum Primum Prodromi Florae Novae Hollandiae London Taylor pp 37 40 Cavanagh Tony Pieroni Margaret 2006 The Dryandras Melbourne Australian Plants Society SGAP Victoria Perth Wildflower Society of Western Australia ISBN 1 876473 54 1 Meissner Carl 1856 XLV Hemiclidia In de Candolle A P ed Prodromus systematis naturalis regni vegetabilis Vol XIV Paris Sumptibus Sociorum Treuttel et Wurtz pp 481 482 Bentham George 1870 Dryandra Flora Australiensis Vol V London L Reeve amp Co pp 562 584 a b George Alex S 1996 New taxa and a new infragenetic classification in Dryandra R Br Proteaceae Grevilleoideae Nuytsia 10 3 313 408 doi 10 58828 nuy00235 S2CID 92008567 George Alex S 1999 Banksia In Wilson Annette ed Flora of Australia Volume 17B Proteaceae 3 Hakea to Dryandra Collingwood Victoria CSIRO Publishing Australian Biological Resources Study pp 175 251 ISBN 0 643 06454 0 Mast Austin R Thiele Kevin 2007 The transfer of Dryandra R Br to Banksia L f Proteaceae Australian Systematic Botany 20 63 71 doi 10 1071 SB06016 External links edit Dryandra subg Hemiclidia R Br A S George Flora of Australia Online Department of the Environment and Heritage Australian Government nbsp Hemiclidia R Br Australian Plant Name Index APNI IBIS database Centre for Plant Biodiversity Research Australian Government Retrieved from https en wikipedia org w index php title Dryandra subg Hemiclidia amp oldid 1156672307, wikipedia, wiki, book, books, library,

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