"Horseshoe shrimp" redirects here. For a species in this class known as the horseshoe shrimp, see Hutchinsoniella. For the horseshoe shrimp better known as tadpole shrimp or shield shrimp, see Notostraca.
The Cephalocarida are a class in the subphylumCrustacea comprising only 12 species. Both the nauplii and the adults are benthic.[1] They were discovered in 1955 by Howard L. Sanders,[2] and are commonly referred to as horseshoe shrimp. They have been grouped together with the Remipedia in the Xenocarida. Although a second family, Lightiellidae, is sometimes used, all cephalocaridans are generally considered to belong in just one family: Hutchinsoniellidae. Fossil records of cephalocaridans has been found in 462 million year old deposits.[3]
These are hermaphroditic and pigmentless crustaceans with an elongated and translucent body that measures 2 to 4 mm (0.079 to 0.157 in) in length. A heart is present, and their exopods and pseudepipodites appears to be used for gas exchange.[5][6] They have a large head, the hind edge of which covers the first thoracic segment. The thorax consists of nine limb-bearing segments (thoracic limb VIII absent in Lightiella), followed by 10 limbless abdominal segments and a telson. In the larva, all the trunk segments are ring-shaped, but more dorsoventrally flattened than in the adults. During growth the anterior segments turns into the thorax and the posterior segments which makes up the abdomen remains ring-shaped.[7] No eyes have been observed in either the adult or larval stages, presumably because of their muddy natural habitat. The second pair of antennae is located behind the mouth; in all other crustaceans the antennae are in front of the mouth at the adult stage, and only their larvae have antennae that have the same location as adult cephalocaridans.[8][9]
The mouth is located behind the large upper lip, flanked by mandibles. The first pair of maxillae is very small, and the second pair has the same structure as the following thoracic legs: a large basal part, equipped with outgrowths on the inner side, used in locomotion, a forked inner branch and two outer lobes - referred to as the "pseudoepipod" and the "exopod". The structural and functional similarity between the maxillae and the legs may be a sign of primitive organization; the maxillae are not specialized, as they are in other crustaceans.[8]
Ecologyedit
Cephalocaridans are found from the intertidal zone down to a depth of 1,500 m (4,900 ft), in all kinds of sediments. Cephalocaridans feed on marine detritus. To bring in food particles, they generate currents with the thoracic appendages like the branchiopods and the malacostracans. Food particles are then passed anteriorly along a ventral groove, leading to the mouthparts.[10]
Referencesedit
^Key for the identification of crustacean nauplii - GfBS
^ abRobert D. Barnes (1982). Invertebrate Zoology. Philadelphia, PA: Holt-Saunders International. p. 672. ISBN978-0-03-056747-6.
^Rediscovery of the horseshoe shrimp Lightiella serendipita Jones, 1961 (Cephalocarida: Hutchinsoniellidae) in San Francisco Bay, California, USA, with a key to the worldwide species of Cephalocarida
^L. A. Zenkevich. "Phylum Arthropoda". The Animal Life [Zhizn' Zhivotnykh]. Vol. 2.
Cephalocarida- University of California Museum of Paleontology
Data related to Cephalocarida at Wikispecies
February 15, 2024
cephalocarida, horseshoe, shrimp, redirects, here, species, this, class, known, horseshoe, shrimp, hutchinsoniella, horseshoe, shrimp, better, known, tadpole, shrimp, shield, shrimp, notostraca, class, subphylum, crustacea, comprising, only, species, both, nau. Horseshoe shrimp redirects here For a species in this class known as the horseshoe shrimp see Hutchinsoniella For the horseshoe shrimp better known as tadpole shrimp or shield shrimp see Notostraca The Cephalocarida are a class in the subphylum Crustacea comprising only 12 species Both the nauplii and the adults are benthic 1 They were discovered in 1955 by Howard L Sanders 2 and are commonly referred to as horseshoe shrimp They have been grouped together with the Remipedia in the Xenocarida Although a second family Lightiellidae is sometimes used all cephalocaridans are generally considered to belong in just one family Hutchinsoniellidae Fossil records of cephalocaridans has been found in 462 million year old deposits 3 CephalocaridaHutchinsoniella macracanthaScientific classificationDomain EukaryotaKingdom AnimaliaPhylum ArthropodaClass CephalocaridaSanders 1955Order BrachypodaBirshteyn 1960Family HutchinsoniellidaeSanders 1955GeneraChiltoniella Hampsonellus Hutchinsoniella Lightiella SandersiellaSynonymsLightiellidae Jones 1961 Contents 1 Taxonomy 2 Description and anatomy 3 Ecology 4 References 5 External linksTaxonomy editClass Cephalocarida Sanders 1955 4 Order Brachypoda Birshteyn 1960 Family Hutchinsoniellidae Sanders 1955 Genus Chiltoniella Knox amp Fenwick 1977 Chiltoniella elongata Knox amp Fenwick 1977 Genus Hampsonellus Hessler amp Wakabara 2000 Hampsonellus brasiliensis Hessler amp Wakabara 2000 Genus Hutchinsoniella Sanders 1955 Hutchinsoniella macracantha Sanders 1955 Genus Lightiella Jones 1961 Lightiella floridana McLaughlin 1976 Lightiella incisa Gooding 1963 Lightiella magdalenina Carcupino et al 2006 Lightiella monniotae Cals amp Delamare Deboutteville 1970 Lightiella serendipita Jones 1961 Genus Sandersiella Shiino 1965 Sandersiella acuminata Shiino 1965 Sandersiella bathyalis Hessler amp Sanders 1973 Sandersiella calmani Hessler amp Sanders 1973 Sandersiella kikuchii Shimomura amp Akiyama 2008Description and anatomy editThese are hermaphroditic and pigmentless crustaceans with an elongated and translucent body that measures 2 to 4 mm 0 079 to 0 157 in in length A heart is present and their exopods and pseudepipodites appears to be used for gas exchange 5 6 They have a large head the hind edge of which covers the first thoracic segment The thorax consists of nine limb bearing segments thoracic limb VIII absent in Lightiella followed by 10 limbless abdominal segments and a telson In the larva all the trunk segments are ring shaped but more dorsoventrally flattened than in the adults During growth the anterior segments turns into the thorax and the posterior segments which makes up the abdomen remains ring shaped 7 No eyes have been observed in either the adult or larval stages presumably because of their muddy natural habitat The second pair of antennae is located behind the mouth in all other crustaceans the antennae are in front of the mouth at the adult stage and only their larvae have antennae that have the same location as adult cephalocaridans 8 9 The mouth is located behind the large upper lip flanked by mandibles The first pair of maxillae is very small and the second pair has the same structure as the following thoracic legs a large basal part equipped with outgrowths on the inner side used in locomotion a forked inner branch and two outer lobes referred to as the pseudoepipod and the exopod The structural and functional similarity between the maxillae and the legs may be a sign of primitive organization the maxillae are not specialized as they are in other crustaceans 8 Ecology editCephalocaridans are found from the intertidal zone down to a depth of 1 500 m 4 900 ft in all kinds of sediments Cephalocaridans feed on marine detritus To bring in food particles they generate currents with the thoracic appendages like the branchiopods and the malacostracans Food particles are then passed anteriorly along a ventral groove leading to the mouthparts 10 References edit Key for the identification of crustacean nauplii GfBS Howard L Sanders 1955 The Cephalocarida a new subclass of Crustacea from Long Island Sound Proceedings of the National Academy of Sciences 41 1 61 66 Bibcode 1955PNAS 41 61S doi 10 1073 pnas 41 1 61 JSTOR 89010 PMC 528024 PMID 16589618 Welsh fossils uncover an ancient marine dwarf world Boxshall G 2010 Hutchinsoniellidae Sanders 1955 World Register of Marine Species Retrieved 2017 12 07 Class Cephalocarida Sanders 19551 in Brill Functional Morphology and Diversity Atlas of Crustacean Larvae a b Robert D Barnes 1982 Invertebrate Zoology Philadelphia PA Holt Saunders International p 672 ISBN 978 0 03 056747 6 Rediscovery of the horseshoe shrimp Lightiella serendipita Jones 1961 Cephalocarida Hutchinsoniellidae in San Francisco Bay California USA with a key to the worldwide species of Cephalocarida L A Zenkevich Phylum Arthropoda The Animal Life Zhizn Zhivotnykh Vol 2 External links edit nbsp Crustaceans portalCephalocarida University of California Museum of Paleontology nbsp Data related to Cephalocarida at Wikispecies Retrieved from https en wikipedia org w index php title Cephalocarida amp oldid 1176932750, wikipedia, wiki, book, books, library,
