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Kasakela chimpanzee community

The Kasekela chimpanzee community (formerly spelled Kasakela[1]) is a habituated community of wild eastern chimpanzees that lives in Gombe National Park near Lake Tanganyika in Tanzania.[2] The community was the subject of Jane Goodall's pioneering study that began in 1960, and studies have continued ever since, becoming the longest continuous study of any animals in their natural habitat.[2][3][4] As a result, the community has been instrumental in the study of chimpanzees and has been popularized in several books and documentaries. The community's popularity was enhanced by Goodall's practice of giving names to the chimpanzees she was observing, in contrast to the typical scientific practice of identifying the subjects by number. Goodall generally used a naming convention in which infants were given names starting with the same letter as their mother, allowing the recognition of matrilineal lines.

One of the most important discoveries that was learned by observing the Kasekela chimpanzee community was the use of tools. On November 4, 1960, Goodall observed a chimpanzee that she had named David Greybeard using a grass stalk as a tool to extract termites from a termite hill. Later, she observed David Greybeard and another chimpanzee named Goliath stripping leaves off twigs to create termite fishing tools. Previously, tool use in chimpanzees was only rarely observed, and tool creation by non-human animals had never been observed. Until then, tool making was considered one of the defining characteristics of being human.[5][6][7] Another important observation occurred a few days earlier, on October 30, 1960. On that day Goodall observed the community's chimpanzees eating meat, dispelling the notion that chimpanzees are vegetarians.[7] A third observation by Goodall in the early 1960s was that male chimpanzees perform a "rain dance," charging, calling, slapping the ground and trees and dragging branches in the rain.[8][9] In the early 1970s the chimpanzees of the community were observed to engage in ongoing coordinated attacks against the chimpanzees of the neighbouring Kahama Chimpanzee Community, ultimately wiping it out.[10][11] According to historian Ian Morris, this "Four Year War" represented the first time scientists had observed chimpanzees "deliberately seek out, attack and leave for dead" chimps from another community, and it has been described as "the first record of lasting 'warfare' among [non-human] primates."[10][12]

Several families within the Kasekela chimpanzee community have been particularly prominent in books and documentaries. The F-family has produced five alpha males for the community, and the matriarch, Flo, played a particularly important role in acknowledging Goodall's acceptance as a human observer by the community. The G-family has produced at least one alpha male, and also the birth of several twins, which are rare among chimpanzees. There are other families as well which include the T-family and S-family (which has produced one alpha male).

F-family edit

Flo edit

Flo (c.1919 – 1972)[13] was the matriarch of the F-family, so named because she and all her matrilineal descendants were given names beginning with the letter "F". In 1962, Flo was one of the first chimpanzees to approach Goodall's camp, along with her infant daughter Fifi. Video of Flo approaching Goodall and allowing Fifi to reach out to touch Goodall's forehead, letting Goodall know she had been accepted, is shown in the IMAX film Jane Goodall's Wild Chimpanzees.[14] Later she brought her sons Figan and Faben, who would later become prominent members of the community to the camp, and when she came into estrus in 1963 she attracted the community's males to Goodall's camp.[14][15]

Flo was a high-ranking female who had at least five known offspring, three sons (Faben, Figan and Flint) and two daughters (Fifi and Flame). Flo was 28 years old when she gave birth to Faben. Most female chimpanzees have their first infant when 12–14 years of age, so it is possible that Flo had at least one infant prior to Faben that either died or, if it was a female, would have transferred to one of the other communities prior to Goodall's arrival to Gombe in 1960. When Flo died in 1972, she was given an obituary in Britain's Sunday Times.[16]

Faben edit

Faben (c.1947 – 1975)[13] was Flo's oldest known offspring.[17] He was a powerful male, but lost the use of one arm in a polio epidemic that ravaged the community in 1966.[17] Despite his disability, he learned to create spectacular bipedal charging displays, which allowed him to regain a high rank within the community.[17] His support was critical to Humphrey's ascension to alpha male rank in 1969, and to his brother Figan's ascension to alpha male in 1972.[18][19] Faben also participated in the conflicts with the rival Kahama community (which was formed by former Kasekela members) that occurred between 1973 and 1975, and was a leader in the killing of several Kahama males, including former Kasekela alpha male Goliath.[17][20] Faben himself died in 1975, possibly as a result of further conflicts with the Kahama community.[17] Following Faben's death, Figan had difficulty maintaining his alpha male position for several months and eventually lost his alpha status to Goblin.[18]

Figan edit

Figan (c.1953 – 1982)[13] was Flo's second son.[21] Goodall considered him to be Gombe's most intelligent chimpanzee.[22] Although younger than Faben, he was able to dominate his older brother after Faben's arm was paralyzed from polio in 1966.[21] With Faben's support, Figan was able to become alpha male in 1972 by defeating the prior alpha Humphrey and an older competitor Evered. At this time, Goodall described him as the most powerful alpha male in Gombe’s recorded history.[21] After Faben's death in 1975, Figan lost his unquestioned alpha status, in that he could be dominated by a coalition of males, but remained the top-ranked male.[21] By 1977, he had regained his alpha status by forging alliances with other males, including his predecessor Humphrey.[21] In 1979, he was toppled from his alpha status by 15-year-old Goblin, whom he had previously supported, but regained his alpha status once again by forging alliances with other males.[21] Although a powerful male, Figan's ability to make alliances with other males was instrumental to allow him to hold on to his alpha status.[23] In 1982, Goblin unseated him again, and Figan disappeared and presumably died a few months later, although when last seen he appeared to be in good health.[21] Although Figan was the alpha male for several years, he is not known for certain to have sired any infants, although he is likely the father of a few.[24]

Fifi edit

Fifi (c.1958 – 2004)[13][25][26] was Flo's oldest daughter.[27][28] Fifi benefited from her high-ranking family, and ascended to the top of the female hierarchy in her community.[29][27][28] Fifi had nine infants, seven of whom survived to independence.[27][28] Her two oldest sons, Freud and Frodo, both have become the community's alpha male, as did another son, Ferdinand. One of her other sons, Faustino, also attained high rank. Her three surviving daughters were Fanni, Flossi and Flirt.[27][28] She also had one son, Fred (1996–1997) and one daughter, Furaha (2002–2004) who died in infancy.[13][30] Fred was fathered by one of Fifi's other sons, Frodo.[31] Five-year-old Fifi was the focal point of the documentary People of the Forest: The Chimps of Gombe, in which her mother and siblings also featured, attempting to gain access to her infant brother Flint.[32] All four of Fifi's surviving sons had all held the alpha and/or beta positions in the community. Grandson Fudge (by Fanni) also attained alpha male status. Late in Frodo's reign as alpha male, Fifi helped defend him against attacks and she suffered some injuries in the process.[33]

Although males are perceived as the more aggressive sex in chimpanzees, studies at Gombe have revealed that aggression among females with each other does indeed occur.[34] One of two contexts for female-female aggressive encounters in Gombe was the presence of newborn infants.[34] Fifi grew to be a formidable female who, although a very successful mother herself, was observed to have carried out ferocious and infanticidal attacks on other, lower-ranking females in her community. Such attacks were observed of Fifi and her adult daughter, Fanni, aggressively harassing fellow Kasekela female, Gremlin, on three different occasions; the first being in 1993 following the birth of her daughter Gaia, and then five years later following the birth of her twins. This was followed by another attack on Gremlin six years later following the birth of her next baby.[34] Researchers could only speculate that the attacks on infants of other females had to do with food competition, as each female strives to secure resources for her own group of offspring and thus the expansion of another female's family unit could deplete resources.[29] The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom in the 1970s,[34] which proved that cannibalism, although deemed rare, does occur among chimpanzees, by both males and, in this particular case, jealous females.

In 2004, Fifi, who was by then 46 years old which made her the oldest living chimpanzee in the Kasekela community at that point in time, disappeared in September with her youngest and ninth offspring, two year old daughter Furaha.[35] During that year she shifted her range to the far north of the Kasekela community territory, and the simultaneous advancement of the neighbouring Mitumba community further south nearing the Kasekela territory border suggests that Fifi and her youngster may have been fatally attacked by the Mitumba territorial males. A female chimpanzee leaving the safety of her community territory poses risks, particularly if she has a vulnerable youngster. Studies on chimpanzee intercommunity relations revealed that chimpanzees are likely to be attacked by the territorial males of a rival community,[36] and the intensity of the attacks depends on the gender of the intruder, and if the stranger is female, it depends on whether or not they are of any reproductive benefit to the males. Young, cycling females are often spared an attack because they are deemed to be potential new mates, but a lactating mother with a young infant is considered of very little use for the males, which results in lethal aggression and fatality.[36] The bodies of Fifi and Furaha were never found, but the confirmed death of fellow Kasekela female, Patti, in 2005, by neighbouring Mitumba community males[37] provides a near definitive conclusion of the fate that befell Fifi and her youngster.

Flint edit

Flint (1964–1972) was Flo's third son, and her first infant born after Goodall began observing the community.[13] Goodall had intended that Flint become the first wild chimpanzee to have his life documented from birth to death.[3] Flint had difficulty weaning, and became distraught and angry when the birth of his sister Flame displaced him as Flo's baby.[38] After Flame disappeared, possibly by the hand of Flint, he regained his enthusiasm, but resumed his needy, infantlike behaviour.[38] After his nephew Freud was born, Flint became fascinated with the new infant and became an important influence of Freud's first year of life.[39] Following the death of his mother Flo in 1972, Flint became depressed and ill; he died about three weeks after her.[16]

Flame edit

Flame (1968–1969) was Flo's daughter.[13] Flo's son Flint, at age four, threw violent fits in order to suckle his mother's breast while newborn Flame was in Flo's arms. Flame disappeared shortly after she was born; there has been speculation she may have been killed by her jealous brother Flint.[40]

Freud edit

Freud (May 1971 – March 2014) was Fifi's oldest son.[41] His father was a male from outside the Kasekela community.[42] As a juvenile, he benefited from a high-ranking mother and an uncle, Figan, who was the alpha male.[41] In 1993, Freud became the alpha male by unseating his childhood playmate Wilkie.[41] While the alpha male, his younger but larger brother Frodo became the second highest-ranking male in the community. In 1997, while Freud was afflicted by sarcoptic mange, Frodo was able to take over the alpha status.[41][43][44] After losing his alpha status, Freud settled into a role as a middle-ranking male, although could still threaten higher-ranking males when aroused.[28][45] He was also able to serve as an important ally to other males.[43]

Although less aggressive than his brother Frodo, Freud is known to have engaged in at least one instance of infanticide, killing a Kasekela infant named Tofiki in 2004.[30] In his old age, his younger brothers sometimes provided him support, such as sharing meat and honey with him.[43] The documentary Chimpanzee Family Fortunes ends with a scene of the orphaned young female Flirt finding solace after several lonely months by getting groomed by her older brother Freud.[46] He fathered at least one infant, the female Cocoa to Candy, born in 2004[47][48][49] Freud's approach to maintaining his alpha status was different from his smaller predecessor Wilkie and his larger successor Frodo.[50] At 99 pounds (45 kg), Freud was larger than the 81-pound (37-kilogram) Wilkie and smaller than the 113-pound (51-kilogram) Frodo.[50] Freud was less aggressive than and groomed other males to help maintain alliances more often than Frodo, but engaged in more contact aggression and less grooming than Wilkie.[28][45][50] While alpha male he was able to intimidate even the larger Frodo with his displays.[44]

Freud died in March 2014, at more than 42 years old, which made him the oldest male whose age was known in the history of the community.[25][43] Freud was one of the first wild chimpanzees whose entire life from birth to old age was documented by researchers.[51]

Frodo edit

Frodo (June 30, 1976 – November 10, 2013) was Fifi's second-oldest son.[52] His father was the relatively low-ranking male Sherry. Even from a young age, Frodo was unusually large and aggressive. He learned to throw rocks as a juvenile and became very accurate with his aim; he sometimes threw them at his human observers, often hitting and bruising them.[53] As an adult, he was one of the largest chimpanzees ever observed in the community, at about 113 pounds (51 kg) and remained quite aggressive.[50][52] He also became an excellent hunter of red colobus monkeys, and was also able to intimidate other chimpanzees into sharing their kills with him if he was unsuccessful.[28][54] From 1990 to 1995, Frodo killed 10% of the red colobus monkeys within his home range.[33] His large size and aggressive nature allowed him to attain high status in the group, while his brother Freud was alpha male. On October 2, 1997, while Freud and other community members were suffering from sarcoptic mange, Frodo attacked and defeated Freud, thus attaining the alpha male position.[28][52][54][44] Researchers speculated that Frodo was unaffected by the mange because he interacted less with the other apes, giving the parasite less opportunity to infect him.[44] After taking over the alpha male position, Frodo demonstrated his new power by protecting his vanquished brother from the other males; typically the former alpha is attacked by the other males and forced to leave the community for a period of time.[55]

As alpha male, Frodo maintained his position largely through intimidation.[28][50][54] He rarely groomed other males, and often demanded that other males groom him.[28][50][54] Frodo remained alpha male until becoming ill himself in 2002;[28][46][54][56] he was then defeated by a coalition of several males and spent most of the next two years alone, recovering from his wounds and illness.[28][46][54][56] He received antibiotics from the researchers, which likely helped his recovery.[57] In 2004, he once again began travelling with the rest of the community, but was unable to re-establish his alpha status.[28][46][54] In his later years, Frodo mellowed. He became less aggressive and spent much time by himself, although he could still intimidate other chimpanzees on occasion.[58] Frodo died in 2013.[59]

Frodo's aggression was not limited to colobus monkeys and other chimpanzees. In May 2002, he killed a 14-month-old human child that the niece of a member of the research team had carried into his territory.[57] As a result, the Tanzanian National Parks Department considered killing Frodo.[57] In 1988, he attacked visiting Far Side cartoonist Gary Larson, leaving him bruised and scratched.[57] Frodo had a history of attacking the researchers observing him; Goodall was attacked by Frodo on multiple occasions and, in 1989, the ape beat her head so violently her neck was nearly broken.[57]

Frodo fathered at least eight infants, second only to group male Wilkie, who fathered ten.[59][60][61] Infants he fathered included:

  • Zeus (1993– ), by Trezia.[25] Zeus was part of a coalition that helped topple his uncle Ferdinand from alpha male status.[62]
  • Titan (1994– ), a large, aggressive adolescent male by Patti[25][63][64]
  • Fred (1996–1997), by Fifi, Frodo's own mother[25][31][42][33]
  • Golden and Glitter (1998– ), female twins and the oldest chimpanzee twins known to have survived in the wild, by Gremlin[25][31][46][44][56]
  • Tarzan (1999– ), Titan's brother by Patti[25]
  • Samwise (2001– ), an aggressive female by Sandi[25]
  • Sinbad (2001– ), Sparrow's youngest son[25]

Fanni edit

Fanni (b. March 19, 1981) is Fifi's oldest daughter.[13] Her father was Goblin.[25][42][65] She had her first infant, Fax, at just 11 years old, making her one of the youngest mothers seen in the community, although Fax did not survive to adulthood.[13][66] She has also had five other children, males Fudge (1996– ), Fundi (2000– ), Fifty (2010– ) and Ficma (2015–) and females Familia (2004– ) and Fadhila (2007– ).[67] Fudge, Fundi and Familia were fathered by Sheldon, who would later become alpha male.[25] Familia disappeared in 2012.[citation needed] Fadhila was fathered by Wilkie, another former alpha male.[25][42] Fifty was also sired by Wilkie.[48][49] As of 2021, Fudge is the alpha male of the community.[55]

Whilst most female chimpanzees leave their natal communities upon reaching sexual maturity,[68] Fanni has remained in her natal Kasekela community alongside her mother (in contrast to her younger sister Flossi). Fanni remained close to her mother Fifi, with whom she formed a close alliance against the other community females. Although males are perceived as the more aggressive sex in chimpanzees, studies at Gombe have revealed that there is substantial aggression among females with each other.[34] One of two contexts for female-female aggressive encounters in Gombe was the presence of newborn infants.[34] Fifi and Fanni were a formidable force and they carried out infanticidal attacks on other, lower-ranking females in her community. Such attacks were observed on fellow Kasekela female, Gremlin, on three different occasions; the first being in 1993 following the birth of her daughter Gaia, and then five years later following the birth of her twins. This was followed by another attack on Gremlin six years later following the birth of her next baby.[34] Researchers could only speculate that the attacks on infants of other females had to do with food competition, as each female strives to secure resources for her own group of offspring and thus the expansion of another female's family unit could deplete resources. The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom in the 1970s,[34] which proved that cannibalism, although deemed rare, does occur among chimpanzees, by both males and, in this particular case, jealous females.

The loss of her mother in 2004 greatly reduced Fanni's dominance and she shifted her range to the southern part of the Kasekela territory, further away from the other community females.[69]

Flossi edit

Flossi (b. February 8, 1985[70]) is Fifi's second-oldest daughter. Her father was Goblin.[47] Unlike most other female chimpanzees in the F-family, upon reaching adulthood she emigrated from the Kasekela community to join the neighbouring Mitumba Chimpanzee Community in 1996, although her younger sister Flirt (who was born after Flossi emigrated) later joined her.[25] Although it is typical for female chimpanzees to emigrate from their natal group upon reaching maturity, within the Kasekela community only about half the females do so. Flossi may have chosen to emigrate to avoid mating attempts from her brother Frodo.[55] Before emigrating from the Kasekela community, Flossi may have innovated the practice of fishing for ants to eat using a stick, which had not been observed before within the Kasekela community (although fishing for termites had been) but soon after the practice spread throughout the community.[71] Flossi has given birth to four offspring, the males Forest (1997– ) and Fansi (2001– ) and the females Flower (2005– ), and Falida (2009– ).[72] Flossi became a high ranking female within the Mitumba community; more typically immigrating females remain at a low rank.[55]

Faustino edit

Faustino (b. May 8, 1989) is Fifi's third oldest son.[13] His father was brother Freud's rival Wilkie.[25][42] At the beginning of the documentary Chimps: So Like Us, the infant Faustino is shown with mother Fifi and sister Flossi trying out facial expressions.[73] Fifi became pregnant less than three years after Faustino was born, an unusually short time for a chimpanzee.[44][74] As a result, Faustino had to be weaned at an unusually young age, leading to a particularly difficult weaning conflict.[44][74] This may account for Faustino being more skittish as a juvenile than his younger brother.[44]

Faustino grew into a large but mellow adult who reminded observers of Freud.[74][75] He had worked his way up to become the beta, or second highest-ranking, male in the community until struck by a near fatal disease in 2005.[74] Although Faustino survived, he lost his high rank.[74] After recovering, Faustino slowly climbed his way back up the hierarchy, but it appears he will never become alpha male. However, he did help his younger brother Ferdinand attain alpha status in 2008, and continued to provide important support for Ferdinand in his brother's role as alpha, although as of early 2014 he would sometimes ally himself with Ferdinand's potential rival Titan.[25][76][77] He is the father of Gaia's son Google, born in 2010.[48][49]

Ferdinand edit

Ferdinand (b. August 19, 1992)[78] is Fifi's fourth oldest son.[13] His father was his uncle Figan's rival Evered, who was 41 years old when Ferdinand was conceived.[25][42] He became the first wild great ape to have his birth recorded.[33] By 2007, he had grown into a large enough male to wound the alpha male Kris, and researchers believed he would eventually become the third of Fifi's sons to become alpha male.[79][80] In 2008, he succeeded in supplanting Kris as alpha male, after wounding Kris badly in a fight.[62][81] Ferdinand maintained his alpha male position primarily through intimidation and threats and even surprising attacking males.[62][55] He would bite rivals on the back to leave a scar.[62][55] By the last two years of his reign Ferdinand's aggressiveness had made him so unpopular with other group males that he spent most of his time apart from them, returning to assert his dominance every so often.[62] During that time a coalition of rival males formed that included his nephews Fudge and Fundi, and their father and uncle Sheldon and Sampson, and another nephew Zeus.[62][55] In October 2016 the coalition attacked Ferdinand when he made one of his occasional appearances to try to assert his continuing dominance.[62] Fudge and Fundi led the attack, with some support from their mother (and Ferdinand's sister) Fanni.[62][55] Ferdinand was badly wounded in the attack and had to leave the group to recover.[62] In his absence, Fudge was able to assert his dominance and attain the alpha position.[62]

Ferdinand has been successful in siring offspring. Infants he fathered included Diaz by Dilly fathered at very young age of 12 in 2005, Tabora (2006– ) by Tanga, and Gizmo (2010–2016) by Gremlin.[48][49][59][61][82][83][84][85]

Flirt edit

Flirt (b. July 1998) is Fifi's youngest surviving daughter.[13] Her father was Kris, who would later become alpha male of the troop.[25] She was a large baby and grew rapidly.[86] This probably allowed her to survive when her mother died before she was seven years old.[46][86] Although she spent a few lonely months after Fifi's death, she would begin to travel with her brothers.[86] The documentary Chimpanzee Family Fortunes ends with Flirt finding solace after months of loneliness by being groomed by eldest brother Freud.[46] She followed in the footsteps of her sister Flossi in eventually emigrating to the Mitumba Chimpanzee Community in 2013.[25]

Fudge edit

Fudge (b. 1996) is Fanni's oldest surviving son. His father is Sheldon, a former alpha male. He is the fifth male of the F-family to reach alpha male status, overthrowing his own maternal uncle Ferdinand in October 2016.[62] Fudge had joined a coalition that included his brother Fundi, his father Sheldon, his uncle Sampson and his cousin Zeus in opposition to Ferdinand.[62] In October 2016 Fudge led an attack by the rival males on Ferdinand.[62][55] Fundi was his primary supporter in the attack and they received some support from their mother Fanni, who is Ferdinand's sister.[62][55] After seriously wounding Ferdinand in the attack, Fudge was able to assert his dominance over the remaining group males to attain alpha male status.[62][55]

Fundi edit

Fundi (b. 2000) is Fanni's second oldest surviving son. His father is the former alpha male Sheldon; Fudge, the alpha male as of 2016, is his full brother. In 2016 Fundi helped Fudge attack their uncle Ferdinand, the previous alpha male, which allowed Fudge to attain his alpha status.[62]

G-family edit

Melissa edit

Melissa (c.1950–1986)[31] was a high-ranking female and mother of long-term alpha male Goblin.[87] Researchers also suspected that she was related to alpha male Humphrey and another male, Mr McGregor.[87] She was afflicted by polio in the 1966 epidemic, and for a while was paralyzed in her neck and shoulders, and was forced to walk bipedally. She would regain use of her arms, although she never fully regained movement of her neck.[87] She gave birth to two sets of twins, one set in 1976 of which neither survived the year, and another set in 1977, of which only one, Gimble, survived to adulthood.[31][87] Her other offspring who survived to adulthood was daughter Gremlin.[31] Her daughter Gremlin and her granddaughter Gaia have also given birth to twins.

Goblin edit

Goblin (September 1964 – August 2004) was Melissa's oldest son.[31][65] He was discovered by Goodall when he was just a few hours old, with a twisted face that led her to name him Goblin.[88] Goblin was protected by Figan as an adolescent, but at the young age of 15 in 1979 he challenged and defeated Figan to become the top-ranking male.[18][89] He was defeated and brutally beaten by a coalition of older males later in the year, giving up his top ranking.[18][89] Goblin regained the top-ranked position in 1982, and was the unambiguous alpha male, able to control even coalitions of rival males, by 1984.[18] Goblin maintained his alpha status until 1989, when he was badly beaten and injured by a coalition of males led by Wilkie.[7][65][73] He was sterilized by a bite to his testicles in the process.[43] He spent time away from the rest of the community recovering from his injuries, during which time Goodall administered medication to help him return to health.[73] He later became Freud's key ally in becoming and retaining his position as alpha male, apparently going so far as to encourage Freud to challenge Wilkie for the role.[55] While Freud was alpha Goblin was able to be a wedge between Freud and Frodo, who had previously been close companions, but when Frodo overtook Freud, Goblin switched his allegiance to Frodo.[55] Goblin died of disease in August 2004.[30]

After recovering from his injuries, Goblin maintained a high-ranking status within the community by forming alliances with the alpha males, especially Freud and Frodo.[65][90] While he was establishing himself as a contender for the alpha position, he would occasionally beat up Goodall.[65] In 1983, when his mother Melissa was in estrus, he forced himself upon her despite her efforts to resist.[89] Fanni and Flossi, by Fifi, and Tanga (1989– ), by Patti, are his daughters.[47][63][65][91]

Gremlin edit

Gremlin (b. 1970) is Melissa's only daughter to survive to adulthood.[31] Her father was Evered, who, more than 20 years later, also fathered the alpha male Ferdinand.[25][42] Goodall has described Gremlin as one of her favourite chimpanzes, due to Gremlin’s patience as a mother and expertise in fishing for termites.[56] She has raised twins, Golden and Glitter, who are the oldest known wild twin chimpanzees.[33] She had male offspring in Getty (1982–1986), Galahad (1988–2000), Gimli (2004– ), Gizmo (2010–2016) and a female offspring in Gaia (1993– ).[31][67] In 2015, she gave birth to Grendel and in 2019, at the age of almost 50, she gave birth to a son, Goodali.[92][55]

Although males are perceived as the more aggressive sex in chimpanzees, studies at Gombe have revealed that aggression between females does occur.[34] Such attacks were observed of Fifi and her adult daughter, Fanni, aggressively harassing fellow Kasekela female, Gremlin, on three different occasions; the first being in 1993 following the birth of her daughter Gaia, and then five years later following the birth of her twins. This was followed by another attack on Gremlin six years later following the birth of her next baby.[34] Researchers could only speculate that the attacks on infants of other females had to do with food competition, as each female strives to secure resources for her own group of offspring and thus the expansion of another female's family unit could deplete resources.[29] The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom in the 1970s,[34] which proved that cannibalism, although deemed rare, does occur among chimpanzees, by both males and, in this particular case, jealous females.

Clever females like Gremlin, who bore the brunt of infanticidal females, employed two counter strategies to avoid attacks on them and newborns.[34] One strategy is being in close proximity to adult males for protection, as Gremlin initially did by sticking close to Frodo (the father of her twins) in 1998.[34] The second strategy used by Gremlin was taking a newborn baby from her daughter, Gaia, in 2006, presumably in fear of it being attacked.[34] Although Gremlin's instincts were justified from her own experiences, the latter case ended with the infant dying just five months later.[34]

Gimble edit

Gimble (b. October 1977) is one of two of Melissa's sons to survive to adulthood.[31] He was one of a set of twins; however, his twin brother Gyre died in 1978.[31] As a twin, he grew into a healthy but small adult.[93] He reached a relatively high rank in the late 1990s but fell to a low-ranking male by 2004.[30] He fathered at least two children, Mambo by Malaika, born in 2004, and Zinda (2006 – October 2012) by Trezia.[25][47][48][49] He disappeared in early 2007.[citation needed]

Galahad edit

Galahad (1988-2000) was Gremlin's third son.[94] His father was a low-ranking male, Atlas.[95][96] Galahad was described by researchers at Gombe as mischievous and playful, often observed terrorizing baboons.[95] In the BBC television program Chimpanzee Diary 1998 Galahad was depicted being supportive of his mother Gremlin, staying near by even though he could not intervene during her attack by other community females trying to take her newborn twins. He was developing into a favourite character for the human observers in Gombe, but was unfortunately struck down by the flu epidemic in 2000 and died in February of that year[95]

Gaia edit

Gaia (b. February 1993) is Gremlin's oldest daughter.[31] Her father was Wilkie.[25][42] As a juvenile, she helped her mother raise the twins Glitter and Golden, and this is shown in a number of documentaries about the community.[28][46][56][97] Gaia gave birth to her first baby in April 2006,[34] but the baby's grandmother Gremlin took the infant from her daughter, presumably in fear of it being attacked.[34] Although Gremlin's instincts were justified from her own experiences of other community females attacking her newborns,[34] the infant died just five months later.[22][34] Gaia's second infant was stillborn, but her third pregnancy resulted in twins.[98] However, once again her mother took over the babies, and one of the twins died at a few days old. She finally gave birth to a son named Google in 2010, fathered by Faustino.[48][49][99][100][67] She gave birth to a son named Gabo in 2015.[67]

Glitter and Golden edit

Glitter and Golden (b. July 1998) are the twin daughters of Gremlin, fathered by Frodo.[31][46][44][56][61] Gaia contributed to the raising of Gremlin’s twins, helping them to become the only known pair of wild chimpanzee twins known that have survived to the age of sexual maturity.[46][56] Each of the pair exhibits a different personality. Glitter is shy and cautious, while Golden is more adventurous and willing to engage in rough and tumble activities.[28][101] In 2011, Glitter gave birth to daughter named Gossamer, who was fathered by Sheldon.[102][67] She gave birth to a son named Ghurubu in 2016.[67] In 2012, Golden gave birth to a daughter named Glamour.[48][49][102] Golden also gave birth to a son named Gombe in 2017. Golden and Glitter have remained extremely close through adulthood.[67]

P-family edit

Passion edit

Passion (c.1951–1982) was a central female of the Kasekela community. She, along with her daughter Pom, captured, killed, and ate about ten newborns at Gombe.[103] Passion possessed a rather domineering and aggressive personality; according to Goodall, Passion was an unnatural and indifferent mother to her three surviving offspring, but she became a very high-ranking female. She became more solitary later in life and died of some type of wasting disease in 1982. In addition to Pom, Prof and Pax, she gave birth to a baby in 1962 which died in infancy. She also had a stillborn baby, or possibly a miscarriage in 1970.

Pom edit

Pom (1965–1983) was Passion's oldest known offspring to survive to adulthood. Pom was instrumental in helping her mother capture and kill community infants.[103] Pom gave birth herself to Pan in 1978, and was very cautious with the infant around her mother. Pan died of injuries from an accident in 1981. Pom eventually disappeared about a year after her mother's death. She may have transferred to another community, but she was never seen again.

Prof edit

Prof (October 27, 1971 – 198?)[26] was a male that survived to adulthood. He and Pom offered their little brother comfort following Passion's death. Prof and Pax associated closely for a number of years. Prof is deceased.

Pax edit

Pax (b. December 16, 1977)[26] was Passion's last offspring. He was only four years old when he was orphaned. He had the support of an elder brother and sister, and had very few behavioural issues following his mother's death. When Pax was very young, his mother was wounded in some type of conflict. Pax lost his testicles as a result of the conflict, which has affected his personality greatly.[104] He is small for a male, doesn't compete in the male hierarchy, and is unable to reproduce.[30][104][85] He also has been chosen by a couple of orphans as a companion.

Pan edit

Pan (October 17, 1978 – 1981)[26] was the only known offspring of Pom. He was injured after being blown from a palm tree by a strong wind and died four days later.

T-family edit

Patti edit

Patti (c.1961–2005) was not born in the Kasekela community but joined the community in 1973.[28] She was an assertive female who spent much time on her own.[28] Her first two infants died before reaching adulthood, but she has successfully raised several chimps since.[28] Besides Tapit, Tita, Tanga, Titan and Tarzan, she had two offspring who died in infancy, including her first born Pibi (1978)[28] and a baby of unknown sex in 1988. She was killed by males from the rival Mitumba Chimpanzee Community in 2005.[1] A female chimpanzee leaving the safety of her community territory poses risks, and studies on chimpanzee intercommunity relations revealed that chimpanzees are likely to be attacked by the territorial males of a rival community,[36] and the intensity of the attacks depends on the gender of the intruder, and if the stranger is female, it depends on whether or not they are of any reproductive benefit to the males. Young, oestrus females are often spared an attack because they are deemed to be potential new mates, but an anestrus female (non-cycling or lactating) is considered of very little use for the males, which results in lethal aggression and fatality.[36]

Tapit edit

Tapit (1979–1983)[26] was Patti's second-oldest son. He had a rough life and was sickly as an infant and died as a juvenile.[28]

Tita edit

Tita (b. January 1984) is Patti's oldest daughter. She gave birth to her first child, Tibi, in 1995; he died in infancy. Another son, Tofiki (fathered by Sheldon), was born in 2000 and was killed in 2004 when Freud attacked him while grooming Tita.[30][85]

Tanga edit

Tanga (b. April 1989) is Patti's second oldest daughter. Her father was Goblin.[91][85] She still lives in the community and gave birth to her first infant, Tom, in 2001.[25] In 2006, she gave birth to a daughter named Tabora.[25] In October 2012, she gave birth to a daughter named Tarime who died in 2013.[25] In 2014, she gave birth to Tukuyu, whose father is mostly likely Fudge.[citation needed]

Titan edit

Titan (b. August 1994) is Patti's third oldest son, fathered by Frodo.[42] Titan displays aggressive behaviour resembling his father Frodo's violent tendencies. Like Frodo, as a youngster he would throw rocks at researchers.[76] Also like his father, Titan is large, and as of 2013 was the largest male in the community.[76] As of 2012, researchers believed that he may have been ready to challenge Ferdinand for alpha status, as he spent a lot of time near Ferdinand and sometimes helped Ferdinand maintain control.[76] However, over the following year, Titan slipped in rank.[76] Researchers suspect that he may be afraid to challenge for the alpha spot, and also that his reluctance to groom other chimps may inhibit him from forming the alliances needed to attain alpha status.[76] As of early 2014, Titan had occasionally been able to ally himself with Ferdinand's older brother Faustino in attempts to challenge Ferdinand, but he has not been able to maintain that alliance reliably, and at times Faustino would also join with Ferdinand to attack Titan.[77] Titan fathered a daughter, Baseke, by Bahati in 2010.[48][49]

Tarzan edit

Tarzan (b. October 1999) is Patti's fourth oldest son. His father was Frodo, making him Titan's full brother.[25]

Tom edit

Tom (b. March 2001) is Tanga's first son, fathered by Kris.[25][85]

Tabora edit

Tabora (b. 2007) is Tanga's first daughter. Her father is Ferdinand.[25]

S-family edit

Sparrow edit

Sparrow (b. 1958) is the oldest female and matriarch of the family.[105] She has had seven offspring, including four sons (Sheldon, Steve, Spud and Sinbad) and three daughters (Sandi, Barbet and Schweini).[22] Four of the seven are alive: Sandi, Sheldon, Sinbad and Schweini. Her son Sheldon became the alpha male of the Kasekela community in 2004. Steve (born 1989) and Spud (born 1996) both died in infancy.

Sandi edit

Sandi (b. October 1973) is Sparrow's oldest daughter and first offspring. Sandi is one of the shyer females of the Gombe chimpanzees. She grew up within the community but her mother was not social and as a result, she still has a timid nature. Sandi is still close to her mother, Sparrow, and the two prefer each other's company rather than anyone else's. Sandi was very nervous of large groups of chimpanzees within the community and often prevented her offspring from playing with the others. However, she has become much more confident and her offspring have thrived.

Sandi has five offspring which include two daughters, Sherehe and Samwise and two sons, Sampson and Siri who are both sired by Apollo.

Barbet edit

Barbet (ca. 1978 – 1992) was Sparrow's second-oldest daughter. She died in 1992.

Sheldon edit

Sheldon (b. May 1983) is Sparrow's oldest son. Sheldon grew up within the community as a subordinate male but within 2004 following the vacancy of the alpha position after former alpha Frodo's departure, when he became the alpha male. Sheldon held on to dominance for only a year and lost the alpha position to another male, Kris, in 2005.[30] He was part of a coalition that helped his son Fudge attain the alpha male position in 2016.[62] He fathered Fanni's first three offspring, Fudge, Fundi and Familia.[25][85] He was a low ranking male when he fathered Fudge and Fundi, but had become the alpha male by the time he fathered Familia.[85] He also fathered Tita's son Tofiki in 2000 while a mid-ranking male.[85]

Schweini edit

Schweini (b. April 1991) is Sparrow's third oldest daughter. Her father is Wilkie.[42][85] She gave birth to her first infant in September 2007 but she died in infancy.[106][25] She gave birth to the female Safi on October 16, 2008, but she died in August 2011.[25] She also gave birth to a son Schall in November 2012.[25] She was brutally attacked by females of the Mitumba chimpanzee community while associating with males of that community in both 2004 and 2005.[106]

Sinbad edit

Sinbad (b. June 2001) is Sparrow's youngest son. His father was Frodo.[25]

Sherehe edit

Sherehe (January 1991 – May 11, 2006) was Sandi's oldest daughter and first offspring.[25] Her father was Beethoven.[85] She gave birth to her first infant, Shangaa in 2004.

Sampson edit

Sampson (b. April 1996) is Sandi's oldest son. His father is Apollo. He is a low-ranking male. Sampson was part of a coalition that included his uncle Sheldon and his cousin Fundi that helped Sampson's cousin Fudge supplant Ferdinand as alpha male in 2016.[62]

Samwise edit

Samwise (b. June 2001) is Sandi's second-oldest daughter. Her father was Frodo.[25]

Male dominance edit

Chimpanzees live in large mixed-sexed communities, in a society that is male dominant[107] and the male chimpanzee who rules above all others (males and females) is referred to as the alpha male. The alpha male is the highest-ranking, most dominant chimpanzee in the entire community, and can control most situations, including situations in which he is opposed by a coalition of other males.[18] Nonetheless, an alpha male typically cannot monopolize breeding opportunities, and in some cases lower-ranking males have been more successful at mating than alpha males, since the alpha male has to spend much energy maintaining his rank.[108] Faben (Figan's older brother with a paralyzed arm), Evered (defeated by Figan in his bid for alpha status who became Ferdinand's father) and Sherry (lower-ranking male who became Frodo's father) are examples of males who successfully devoted their energy to breeding success despite not attaining alpha status.[108] However, a very secure alpha male, such as Frodo, can use his status to increase his access to females in estrus, especially during the period when she is at peak fertility.[108]

Goliath was the first chimpanzee Goodall recognized as the alpha male.[7][109] In 1964, Mike, who was previously a low-ranking male, deposed him.[110][111] Goliath remained a high-ranking male for several more years, but fell in rank as he aged.[109] Eventually, Goliath was one of the chimpanzees who left the Kasekela community to form the Kahama community.[109] However, in 1975 he was brutally attacked by a group of Kasekela males led by Faben, and died as a result of his injuries.[20][109]

Mike figured out that incorporating cans and other objects left by the human researchers into his displays would make them more effective. These displays indeed allowed him to take over the alpha male rank in 1964.[110][111] By 1969, a group of younger males including Humphrey, Faben, Figan and Evered challenged and defeated him, making Humphrey the alpha male and returning Mike to his low-ranking position.[18]

From 1972 through 1989, two males, Figan and Goblin, held the alpha position. Figan was able to defeat Humphrey when Faben became his ally. For a period after Faben's death in 1975, Figan remained the top-ranking male, but was unable to control coalitions of other males when they joined together, making his status as alpha male ambiguous. Within a few months, Figan reestablished himself as the unambiguous alpha by forging alliances with other high-ranking males, such as Humphrey. In 1979, 15-year-old Goblin, who had previously been protected by Figan challenged him for the alpha position. Goblin temporarily held the top-ranking position in 1979, although could not control coalitions of other males, before the more senior males brutally beat him and restored Figan to the alpha position. By 1982, Goblin had reestablished himself as the top-ranking male; by 1984, he was unambiguously the alpha male, able to control situations even against coalitions of other males.[18]

In 1989, Wilkie defeated Goblin and attained the alpha position.[73] Wilkie attained this position despite being one of the smallest males in the community, at 37 kilograms (82 lb).[112] According to researchers at the University of Minnesota's Jane Goodall Institute Center for Primate Studies, Wilkie attained his position primarily by becoming popular by obsessively grooming other males.[104][112] Unlike most males, Wilkie also groomed females.[112] Wilkie also made effective use of charging displays.[104] Wilkie was overthrown in 1992 by his childhood playmate Freud, who was in turn deposed by his brother Frodo in 1997 while Freud was ill.[41][52] Freud, at 44.8 kg (99 lb), was larger than Wilkie and relied on a mixture of moderate force and grooming to become alpha and maintain his status.[104][112] Frodo, at 51.2 kg (113 lb), was one of the largest males ever seen in the group, relied primarily on brute force to attain and maintain alpha status.[104][112] After Frodo became ill in 2002, he was brutally beaten by a coalition of males led by Sheldon and spent two years travelling away from the rest of the community. No chimpanzee was able to establish himself as an alpha male until 2004. Sheldon briefly held the top-ranking male position in 2004; however, by 2005, Kris had defeated all competitors, including Frodo, and established himself as the unambiguous alpha male.[46][56] In 2008, Kris was supplanted as alpha male by Ferdinand, the youngest surviving brother of Freud and Frodo.[82] Unlike most deposed alpha males, who tend to stay away from the group for a while in order to avoid further injury while the new alpha is consolidating his position, Kris stayed with the group after being deposed and eventually died of his injuries.[25][43][59][61][83][113]

In October 2016, Ferdinand was attacked by a coalition of males led by his nephews Fudge and Fundi and also included the former alpha male Sheldon.[62] Ferdinand was badly injured in the attack and Fudge asserted himself as the new alpha male.[62]

Name Reign Lifespan Notes Reference
Goliath 1961–1964 ?–1975 [7][18]
Mike 1964–1969 ?–1975 [7][18][110][111]
Humphrey 1969–1971 ca. 1947–1980 [7][18][110][111]
vacant 1971–1972 After Humphrey was deposed as alpha in 1971, no chimpanzee clearly held the highest rank until Figan established himself as alpha male over Humphrey and Evered in 1972. [18][25]
Figan 1972–1979 1953–1982 For a nine-month period in 1975 and 1976, while Figan was clearly the highest-ranking male, he was not an unambiguous alpha male. [16][18]
Goblin 1979 1964–2004 In 1979, Goblin was able to oust Figan and become the top-ranked male, although he was not able to establish himself as an unambiguous alpha male before Figan regained his status. [18][90]
Figan 1979–1982 1953–1982 [16][18]
Goblin 1982–1989 1964–2004 Although Goblin established himself as the highest-ranking male during 1982, he did not become the unambiguous alpha male until 1984. [7][18][73][89]
Wilkie 1989–1993 1972–2012 [7][73][112][114][115]
Freud 1993–1997 1971–2014 Nephew of Figan [7][41][114]
Frodo 1997–2002 1976–2013 Replaced his brother Freud; nephew of Figan [7][52][114][116]
vacant 2002–2004 After Frodo was ousted as alpha male during his illness in 2002, no male established himself as unambiguously the highest-ranking male until 2004. [46]
Sheldon 2004–2005 1982– Was part of a coalition that helped his son Fudge become alpha male in 2016 [46][52][115][116][117][118][62]
Kris 2005–2008 1982–2010 [7][43][46][52][115][118]
Ferdinand 2008–2016 1992– Youngest brother of Freud and Frodo; nephew of Figan; displaced by his sister's son Fudge. [25][82][62]
Fudge 2016– 1996– Nephew of Freud, Frodo and Ferdinand, grandnephew of Figan, son of Sheldon, and grandson of Goblin. [62]

Female dominance edit

Even though chimpanzees live in a male-dominant society,[107] the female chimpanzees in the Kasekela community have evolved their own pecking order, in which the top-ranking female has priority access to food, mates and sleeping sites. Studies in Gombe have discovered that the rank of a mother directly correlates to infant survival, fast maturing of daughters and faster production of offspring.[119] Jane Goodall considered elderly female Flo, the original matriarch of what is now the famous 'F' family of Gombe, to be the most dominant of the females in the Kasekela community at the time that the study began (in the 1960s)[120] and this was evident in her interaction with other females, where she was never hesitant about asserting herself towards new females.[121] She was a successful mother and fierce defender of her offspring, one of whom would succeed her.

The dominance of a female to other community females depends largely on family ties.[29][122] Fifi was a popular female in Gombe, not only from her high-ranking family but also from her close ties with the males. She inherited her mother's superior status[120] and because her older brothers Faben and Figan were among the top big males, she grew up with enormous confidence.[29] As a mother of nine, Fifi was fiercely defensive of her offspring[69] and her authority was rarely questioned. This was also possible because of her ability to fearlessly navigate a male-dominant society,[69] helped in part from the fact that her adult sons Freud and Frodo grew to become the alpha males in the community. Having produced the most offspring out of any female during her time, Fifi was regarded as Gombe's most fertile female[123] and her ability to have reproduced as frequently as she did, and have enough resources and social standing to ensure the survival of seven of her nine children validates the studies of how the social rank of a chimpanzee mother directly correlates to infant survival, fast maturing of daughters and faster production of offspring.[119]

Just as her mother's high status rubbed off on her,[29] Fifi's high standing in the community benefited her daughters. Although second daughter Flossi emigrated to dominate another chimp community[34] her eldest daughter Fanni remained with her mother, and together they became a formidable force in Gombe. Observers in Gombe discovered that the linear hierarchy among females becomes apparent in aggressive interactions involving infanticide.[34] One of two contexts for female-female aggressive encounters in Gombe was the presence of newborn infants.[34] Matriarch Fifi was known to be a feisty female who, although a very successful mother herself, was observed to have carried out ferocious and infanticidal attacks on other, lower-ranking females in her community. Such attacks were observed of Fifi and her adult daughter, Fanni, aggressively harassing lower-ranking Kasekela female, Gremlin, on three different occasions, with the goal of killing her newborn infants[34] Researchers could only speculate that the attacks on infants of other females had to do with food competition, as each female strives to secure resources for her own group of offspring and thus the expansion of another female's family unit could deplete resources.[29] The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom, a dominant pair who only attacked subordinate females, in the 1970s,[34] which proved that cannibalism, although deemed rare, does occur among chimpanzees, by both males and, in this particular case, jealous females.

The loss of her mother Fifi in 2004 greatly reduced the dominance of Fanni, now the oldest surviving female of the once mighty 'F' group, towards the other females, and this is despite having high-ranking male relatives in the community, including her son Fudge who has been the top male since 2016[124] and her high-ranking brothers Faustino and Ferdidand (Fudge's predecessor). Fanni shifted her range to the southern part of the Kasekela territory, further away from the other community females,[69] particularly Gremlin's growing 'G' family, which now makes up a large portion of the Kasekela community and who occupy the top of the hierarchy.[69]

Other communities of chimpanzees at Gombe Stream National Park edit

Within the Gombe Stream National Park live other chimpanzee communities other than the Kasekela, which also inhabit certain home ranges. The two other communities currently bordering the Kasekela community include:

One extinct community of Gombe was:

The Kahama community came about from a split within the Kasekela community.[11][20] In the early 1970s several of the Kasekela chimpanzees, including 8 adult males, kept to the northern portion of the Kasekela range, while others, including 7 adult males, kept to the southern portion of the range.[11][20] The two groups became hostile towards each other, and had less and less to do with each other, and eventually the southern community became totally separate, and was referred to by researchers as the Kahama community.[11][20] By 1974, the Kasekela males began to attack the Kahama chimpanzees in what was referred to as the "Four Year War."[11][20] Figan, Faben, Humphrey, Frodo's father Sherry, and Sherry's brother Jomeo were often among the ringleaders of the attacks.[20] Researchers described these attacks as being of the nature of raids, as opposed to simply defending their territory.[11] The Kasekela chimpanzees did not collect food during these "raids," although at least three Kahama females were brought back to the Kasekela community.[11][12][20] Kahama chimpanzees were brutally attacked during these raids and often were killed.[11][20] Former Kasekela alpha male Goliath, who had joined Kahama and was then very old, was killed in one such attack in 1975.[20] Kahama's alpha male Charley was killed in 1977.[20] By 1977 the Kahama community had been completely wiped out, and the Kasekela community absorbed its territory.[20]

Annexing the Kahama territory brought the Kasekela community in contact with the Kalande Chimpanzee Community.[11][20] The Kalande community may have been attacking Kahama chimpanzees during the Four Year War, and after Kahama was wiped out they began to attack Kasekela chimpanzees.[11][20] At least one Kasekela female was badly injured in an attack by Kalande chimpanzees, and at least two infants were killed in such attacks.[11][20] Former Kasekela alpha male Humphrey died in 1981, possibly from injuries from an attack by Kalande chimpanzees.[11]

Kasekela has had interactions with the Mitumba Chimpanzee Community as well. Some of these interactions have been hostile, and some Mitumba chimpanzees are known to have been killed by Kasekela chimpanzees.[30][125] At least two Kasekela females, Fifi's daughters Flossi and Flirt, have emigrated from Kasekela to join the Mitumba community.[25][125] Based on circumstantial evidence researchers believed that it is possible that Fifi herself may have died as a result of an attack by Mitumba chimpanzees.[30]

Books about the community edit

  • Goodall, J. (1971). In the Shadow of Man. Houghton Mifflin. ISBN 978-0-395-33145-3.
  • Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. ISBN 978-0-674-11649-8.
  • Goodall, J. (1990). Through a Window. Houghton-Mifflin. ISBN 978-0-395-59925-9.
  • Goodall, J. (1997). The Chimpanzee Family Book. North-South Books. ISBN 978-1-55858-803-5.
  • Shah, A.; Rogers, F. (2014). Tales from Gombe. Firefly. ISBN 978-1-77085-468-0.

Films about the community edit

Further reading edit

  • Morris, Ian (2014). War! What Is It Good For?: The Role of Conflict and the Progress of Civilisation from Primates to Robots. MacMillan. ISBN 978-1-84765-454-0.

See also edit

References edit

  1. ^ a b Wilson, M.L. (2012). "Long-term Studies of the Chimpanzees of Gombe National Park, Tanzania". In Kappeler, Peter M.; Watts, David P. (eds.). Long-term Field Studies of Primates. Springer. pp. 357–384. ISBN 9783642225130.
  2. ^ a b Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. p. 84. ISBN 978-0-674-11649-8.
  3. ^ a b Morgen, Brett (2018). Jane (film). National Geographic.
  4. ^ Wilson, Michael L.; et al. (2020). "Research and conservation in the greater Gombe ecosystem: challenges and opportunities". Biological Conservation. 252: 108853. Bibcode:2020BCons.25208853W. doi:10.1016/j.biocon.2020.108853. PMC 7743041. PMID 33343005.
  5. ^ Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. pp. 535–539. ISBN 978-0-674-11649-8.
  6. ^ Goodall, J. (1971). In the Shadow of Man. Houghton Mifflin. pp. 35–37. ISBN 978-0-395-33145-3.
  7. ^ a b c d e f g h i j k l . The Jane Goodall Institute. Archived from the original on December 7, 2008. Retrieved 2009-02-28.
  8. ^ "Scientist Makes Unique Record of Chimpanzees". Greeley Daily Tribune. March 19, 1964. p. 34. Retrieved 2019-01-26 – via newspapers.com.
  9. ^ Harriman, Jane (September 2, 1963). "Wild Chimp is a Friendly Chap". Boston Globe. p. 44. Retrieved 2019-01-26 – via newspapers.com.
  10. ^ a b Eppinga, J. (2009). They Made Their Mark: An Illustrated History of the Society of Woman Geographers. Globe Pequot. pp. 218–220. ISBN 978-0-7627-4597-5.
  11. ^ a b c d e f g h i j k l Wrangham, R.; Peterson, D. (1996). "Demonic Males". The Washington Post. Retrieved 2015-01-14.
  12. ^ a b Morris, I. (2014). "Red in Tooth and Claw: Why the Chimps of Gombe Went to War". War! What Is It Good For?: Conflict and the Progress of Civilization from Primates to Robots. Farrar, Straus and Giroux. ISBN 978-0-374-28600-2.
  13. ^ a b c d e f g h i j k l . University of Minnesota JGI Center for Primate Studies. Archived from the original on 2009-02-25. Retrieved 2009-03-03.
  14. ^ a b Lickley, D., Low, S., Goodall, J. (2002). Jane Goodall's Wild Chimpanzees (IMAX).
  15. ^ Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. pp. 65–66. ISBN 978-0-674-11649-8.
  16. ^ a b c d . University of Minnesota JGI Center for Primate Studies. Archived from the original on 2009-02-25. Retrieved 2009-02-28.
  17. ^ a b c d e Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. p. 64. ISBN 978-0-674-11649-8.
  18. ^ a b c d e f g h i j k l m n o p Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. pp. 424–429. ISBN 978-0-674-11649-8.
  19. ^ Goodall, J. (1990). Through a Window. Houghton-Mifflin. pp. 44–50. ISBN 978-0-395-59925-9.
  20. ^ a b c d e f g h i j k l m n o Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. pp. 503–514. ISBN 978-0-674-11649-8.
  21. ^ a b c d e f g Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. pp. 64–65. ISBN 978-0-674-11649-8.
  22. ^ a b c Quammen, David (August 2014). . National Geographic. Archived from the original on December 5, 2017. Retrieved 2017-12-04.
  23. ^ Goodall, J. (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. pp. 178–184. ISBN 978-0-674-11649-8.
  24. ^ Jane Goodall (1986). The Chimpanzees of Gombe: Patterns of Behavior. The Belknap Press of Harvard University Press. p. 625. ISBN 978-0-674-11649-8.
  25. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al Shah, A.; Rogers, F. (2014). Tales from Gombe. Firefly. pp. 318–320. ISBN 978-1-77085-468-0.
  26. ^ a b c d e Brent, L.; et al. (December 1997). "The Influence of Siblings on Wild Infant Chimpanzee Social Interaction". Behaviour. 134 (15/16): 1189–1210. doi:10.1163/156853997x00124. JSTOR 4535501. – via JSTOR (subscription required)
  27. ^ a b c d . University of Minnesota JGI Center for Primate Studies. Archived from the original on 2009-03-28. Retrieved 2009-03-03.
  28. ^ a b c d e f g h i j k l m n o p q r s t "Gombe Chimpanzee Biographies" (PDF). wildchimpanzees.org. Retrieved 2014-09-18.
  29. ^ a b c d e f g Subramanian, S (4 May 2022). "The Queens of Kasekela". Medium. Retrieved 19 July 2023.
  30. ^ a b c d e f g h i Wilson, M.L. "Gombe Stream Research Centre 2004 Annual Report" (PDF). Retrieved 2015-01-11.
  31. ^ a b c d e f g h i j k l m . University of Minnesota JGI Center for Primate Studies. Archived from the original on 2009-02-26. Retrieved 2009-03-04.
  32. ^ van Lawick, H., Sutherland, D. (1991). People of the Forest: The Chimps of Gombe. Discovery Channel.
  33. ^ a b c d e . National Geographic Wild. June 13, 2014. Archived from the original on December 14, 2017. Retrieved 2017-12-13.
  34. ^ a b c d e f g h i j k l m n o p q r s t u v w x Pusey, A; Murray, C; Wallauer, W; Wilson, M; Wroblewski, E; Goodall, J. Severe Aggression Among Female Pan troglodytes schweinfurthii at Gombe National Park, Tanzania (PDF). VIVAMUS.
  35. ^ Wilson, M. L; Kamenya, S; Collins, D. A; Wallauer, W. R (2004). Gombe Stream Research Centre2004 Annual Report. Kigoma, Tanzania: Jane Goodall Institute. p. 16. Retrieved 17 July 2023.
  36. ^ a b c d Smuts, B (1992). "Male aggression against women - An evolutionary perspective" (PDF). Human Nature. 3 (1): 7, 8. doi:10.1007/BF02692265. PMID 24222394. Retrieved 20 July 2023.
  37. ^ Wilson, M. L; Collins, D. A; Wallauer, W. R; Kamenya, S (2005). Gombe Stream Research Centre 2005 Annual Report (PDF). Kigoma, Tanzania: Jane Goodall Institute. p. 20. Retrieved 17 July 2023.
  38. ^ a b Goodall, J. (1971). In the Shadow of Man. Houghton Mifflin. pp. 232–236. ISBN 978-0-395-33145-3.
  39. ^ Goodall, J. (1990). Through a Window. Houghton-Mifflin. pp. 112–113. ISBN 978-0-395-59925-9.
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kasakela, chimpanzee, community, kasekela, chimpanzee, community, formerly, spelled, kasakela, habituated, community, wild, eastern, chimpanzees, that, lives, gombe, national, park, near, lake, tanganyika, tanzania, community, subject, jane, goodall, pioneerin. The Kasekela chimpanzee community formerly spelled Kasakela 1 is a habituated community of wild eastern chimpanzees that lives in Gombe National Park near Lake Tanganyika in Tanzania 2 The community was the subject of Jane Goodall s pioneering study that began in 1960 and studies have continued ever since becoming the longest continuous study of any animals in their natural habitat 2 3 4 As a result the community has been instrumental in the study of chimpanzees and has been popularized in several books and documentaries The community s popularity was enhanced by Goodall s practice of giving names to the chimpanzees she was observing in contrast to the typical scientific practice of identifying the subjects by number Goodall generally used a naming convention in which infants were given names starting with the same letter as their mother allowing the recognition of matrilineal lines One of the most important discoveries that was learned by observing the Kasekela chimpanzee community was the use of tools On November 4 1960 Goodall observed a chimpanzee that she had named David Greybeard using a grass stalk as a tool to extract termites from a termite hill Later she observed David Greybeard and another chimpanzee named Goliath stripping leaves off twigs to create termite fishing tools Previously tool use in chimpanzees was only rarely observed and tool creation by non human animals had never been observed Until then tool making was considered one of the defining characteristics of being human 5 6 7 Another important observation occurred a few days earlier on October 30 1960 On that day Goodall observed the community s chimpanzees eating meat dispelling the notion that chimpanzees are vegetarians 7 A third observation by Goodall in the early 1960s was that male chimpanzees perform a rain dance charging calling slapping the ground and trees and dragging branches in the rain 8 9 In the early 1970s the chimpanzees of the community were observed to engage in ongoing coordinated attacks against the chimpanzees of the neighbouring Kahama Chimpanzee Community ultimately wiping it out 10 11 According to historian Ian Morris this Four Year War represented the first time scientists had observed chimpanzees deliberately seek out attack and leave for dead chimps from another community and it has been described as the first record of lasting warfare among non human primates 10 12 Several families within the Kasekela chimpanzee community have been particularly prominent in books and documentaries The F family has produced five alpha males for the community and the matriarch Flo played a particularly important role in acknowledging Goodall s acceptance as a human observer by the community The G family has produced at least one alpha male and also the birth of several twins which are rare among chimpanzees There are other families as well which include the T family and S family which has produced one alpha male Contents 1 F family 1 1 Flo 1 2 Faben 1 3 Figan 1 4 Fifi 1 5 Flint 1 6 Flame 1 7 Freud 1 8 Frodo 1 9 Fanni 1 10 Flossi 1 11 Faustino 1 12 Ferdinand 1 13 Flirt 1 14 Fudge 1 15 Fundi 2 G family 2 1 Melissa 2 2 Goblin 2 3 Gremlin 2 4 Gimble 2 5 Galahad 2 6 Gaia 2 7 Glitter and Golden 3 P family 3 1 Passion 3 2 Pom 3 3 Prof 3 4 Pax 3 5 Pan 4 T family 4 1 Patti 4 2 Tapit 4 3 Tita 4 4 Tanga 4 5 Titan 4 6 Tarzan 4 7 Tom 4 8 Tabora 5 S family 5 1 Sparrow 5 2 Sandi 5 3 Barbet 5 4 Sheldon 5 5 Schweini 5 6 Sinbad 5 7 Sherehe 5 8 Sampson 5 9 Samwise 6 Male dominance 7 Female dominance 8 Other communities of chimpanzees at Gombe Stream National Park 9 Books about the community 10 Films about the community 11 Further reading 12 See also 13 ReferencesF family editFlo edit Flo c 1919 1972 13 was the matriarch of the F family so named because she and all her matrilineal descendants were given names beginning with the letter F In 1962 Flo was one of the first chimpanzees to approach Goodall s camp along with her infant daughter Fifi Video of Flo approaching Goodall and allowing Fifi to reach out to touch Goodall s forehead letting Goodall know she had been accepted is shown in the IMAX film Jane Goodall s Wild Chimpanzees 14 Later she brought her sons Figan and Faben who would later become prominent members of the community to the camp and when she came into estrus in 1963 she attracted the community s males to Goodall s camp 14 15 Flo was a high ranking female who had at least five known offspring three sons Faben Figan and Flint and two daughters Fifi and Flame Flo was 28 years old when she gave birth to Faben Most female chimpanzees have their first infant when 12 14 years of age so it is possible that Flo had at least one infant prior to Faben that either died or if it was a female would have transferred to one of the other communities prior to Goodall s arrival to Gombe in 1960 When Flo died in 1972 she was given an obituary in Britain s Sunday Times 16 Faben edit Faben c 1947 1975 13 was Flo s oldest known offspring 17 He was a powerful male but lost the use of one arm in a polio epidemic that ravaged the community in 1966 17 Despite his disability he learned to create spectacular bipedal charging displays which allowed him to regain a high rank within the community 17 His support was critical to Humphrey s ascension to alpha male rank in 1969 and to his brother Figan s ascension to alpha male in 1972 18 19 Faben also participated in the conflicts with the rival Kahama community which was formed by former Kasekela members that occurred between 1973 and 1975 and was a leader in the killing of several Kahama males including former Kasekela alpha male Goliath 17 20 Faben himself died in 1975 possibly as a result of further conflicts with the Kahama community 17 Following Faben s death Figan had difficulty maintaining his alpha male position for several months and eventually lost his alpha status to Goblin 18 Figan edit Figan c 1953 1982 13 was Flo s second son 21 Goodall considered him to be Gombe s most intelligent chimpanzee 22 Although younger than Faben he was able to dominate his older brother after Faben s arm was paralyzed from polio in 1966 21 With Faben s support Figan was able to become alpha male in 1972 by defeating the prior alpha Humphrey and an older competitor Evered At this time Goodall described him as the most powerful alpha male in Gombe s recorded history 21 After Faben s death in 1975 Figan lost his unquestioned alpha status in that he could be dominated by a coalition of males but remained the top ranked male 21 By 1977 he had regained his alpha status by forging alliances with other males including his predecessor Humphrey 21 In 1979 he was toppled from his alpha status by 15 year old Goblin whom he had previously supported but regained his alpha status once again by forging alliances with other males 21 Although a powerful male Figan s ability to make alliances with other males was instrumental to allow him to hold on to his alpha status 23 In 1982 Goblin unseated him again and Figan disappeared and presumably died a few months later although when last seen he appeared to be in good health 21 Although Figan was the alpha male for several years he is not known for certain to have sired any infants although he is likely the father of a few 24 Fifi edit Fifi c 1958 2004 13 25 26 was Flo s oldest daughter 27 28 Fifi benefited from her high ranking family and ascended to the top of the female hierarchy in her community 29 27 28 Fifi had nine infants seven of whom survived to independence 27 28 Her two oldest sons Freud and Frodo both have become the community s alpha male as did another son Ferdinand One of her other sons Faustino also attained high rank Her three surviving daughters were Fanni Flossi and Flirt 27 28 She also had one son Fred 1996 1997 and one daughter Furaha 2002 2004 who died in infancy 13 30 Fred was fathered by one of Fifi s other sons Frodo 31 Five year old Fifi was the focal point of the documentary People of the Forest The Chimps of Gombe in which her mother and siblings also featured attempting to gain access to her infant brother Flint 32 All four of Fifi s surviving sons had all held the alpha and or beta positions in the community Grandson Fudge by Fanni also attained alpha male status Late in Frodo s reign as alpha male Fifi helped defend him against attacks and she suffered some injuries in the process 33 Although males are perceived as the more aggressive sex in chimpanzees studies at Gombe have revealed that aggression among females with each other does indeed occur 34 One of two contexts for female female aggressive encounters in Gombe was the presence of newborn infants 34 Fifi grew to be a formidable female who although a very successful mother herself was observed to have carried out ferocious and infanticidal attacks on other lower ranking females in her community Such attacks were observed of Fifi and her adult daughter Fanni aggressively harassing fellow Kasekela female Gremlin on three different occasions the first being in 1993 following the birth of her daughter Gaia and then five years later following the birth of her twins This was followed by another attack on Gremlin six years later following the birth of her next baby 34 Researchers could only speculate that the attacks on infants of other females had to do with food competition as each female strives to secure resources for her own group of offspring and thus the expansion of another female s family unit could deplete resources 29 The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom in the 1970s 34 which proved that cannibalism although deemed rare does occur among chimpanzees by both males and in this particular case jealous females In 2004 Fifi who was by then 46 years old which made her the oldest living chimpanzee in the Kasekela community at that point in time disappeared in September with her youngest and ninth offspring two year old daughter Furaha 35 During that year she shifted her range to the far north of the Kasekela community territory and the simultaneous advancement of the neighbouring Mitumba community further south nearing the Kasekela territory border suggests that Fifi and her youngster may have been fatally attacked by the Mitumba territorial males A female chimpanzee leaving the safety of her community territory poses risks particularly if she has a vulnerable youngster Studies on chimpanzee intercommunity relations revealed that chimpanzees are likely to be attacked by the territorial males of a rival community 36 and the intensity of the attacks depends on the gender of the intruder and if the stranger is female it depends on whether or not they are of any reproductive benefit to the males Young cycling females are often spared an attack because they are deemed to be potential new mates but a lactating mother with a young infant is considered of very little use for the males which results in lethal aggression and fatality 36 The bodies of Fifi and Furaha were never found but the confirmed death of fellow Kasekela female Patti in 2005 by neighbouring Mitumba community males 37 provides a near definitive conclusion of the fate that befell Fifi and her youngster Flint edit Flint 1964 1972 was Flo s third son and her first infant born after Goodall began observing the community 13 Goodall had intended that Flint become the first wild chimpanzee to have his life documented from birth to death 3 Flint had difficulty weaning and became distraught and angry when the birth of his sister Flame displaced him as Flo s baby 38 After Flame disappeared possibly by the hand of Flint he regained his enthusiasm but resumed his needy infantlike behaviour 38 After his nephew Freud was born Flint became fascinated with the new infant and became an important influence of Freud s first year of life 39 Following the death of his mother Flo in 1972 Flint became depressed and ill he died about three weeks after her 16 Flame edit Flame 1968 1969 was Flo s daughter 13 Flo s son Flint at age four threw violent fits in order to suckle his mother s breast while newborn Flame was in Flo s arms Flame disappeared shortly after she was born there has been speculation she may have been killed by her jealous brother Flint 40 Freud edit Freud May 1971 March 2014 was Fifi s oldest son 41 His father was a male from outside the Kasekela community 42 As a juvenile he benefited from a high ranking mother and an uncle Figan who was the alpha male 41 In 1993 Freud became the alpha male by unseating his childhood playmate Wilkie 41 While the alpha male his younger but larger brother Frodo became the second highest ranking male in the community In 1997 while Freud was afflicted by sarcoptic mange Frodo was able to take over the alpha status 41 43 44 After losing his alpha status Freud settled into a role as a middle ranking male although could still threaten higher ranking males when aroused 28 45 He was also able to serve as an important ally to other males 43 Although less aggressive than his brother Frodo Freud is known to have engaged in at least one instance of infanticide killing a Kasekela infant named Tofiki in 2004 30 In his old age his younger brothers sometimes provided him support such as sharing meat and honey with him 43 The documentary Chimpanzee Family Fortunes ends with a scene of the orphaned young female Flirt finding solace after several lonely months by getting groomed by her older brother Freud 46 He fathered at least one infant the female Cocoa to Candy born in 2004 47 48 49 Freud s approach to maintaining his alpha status was different from his smaller predecessor Wilkie and his larger successor Frodo 50 At 99 pounds 45 kg Freud was larger than the 81 pound 37 kilogram Wilkie and smaller than the 113 pound 51 kilogram Frodo 50 Freud was less aggressive than and groomed other males to help maintain alliances more often than Frodo but engaged in more contact aggression and less grooming than Wilkie 28 45 50 While alpha male he was able to intimidate even the larger Frodo with his displays 44 Freud died in March 2014 at more than 42 years old which made him the oldest male whose age was known in the history of the community 25 43 Freud was one of the first wild chimpanzees whose entire life from birth to old age was documented by researchers 51 Frodo edit Frodo June 30 1976 November 10 2013 was Fifi s second oldest son 52 His father was the relatively low ranking male Sherry Even from a young age Frodo was unusually large and aggressive He learned to throw rocks as a juvenile and became very accurate with his aim he sometimes threw them at his human observers often hitting and bruising them 53 As an adult he was one of the largest chimpanzees ever observed in the community at about 113 pounds 51 kg and remained quite aggressive 50 52 He also became an excellent hunter of red colobus monkeys and was also able to intimidate other chimpanzees into sharing their kills with him if he was unsuccessful 28 54 From 1990 to 1995 Frodo killed 10 of the red colobus monkeys within his home range 33 His large size and aggressive nature allowed him to attain high status in the group while his brother Freud was alpha male On October 2 1997 while Freud and other community members were suffering from sarcoptic mange Frodo attacked and defeated Freud thus attaining the alpha male position 28 52 54 44 Researchers speculated that Frodo was unaffected by the mange because he interacted less with the other apes giving the parasite less opportunity to infect him 44 After taking over the alpha male position Frodo demonstrated his new power by protecting his vanquished brother from the other males typically the former alpha is attacked by the other males and forced to leave the community for a period of time 55 As alpha male Frodo maintained his position largely through intimidation 28 50 54 He rarely groomed other males and often demanded that other males groom him 28 50 54 Frodo remained alpha male until becoming ill himself in 2002 28 46 54 56 he was then defeated by a coalition of several males and spent most of the next two years alone recovering from his wounds and illness 28 46 54 56 He received antibiotics from the researchers which likely helped his recovery 57 In 2004 he once again began travelling with the rest of the community but was unable to re establish his alpha status 28 46 54 In his later years Frodo mellowed He became less aggressive and spent much time by himself although he could still intimidate other chimpanzees on occasion 58 Frodo died in 2013 59 Frodo s aggression was not limited to colobus monkeys and other chimpanzees In May 2002 he killed a 14 month old human child that the niece of a member of the research team had carried into his territory 57 As a result the Tanzanian National Parks Department considered killing Frodo 57 In 1988 he attacked visiting Far Side cartoonist Gary Larson leaving him bruised and scratched 57 Frodo had a history of attacking the researchers observing him Goodall was attacked by Frodo on multiple occasions and in 1989 the ape beat her head so violently her neck was nearly broken 57 Frodo fathered at least eight infants second only to group male Wilkie who fathered ten 59 60 61 Infants he fathered included Zeus 1993 by Trezia 25 Zeus was part of a coalition that helped topple his uncle Ferdinand from alpha male status 62 Titan 1994 a large aggressive adolescent male by Patti 25 63 64 Fred 1996 1997 by Fifi Frodo s own mother 25 31 42 33 Golden and Glitter 1998 female twins and the oldest chimpanzee twins known to have survived in the wild by Gremlin 25 31 46 44 56 Tarzan 1999 Titan s brother by Patti 25 Samwise 2001 an aggressive female by Sandi 25 Sinbad 2001 Sparrow s youngest son 25 Fanni edit Fanni b March 19 1981 is Fifi s oldest daughter 13 Her father was Goblin 25 42 65 She had her first infant Fax at just 11 years old making her one of the youngest mothers seen in the community although Fax did not survive to adulthood 13 66 She has also had five other children males Fudge 1996 Fundi 2000 Fifty 2010 and Ficma 2015 and females Familia 2004 and Fadhila 2007 67 Fudge Fundi and Familia were fathered by Sheldon who would later become alpha male 25 Familia disappeared in 2012 citation needed Fadhila was fathered by Wilkie another former alpha male 25 42 Fifty was also sired by Wilkie 48 49 As of 2021 Fudge is the alpha male of the community 55 Whilst most female chimpanzees leave their natal communities upon reaching sexual maturity 68 Fanni has remained in her natal Kasekela community alongside her mother in contrast to her younger sister Flossi Fanni remained close to her mother Fifi with whom she formed a close alliance against the other community females Although males are perceived as the more aggressive sex in chimpanzees studies at Gombe have revealed that there is substantial aggression among females with each other 34 One of two contexts for female female aggressive encounters in Gombe was the presence of newborn infants 34 Fifi and Fanni were a formidable force and they carried out infanticidal attacks on other lower ranking females in her community Such attacks were observed on fellow Kasekela female Gremlin on three different occasions the first being in 1993 following the birth of her daughter Gaia and then five years later following the birth of her twins This was followed by another attack on Gremlin six years later following the birth of her next baby 34 Researchers could only speculate that the attacks on infants of other females had to do with food competition as each female strives to secure resources for her own group of offspring and thus the expansion of another female s family unit could deplete resources The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom in the 1970s 34 which proved that cannibalism although deemed rare does occur among chimpanzees by both males and in this particular case jealous females The loss of her mother in 2004 greatly reduced Fanni s dominance and she shifted her range to the southern part of the Kasekela territory further away from the other community females 69 Flossi edit Flossi b February 8 1985 70 is Fifi s second oldest daughter Her father was Goblin 47 Unlike most other female chimpanzees in the F family upon reaching adulthood she emigrated from the Kasekela community to join the neighbouring Mitumba Chimpanzee Community in 1996 although her younger sister Flirt who was born after Flossi emigrated later joined her 25 Although it is typical for female chimpanzees to emigrate from their natal group upon reaching maturity within the Kasekela community only about half the females do so Flossi may have chosen to emigrate to avoid mating attempts from her brother Frodo 55 Before emigrating from the Kasekela community Flossi may have innovated the practice of fishing for ants to eat using a stick which had not been observed before within the Kasekela community although fishing for termites had been but soon after the practice spread throughout the community 71 Flossi has given birth to four offspring the males Forest 1997 and Fansi 2001 and the females Flower 2005 and Falida 2009 72 Flossi became a high ranking female within the Mitumba community more typically immigrating females remain at a low rank 55 Faustino edit Faustino b May 8 1989 is Fifi s third oldest son 13 His father was brother Freud s rival Wilkie 25 42 At the beginning of the documentary Chimps So Like Us the infant Faustino is shown with mother Fifi and sister Flossi trying out facial expressions 73 Fifi became pregnant less than three years after Faustino was born an unusually short time for a chimpanzee 44 74 As a result Faustino had to be weaned at an unusually young age leading to a particularly difficult weaning conflict 44 74 This may account for Faustino being more skittish as a juvenile than his younger brother 44 Faustino grew into a large but mellow adult who reminded observers of Freud 74 75 He had worked his way up to become the beta or second highest ranking male in the community until struck by a near fatal disease in 2005 74 Although Faustino survived he lost his high rank 74 After recovering Faustino slowly climbed his way back up the hierarchy but it appears he will never become alpha male However he did help his younger brother Ferdinand attain alpha status in 2008 and continued to provide important support for Ferdinand in his brother s role as alpha although as of early 2014 he would sometimes ally himself with Ferdinand s potential rival Titan 25 76 77 He is the father of Gaia s son Google born in 2010 48 49 Ferdinand edit Ferdinand b August 19 1992 78 is Fifi s fourth oldest son 13 His father was his uncle Figan s rival Evered who was 41 years old when Ferdinand was conceived 25 42 He became the first wild great ape to have his birth recorded 33 By 2007 he had grown into a large enough male to wound the alpha male Kris and researchers believed he would eventually become the third of Fifi s sons to become alpha male 79 80 In 2008 he succeeded in supplanting Kris as alpha male after wounding Kris badly in a fight 62 81 Ferdinand maintained his alpha male position primarily through intimidation and threats and even surprising attacking males 62 55 He would bite rivals on the back to leave a scar 62 55 By the last two years of his reign Ferdinand s aggressiveness had made him so unpopular with other group males that he spent most of his time apart from them returning to assert his dominance every so often 62 During that time a coalition of rival males formed that included his nephews Fudge and Fundi and their father and uncle Sheldon and Sampson and another nephew Zeus 62 55 In October 2016 the coalition attacked Ferdinand when he made one of his occasional appearances to try to assert his continuing dominance 62 Fudge and Fundi led the attack with some support from their mother and Ferdinand s sister Fanni 62 55 Ferdinand was badly wounded in the attack and had to leave the group to recover 62 In his absence Fudge was able to assert his dominance and attain the alpha position 62 Ferdinand has been successful in siring offspring Infants he fathered included Diaz by Dilly fathered at very young age of 12 in 2005 Tabora 2006 by Tanga and Gizmo 2010 2016 by Gremlin 48 49 59 61 82 83 84 85 Flirt edit Flirt b July 1998 is Fifi s youngest surviving daughter 13 Her father was Kris who would later become alpha male of the troop 25 She was a large baby and grew rapidly 86 This probably allowed her to survive when her mother died before she was seven years old 46 86 Although she spent a few lonely months after Fifi s death she would begin to travel with her brothers 86 The documentary Chimpanzee Family Fortunes ends with Flirt finding solace after months of loneliness by being groomed by eldest brother Freud 46 She followed in the footsteps of her sister Flossi in eventually emigrating to the Mitumba Chimpanzee Community in 2013 25 Fudge edit Fudge b 1996 is Fanni s oldest surviving son His father is Sheldon a former alpha male He is the fifth male of the F family to reach alpha male status overthrowing his own maternal uncle Ferdinand in October 2016 62 Fudge had joined a coalition that included his brother Fundi his father Sheldon his uncle Sampson and his cousin Zeus in opposition to Ferdinand 62 In October 2016 Fudge led an attack by the rival males on Ferdinand 62 55 Fundi was his primary supporter in the attack and they received some support from their mother Fanni who is Ferdinand s sister 62 55 After seriously wounding Ferdinand in the attack Fudge was able to assert his dominance over the remaining group males to attain alpha male status 62 55 Fundi edit Fundi b 2000 is Fanni s second oldest surviving son His father is the former alpha male Sheldon Fudge the alpha male as of 2016 is his full brother In 2016 Fundi helped Fudge attack their uncle Ferdinand the previous alpha male which allowed Fudge to attain his alpha status 62 G family editMelissa edit Melissa c 1950 1986 31 was a high ranking female and mother of long term alpha male Goblin 87 Researchers also suspected that she was related to alpha male Humphrey and another male Mr McGregor 87 She was afflicted by polio in the 1966 epidemic and for a while was paralyzed in her neck and shoulders and was forced to walk bipedally She would regain use of her arms although she never fully regained movement of her neck 87 She gave birth to two sets of twins one set in 1976 of which neither survived the year and another set in 1977 of which only one Gimble survived to adulthood 31 87 Her other offspring who survived to adulthood was daughter Gremlin 31 Her daughter Gremlin and her granddaughter Gaia have also given birth to twins Goblin edit Goblin September 1964 August 2004 was Melissa s oldest son 31 65 He was discovered by Goodall when he was just a few hours old with a twisted face that led her to name him Goblin 88 Goblin was protected by Figan as an adolescent but at the young age of 15 in 1979 he challenged and defeated Figan to become the top ranking male 18 89 He was defeated and brutally beaten by a coalition of older males later in the year giving up his top ranking 18 89 Goblin regained the top ranked position in 1982 and was the unambiguous alpha male able to control even coalitions of rival males by 1984 18 Goblin maintained his alpha status until 1989 when he was badly beaten and injured by a coalition of males led by Wilkie 7 65 73 He was sterilized by a bite to his testicles in the process 43 He spent time away from the rest of the community recovering from his injuries during which time Goodall administered medication to help him return to health 73 He later became Freud s key ally in becoming and retaining his position as alpha male apparently going so far as to encourage Freud to challenge Wilkie for the role 55 While Freud was alpha Goblin was able to be a wedge between Freud and Frodo who had previously been close companions but when Frodo overtook Freud Goblin switched his allegiance to Frodo 55 Goblin died of disease in August 2004 30 After recovering from his injuries Goblin maintained a high ranking status within the community by forming alliances with the alpha males especially Freud and Frodo 65 90 While he was establishing himself as a contender for the alpha position he would occasionally beat up Goodall 65 In 1983 when his mother Melissa was in estrus he forced himself upon her despite her efforts to resist 89 Fanni and Flossi by Fifi and Tanga 1989 by Patti are his daughters 47 63 65 91 Gremlin edit Gremlin b 1970 is Melissa s only daughter to survive to adulthood 31 Her father was Evered who more than 20 years later also fathered the alpha male Ferdinand 25 42 Goodall has described Gremlin as one of her favourite chimpanzes due to Gremlin s patience as a mother and expertise in fishing for termites 56 She has raised twins Golden and Glitter who are the oldest known wild twin chimpanzees 33 She had male offspring in Getty 1982 1986 Galahad 1988 2000 Gimli 2004 Gizmo 2010 2016 and a female offspring in Gaia 1993 31 67 In 2015 she gave birth to Grendel and in 2019 at the age of almost 50 she gave birth to a son Goodali 92 55 Although males are perceived as the more aggressive sex in chimpanzees studies at Gombe have revealed that aggression between females does occur 34 Such attacks were observed of Fifi and her adult daughter Fanni aggressively harassing fellow Kasekela female Gremlin on three different occasions the first being in 1993 following the birth of her daughter Gaia and then five years later following the birth of her twins This was followed by another attack on Gremlin six years later following the birth of her next baby 34 Researchers could only speculate that the attacks on infants of other females had to do with food competition as each female strives to secure resources for her own group of offspring and thus the expansion of another female s family unit could deplete resources 29 The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom in the 1970s 34 which proved that cannibalism although deemed rare does occur among chimpanzees by both males and in this particular case jealous females Clever females like Gremlin who bore the brunt of infanticidal females employed two counter strategies to avoid attacks on them and newborns 34 One strategy is being in close proximity to adult males for protection as Gremlin initially did by sticking close to Frodo the father of her twins in 1998 34 The second strategy used by Gremlin was taking a newborn baby from her daughter Gaia in 2006 presumably in fear of it being attacked 34 Although Gremlin s instincts were justified from her own experiences the latter case ended with the infant dying just five months later 34 Gimble edit Gimble b October 1977 is one of two of Melissa s sons to survive to adulthood 31 He was one of a set of twins however his twin brother Gyre died in 1978 31 As a twin he grew into a healthy but small adult 93 He reached a relatively high rank in the late 1990s but fell to a low ranking male by 2004 30 He fathered at least two children Mambo by Malaika born in 2004 and Zinda 2006 October 2012 by Trezia 25 47 48 49 He disappeared in early 2007 citation needed Galahad edit Galahad 1988 2000 was Gremlin s third son 94 His father was a low ranking male Atlas 95 96 Galahad was described by researchers at Gombe as mischievous and playful often observed terrorizing baboons 95 In the BBC television program Chimpanzee Diary 1998 Galahad was depicted being supportive of his mother Gremlin staying near by even though he could not intervene during her attack by other community females trying to take her newborn twins He was developing into a favourite character for the human observers in Gombe but was unfortunately struck down by the flu epidemic in 2000 and died in February of that year 95 Gaia edit Gaia b February 1993 is Gremlin s oldest daughter 31 Her father was Wilkie 25 42 As a juvenile she helped her mother raise the twins Glitter and Golden and this is shown in a number of documentaries about the community 28 46 56 97 Gaia gave birth to her first baby in April 2006 34 but the baby s grandmother Gremlin took the infant from her daughter presumably in fear of it being attacked 34 Although Gremlin s instincts were justified from her own experiences of other community females attacking her newborns 34 the infant died just five months later 22 34 Gaia s second infant was stillborn but her third pregnancy resulted in twins 98 However once again her mother took over the babies and one of the twins died at a few days old She finally gave birth to a son named Google in 2010 fathered by Faustino 48 49 99 100 67 She gave birth to a son named Gabo in 2015 67 Glitter and Golden edit Glitter and Golden b July 1998 are the twin daughters of Gremlin fathered by Frodo 31 46 44 56 61 Gaia contributed to the raising of Gremlin s twins helping them to become the only known pair of wild chimpanzee twins known that have survived to the age of sexual maturity 46 56 Each of the pair exhibits a different personality Glitter is shy and cautious while Golden is more adventurous and willing to engage in rough and tumble activities 28 101 In 2011 Glitter gave birth to daughter named Gossamer who was fathered by Sheldon 102 67 She gave birth to a son named Ghurubu in 2016 67 In 2012 Golden gave birth to a daughter named Glamour 48 49 102 Golden also gave birth to a son named Gombe in 2017 Golden and Glitter have remained extremely close through adulthood 67 P family editPassion edit Passion c 1951 1982 was a central female of the Kasekela community She along with her daughter Pom captured killed and ate about ten newborns at Gombe 103 Passion possessed a rather domineering and aggressive personality according to Goodall Passion was an unnatural and indifferent mother to her three surviving offspring but she became a very high ranking female She became more solitary later in life and died of some type of wasting disease in 1982 In addition to Pom Prof and Pax she gave birth to a baby in 1962 which died in infancy She also had a stillborn baby or possibly a miscarriage in 1970 Pom edit Pom 1965 1983 was Passion s oldest known offspring to survive to adulthood Pom was instrumental in helping her mother capture and kill community infants 103 Pom gave birth herself to Pan in 1978 and was very cautious with the infant around her mother Pan died of injuries from an accident in 1981 Pom eventually disappeared about a year after her mother s death She may have transferred to another community but she was never seen again Prof edit Prof October 27 1971 198 26 was a male that survived to adulthood He and Pom offered their little brother comfort following Passion s death Prof and Pax associated closely for a number of years Prof is deceased Pax edit Pax b December 16 1977 26 was Passion s last offspring He was only four years old when he was orphaned He had the support of an elder brother and sister and had very few behavioural issues following his mother s death When Pax was very young his mother was wounded in some type of conflict Pax lost his testicles as a result of the conflict which has affected his personality greatly 104 He is small for a male doesn t compete in the male hierarchy and is unable to reproduce 30 104 85 He also has been chosen by a couple of orphans as a companion Pan edit Pan October 17 1978 1981 26 was the only known offspring of Pom He was injured after being blown from a palm tree by a strong wind and died four days later T family editPatti edit Patti c 1961 2005 was not born in the Kasekela community but joined the community in 1973 28 She was an assertive female who spent much time on her own 28 Her first two infants died before reaching adulthood but she has successfully raised several chimps since 28 Besides Tapit Tita Tanga Titan and Tarzan she had two offspring who died in infancy including her first born Pibi 1978 28 and a baby of unknown sex in 1988 She was killed by males from the rival Mitumba Chimpanzee Community in 2005 1 A female chimpanzee leaving the safety of her community territory poses risks and studies on chimpanzee intercommunity relations revealed that chimpanzees are likely to be attacked by the territorial males of a rival community 36 and the intensity of the attacks depends on the gender of the intruder and if the stranger is female it depends on whether or not they are of any reproductive benefit to the males Young oestrus females are often spared an attack because they are deemed to be potential new mates but an anestrus female non cycling or lactating is considered of very little use for the males which results in lethal aggression and fatality 36 Tapit edit Tapit 1979 1983 26 was Patti s second oldest son He had a rough life and was sickly as an infant and died as a juvenile 28 Tita edit Tita b January 1984 is Patti s oldest daughter She gave birth to her first child Tibi in 1995 he died in infancy Another son Tofiki fathered by Sheldon was born in 2000 and was killed in 2004 when Freud attacked him while grooming Tita 30 85 Tanga edit Tanga b April 1989 is Patti s second oldest daughter Her father was Goblin 91 85 She still lives in the community and gave birth to her first infant Tom in 2001 25 In 2006 she gave birth to a daughter named Tabora 25 In October 2012 she gave birth to a daughter named Tarime who died in 2013 25 In 2014 she gave birth to Tukuyu whose father is mostly likely Fudge citation needed Titan edit Titan b August 1994 is Patti s third oldest son fathered by Frodo 42 Titan displays aggressive behaviour resembling his father Frodo s violent tendencies Like Frodo as a youngster he would throw rocks at researchers 76 Also like his father Titan is large and as of 2013 was the largest male in the community 76 As of 2012 researchers believed that he may have been ready to challenge Ferdinand for alpha status as he spent a lot of time near Ferdinand and sometimes helped Ferdinand maintain control 76 However over the following year Titan slipped in rank 76 Researchers suspect that he may be afraid to challenge for the alpha spot and also that his reluctance to groom other chimps may inhibit him from forming the alliances needed to attain alpha status 76 As of early 2014 Titan had occasionally been able to ally himself with Ferdinand s older brother Faustino in attempts to challenge Ferdinand but he has not been able to maintain that alliance reliably and at times Faustino would also join with Ferdinand to attack Titan 77 Titan fathered a daughter Baseke by Bahati in 2010 48 49 Tarzan edit Tarzan b October 1999 is Patti s fourth oldest son His father was Frodo making him Titan s full brother 25 Tom edit Tom b March 2001 is Tanga s first son fathered by Kris 25 85 Tabora edit Tabora b 2007 is Tanga s first daughter Her father is Ferdinand 25 S family editSparrow edit Sparrow b 1958 is the oldest female and matriarch of the family 105 She has had seven offspring including four sons Sheldon Steve Spud and Sinbad and three daughters Sandi Barbet and Schweini 22 Four of the seven are alive Sandi Sheldon Sinbad and Schweini Her son Sheldon became the alpha male of the Kasekela community in 2004 Steve born 1989 and Spud born 1996 both died in infancy Sandi edit Sandi b October 1973 is Sparrow s oldest daughter and first offspring Sandi is one of the shyer females of the Gombe chimpanzees She grew up within the community but her mother was not social and as a result she still has a timid nature Sandi is still close to her mother Sparrow and the two prefer each other s company rather than anyone else s Sandi was very nervous of large groups of chimpanzees within the community and often prevented her offspring from playing with the others However she has become much more confident and her offspring have thrived Sandi has five offspring which include two daughters Sherehe and Samwise and two sons Sampson and Siri who are both sired by Apollo Barbet edit Barbet ca 1978 1992 was Sparrow s second oldest daughter She died in 1992 Sheldon edit Sheldon b May 1983 is Sparrow s oldest son Sheldon grew up within the community as a subordinate male but within 2004 following the vacancy of the alpha position after former alpha Frodo s departure when he became the alpha male Sheldon held on to dominance for only a year and lost the alpha position to another male Kris in 2005 30 He was part of a coalition that helped his son Fudge attain the alpha male position in 2016 62 He fathered Fanni s first three offspring Fudge Fundi and Familia 25 85 He was a low ranking male when he fathered Fudge and Fundi but had become the alpha male by the time he fathered Familia 85 He also fathered Tita s son Tofiki in 2000 while a mid ranking male 85 Schweini edit Schweini b April 1991 is Sparrow s third oldest daughter Her father is Wilkie 42 85 She gave birth to her first infant in September 2007 but she died in infancy 106 25 She gave birth to the female Safi on October 16 2008 but she died in August 2011 25 She also gave birth to a son Schall in November 2012 25 She was brutally attacked by females of the Mitumba chimpanzee community while associating with males of that community in both 2004 and 2005 106 Sinbad edit Sinbad b June 2001 is Sparrow s youngest son His father was Frodo 25 Sherehe edit Sherehe January 1991 May 11 2006 was Sandi s oldest daughter and first offspring 25 Her father was Beethoven 85 She gave birth to her first infant Shangaa in 2004 Sampson edit Sampson b April 1996 is Sandi s oldest son His father is Apollo He is a low ranking male Sampson was part of a coalition that included his uncle Sheldon and his cousin Fundi that helped Sampson s cousin Fudge supplant Ferdinand as alpha male in 2016 62 Samwise edit Samwise b June 2001 is Sandi s second oldest daughter Her father was Frodo 25 Male dominance editChimpanzees live in large mixed sexed communities in a society that is male dominant 107 and the male chimpanzee who rules above all others males and females is referred to as the alpha male The alpha male is the highest ranking most dominant chimpanzee in the entire community and can control most situations including situations in which he is opposed by a coalition of other males 18 Nonetheless an alpha male typically cannot monopolize breeding opportunities and in some cases lower ranking males have been more successful at mating than alpha males since the alpha male has to spend much energy maintaining his rank 108 Faben Figan s older brother with a paralyzed arm Evered defeated by Figan in his bid for alpha status who became Ferdinand s father and Sherry lower ranking male who became Frodo s father are examples of males who successfully devoted their energy to breeding success despite not attaining alpha status 108 However a very secure alpha male such as Frodo can use his status to increase his access to females in estrus especially during the period when she is at peak fertility 108 Goliath was the first chimpanzee Goodall recognized as the alpha male 7 109 In 1964 Mike who was previously a low ranking male deposed him 110 111 Goliath remained a high ranking male for several more years but fell in rank as he aged 109 Eventually Goliath was one of the chimpanzees who left the Kasekela community to form the Kahama community 109 However in 1975 he was brutally attacked by a group of Kasekela males led by Faben and died as a result of his injuries 20 109 Mike figured out that incorporating cans and other objects left by the human researchers into his displays would make them more effective These displays indeed allowed him to take over the alpha male rank in 1964 110 111 By 1969 a group of younger males including Humphrey Faben Figan and Evered challenged and defeated him making Humphrey the alpha male and returning Mike to his low ranking position 18 From 1972 through 1989 two males Figan and Goblin held the alpha position Figan was able to defeat Humphrey when Faben became his ally For a period after Faben s death in 1975 Figan remained the top ranking male but was unable to control coalitions of other males when they joined together making his status as alpha male ambiguous Within a few months Figan reestablished himself as the unambiguous alpha by forging alliances with other high ranking males such as Humphrey In 1979 15 year old Goblin who had previously been protected by Figan challenged him for the alpha position Goblin temporarily held the top ranking position in 1979 although could not control coalitions of other males before the more senior males brutally beat him and restored Figan to the alpha position By 1982 Goblin had reestablished himself as the top ranking male by 1984 he was unambiguously the alpha male able to control situations even against coalitions of other males 18 In 1989 Wilkie defeated Goblin and attained the alpha position 73 Wilkie attained this position despite being one of the smallest males in the community at 37 kilograms 82 lb 112 According to researchers at the University of Minnesota s Jane Goodall Institute Center for Primate Studies Wilkie attained his position primarily by becoming popular by obsessively grooming other males 104 112 Unlike most males Wilkie also groomed females 112 Wilkie also made effective use of charging displays 104 Wilkie was overthrown in 1992 by his childhood playmate Freud who was in turn deposed by his brother Frodo in 1997 while Freud was ill 41 52 Freud at 44 8 kg 99 lb was larger than Wilkie and relied on a mixture of moderate force and grooming to become alpha and maintain his status 104 112 Frodo at 51 2 kg 113 lb was one of the largest males ever seen in the group relied primarily on brute force to attain and maintain alpha status 104 112 After Frodo became ill in 2002 he was brutally beaten by a coalition of males led by Sheldon and spent two years travelling away from the rest of the community No chimpanzee was able to establish himself as an alpha male until 2004 Sheldon briefly held the top ranking male position in 2004 however by 2005 Kris had defeated all competitors including Frodo and established himself as the unambiguous alpha male 46 56 In 2008 Kris was supplanted as alpha male by Ferdinand the youngest surviving brother of Freud and Frodo 82 Unlike most deposed alpha males who tend to stay away from the group for a while in order to avoid further injury while the new alpha is consolidating his position Kris stayed with the group after being deposed and eventually died of his injuries 25 43 59 61 83 113 In October 2016 Ferdinand was attacked by a coalition of males led by his nephews Fudge and Fundi and also included the former alpha male Sheldon 62 Ferdinand was badly injured in the attack and Fudge asserted himself as the new alpha male 62 Name Reign Lifespan Notes ReferenceGoliath 1961 1964 1975 7 18 Mike 1964 1969 1975 7 18 110 111 Humphrey 1969 1971 ca 1947 1980 7 18 110 111 vacant 1971 1972 After Humphrey was deposed as alpha in 1971 no chimpanzee clearly held the highest rank until Figan established himself as alpha male over Humphrey and Evered in 1972 18 25 Figan 1972 1979 1953 1982 For a nine month period in 1975 and 1976 while Figan was clearly the highest ranking male he was not an unambiguous alpha male 16 18 Goblin 1979 1964 2004 In 1979 Goblin was able to oust Figan and become the top ranked male although he was not able to establish himself as an unambiguous alpha male before Figan regained his status 18 90 Figan 1979 1982 1953 1982 16 18 Goblin 1982 1989 1964 2004 Although Goblin established himself as the highest ranking male during 1982 he did not become the unambiguous alpha male until 1984 7 18 73 89 Wilkie 1989 1993 1972 2012 7 73 112 114 115 Freud 1993 1997 1971 2014 Nephew of Figan 7 41 114 Frodo 1997 2002 1976 2013 Replaced his brother Freud nephew of Figan 7 52 114 116 vacant 2002 2004 After Frodo was ousted as alpha male during his illness in 2002 no male established himself as unambiguously the highest ranking male until 2004 46 Sheldon 2004 2005 1982 Was part of a coalition that helped his son Fudge become alpha male in 2016 46 52 115 116 117 118 62 Kris 2005 2008 1982 2010 7 43 46 52 115 118 Ferdinand 2008 2016 1992 Youngest brother of Freud and Frodo nephew of Figan displaced by his sister s son Fudge 25 82 62 Fudge 2016 1996 Nephew of Freud Frodo and Ferdinand grandnephew of Figan son of Sheldon and grandson of Goblin 62 Female dominance editEven though chimpanzees live in a male dominant society 107 the female chimpanzees in the Kasekela community have evolved their own pecking order in which the top ranking female has priority access to food mates and sleeping sites Studies in Gombe have discovered that the rank of a mother directly correlates to infant survival fast maturing of daughters and faster production of offspring 119 Jane Goodall considered elderly female Flo the original matriarch of what is now the famous F family of Gombe to be the most dominant of the females in the Kasekela community at the time that the study began in the 1960s 120 and this was evident in her interaction with other females where she was never hesitant about asserting herself towards new females 121 She was a successful mother and fierce defender of her offspring one of whom would succeed her The dominance of a female to other community females depends largely on family ties 29 122 Fifi was a popular female in Gombe not only from her high ranking family but also from her close ties with the males She inherited her mother s superior status 120 and because her older brothers Faben and Figan were among the top big males she grew up with enormous confidence 29 As a mother of nine Fifi was fiercely defensive of her offspring 69 and her authority was rarely questioned This was also possible because of her ability to fearlessly navigate a male dominant society 69 helped in part from the fact that her adult sons Freud and Frodo grew to become the alpha males in the community Having produced the most offspring out of any female during her time Fifi was regarded as Gombe s most fertile female 123 and her ability to have reproduced as frequently as she did and have enough resources and social standing to ensure the survival of seven of her nine children validates the studies of how the social rank of a chimpanzee mother directly correlates to infant survival fast maturing of daughters and faster production of offspring 119 Just as her mother s high status rubbed off on her 29 Fifi s high standing in the community benefited her daughters Although second daughter Flossi emigrated to dominate another chimp community 34 her eldest daughter Fanni remained with her mother and together they became a formidable force in Gombe Observers in Gombe discovered that the linear hierarchy among females becomes apparent in aggressive interactions involving infanticide 34 One of two contexts for female female aggressive encounters in Gombe was the presence of newborn infants 34 Matriarch Fifi was known to be a feisty female who although a very successful mother herself was observed to have carried out ferocious and infanticidal attacks on other lower ranking females in her community Such attacks were observed of Fifi and her adult daughter Fanni aggressively harassing lower ranking Kasekela female Gremlin on three different occasions with the goal of killing her newborn infants 34 Researchers could only speculate that the attacks on infants of other females had to do with food competition as each female strives to secure resources for her own group of offspring and thus the expansion of another female s family unit could deplete resources 29 The attacks by Fifi and Fanni mirror those of previous perpetrators Passion and Pom a dominant pair who only attacked subordinate females in the 1970s 34 which proved that cannibalism although deemed rare does occur among chimpanzees by both males and in this particular case jealous females The loss of her mother Fifi in 2004 greatly reduced the dominance of Fanni now the oldest surviving female of the once mighty F group towards the other females and this is despite having high ranking male relatives in the community including her son Fudge who has been the top male since 2016 124 and her high ranking brothers Faustino and Ferdidand Fudge s predecessor Fanni shifted her range to the southern part of the Kasekela territory further away from the other community females 69 particularly Gremlin s growing G family which now makes up a large portion of the Kasekela community and who occupy the top of the hierarchy 69 Other communities of chimpanzees at Gombe Stream National Park editWithin the Gombe Stream National Park live other chimpanzee communities other than the Kasekela which also inhabit certain home ranges The two other communities currently bordering the Kasekela community include Mitumba Chimpanzee Community 125 Kalande Chimpanzee Community 126 One extinct community of Gombe was Kahama Chimpanzee CommunityThe Kahama community came about from a split within the Kasekela community 11 20 In the early 1970s several of the Kasekela chimpanzees including 8 adult males kept to the northern portion of the Kasekela range while others including 7 adult males kept to the southern portion of the range 11 20 The two groups became hostile towards each other and had less and less to do with each other and eventually the southern community became totally separate and was referred to by researchers as the Kahama community 11 20 By 1974 the Kasekela males began to attack the Kahama chimpanzees in what was referred to as the Four Year War 11 20 Figan Faben Humphrey Frodo s father Sherry and Sherry s brother Jomeo were often among the ringleaders of the attacks 20 Researchers described these attacks as being of the nature of raids as opposed to simply defending their territory 11 The Kasekela chimpanzees did not collect food during these raids although at least three Kahama females were brought back to the Kasekela community 11 12 20 Kahama chimpanzees were brutally attacked during these raids and often were killed 11 20 Former Kasekela alpha male Goliath who had joined Kahama and was then very old was killed in one such attack in 1975 20 Kahama s alpha male Charley was killed in 1977 20 By 1977 the Kahama community had been completely wiped out and the Kasekela community absorbed its territory 20 Annexing the Kahama territory brought the Kasekela community in contact with the Kalande Chimpanzee Community 11 20 The Kalande community may have been attacking Kahama chimpanzees during the Four Year War and after Kahama was wiped out they began to attack Kasekela chimpanzees 11 20 At least one Kasekela female was badly injured in an attack by Kalande chimpanzees and at least two infants were killed in such attacks 11 20 Former Kasekela alpha male Humphrey died in 1981 possibly from injuries from an attack by Kalande chimpanzees 11 Kasekela has had interactions with the Mitumba Chimpanzee Community as well Some of these interactions have been hostile and some Mitumba chimpanzees are known to have been killed by Kasekela chimpanzees 30 125 At least two Kasekela females Fifi s daughters Flossi and Flirt have emigrated from Kasekela to join the Mitumba community 25 125 Based on circumstantial evidence researchers believed that it is possible that Fifi herself may have died as a result of an attack by Mitumba chimpanzees 30 Books about the community editGoodall J 1971 In the Shadow of Man Houghton Mifflin ISBN 978 0 395 33145 3 Goodall J 1986 The Chimpanzees of Gombe Patterns of Behavior The Belknap Press of Harvard University Press ISBN 978 0 674 11649 8 Goodall J 1990 Through a Window Houghton Mifflin ISBN 978 0 395 59925 9 Goodall J 1997 The Chimpanzee Family Book North South Books ISBN 978 1 55858 803 5 Shah A Rogers F 2014 Tales from Gombe Firefly ISBN 978 1 77085 468 0 Films about the community editvan Lawick H Jampel B 1984 Among the Wild Chimpanzees National Geographic Hallet D Goodall J 1990 Jane Goodall My Life with the Chimpanzees National Geographic Goodman K amp Simon K 1990 Chimps So Like Us DVD HBO van Lawick H Sutherland D 1991 People of the Forest The Chimps of Gombe Discovery Channel Lickley D Low S Goodall J 2002 Jane Goodall s Wild Chimpanzees IMAX Bristow M Goodall J 2004 Return to Gombe Discovery Channel D Auvergne L 2006 Chimpanzee Family Fortunes BBC Brett Morgen 2017 Jane National Geographic Further reading editMorris Ian 2014 War What Is It Good For The Role of Conflict and the Progress of Civilisation from Primates to Robots MacMillan ISBN 978 1 84765 454 0 See also editGombe Chimpanzee War List of individual apesReferences edit a b Wilson M L 2012 Long term Studies of the Chimpanzees of Gombe National Park Tanzania In Kappeler Peter M Watts David P eds Long term Field Studies of Primates Springer pp 357 384 ISBN 9783642225130 a b Goodall J 1986 The Chimpanzees of Gombe Patterns of Behavior The Belknap Press of Harvard University Press p 84 ISBN 978 0 674 11649 8 a b Morgen Brett 2018 Jane film National Geographic Wilson Michael L et al 2020 Research and conservation in the greater Gombe ecosystem challenges and opportunities Biological Conservation 252 108853 Bibcode 2020BCons 25208853W doi 10 1016 j biocon 2020 108853 PMC 7743041 PMID 33343005 Goodall J 1986 The Chimpanzees of Gombe Patterns of Behavior The Belknap Press of Harvard University Press pp 535 539 ISBN 978 0 674 11649 8 Goodall J 1971 In the Shadow of Man Houghton Mifflin pp 35 37 ISBN 978 0 395 33145 3 a b c d e f g h i j k l Gombe Timeline The Jane Goodall Institute Archived from the original on December 7 2008 Retrieved 2009 02 28 Scientist Makes Unique Record of Chimpanzees Greeley Daily Tribune March 19 1964 p 34 Retrieved 2019 01 26 via newspapers com Harriman Jane September 2 1963 Wild Chimp is a Friendly Chap Boston Globe p 44 Retrieved 2019 01 26 via newspapers com a b Eppinga J 2009 They Made Their Mark An Illustrated History of the Society of Woman Geographers Globe Pequot pp 218 220 ISBN 978 0 7627 4597 5 a b c d e f g h i j k l Wrangham R Peterson D 1996 Demonic Males The Washington Post Retrieved 2015 01 14 a b Morris I 2014 Red in Tooth and Claw Why the Chimps of Gombe Went to War War What Is It Good For Conflict and the Progress of Civilization from Primates to Robots Farrar Straus and Giroux ISBN 978 0 374 28600 2 a b c d e f g h i j k l Flo s Family Tree University of Minnesota JGI Center for Primate Studies Archived from the original on 2009 02 25 Retrieved 2009 03 03 a b Lickley D Low S Goodall J 2002 Jane Goodall s Wild Chimpanzees IMAX Goodall J 1986 The Chimpanzees of Gombe Patterns of Behavior The Belknap Press of Harvard University Press pp 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